Skip to main content
Log in

Deer of the genus Megaloceros (Mammalia, Cervidae) from the Early Pleistocene of Ciscaucasia

  • Published:
Paleontological Journal Aims and scope Submit manuscript

Abstract

A new deer species, Megaloceros stavropolensis sp. nov., from the pre-Apsheronian sandy–clayey deltaic deposits of the Georgievsk sand pit (village of Podgornoe, Stavropol Region) is described. The fauna of large and small mammals from the bone beds of this locality is dated terminal Villanian, end of the MNQ 17 Zone, beginning of the Late Villafranchian, and assigned to the beginning of the Psekups Mammalian Assemblage. This is the earliest known member of the genus, which possesses antlers with well-developed palmation. The middle tine is absent and the posterior tine is a part of the antler palmation. Other specimens of Early Pleistocene Megaloceros are represented by fragments.

This is a preview of subscription content, log in via an institution to check access.

Access this article

Price excludes VAT (USA)
Tax calculation will be finalised during checkout.

Instant access to the full article PDF.

Similar content being viewed by others

References

  • Abbazzi, L., Remarks on the validity of the generic name Praemegaceros Portis 1920 and an overview on Praemegaceros species in Italy, Rend. Fis. Accad. Lincei. Ser., 2004, vol. 15, pp. 115–132.

    Article  Google Scholar 

  • Ambrosetti, P., Cromerian fauna of the Rome area, Quaternaria, 1967, vol. 9, pp. 267–283.

    Google Scholar 

  • Azzaroli, A., The deer of the Weybourn Crag and Forest Bed of Norfolk, Bull. Brit. Mus. Natur. Hist., 1953, vol. 2, no. 1, pp. 1–96.

    Google Scholar 

  • Azzaroli, A., Forest Bed elks and giant deer revisited, Zool. J. Linn. Soc., 1994, vol. 112, pp. 119–133.

    Article  Google Scholar 

  • Baigusheva, V.S., Fossil theriofauna of the Liventsovka sand pit (northeastern Sea of Azov Region), Tr. Zool. Inst. Akad. Nauk SSSR, 1971, vol. 69, no. 49, pp. 5–28.

    Google Scholar 

  • Baigusheva, B.C. and Titov, V.V., Review of records of ancient elephants based on the material of the Sea of Azov Region, in Mamont i ego okruzhenie: 200 let izucheniya (Mammoth and Its Environs: 200 Years of Studying), Moscow: GEOS, 2001, pp. 71–81.

    Google Scholar 

  • Baigusheva, V. and Titov, V., The evolution of eastern European meridionaloid elephants’ dental characteristics, Quatern. Int., 2012, vol. 255, pp. 206–216.

    Article  Google Scholar 

  • Baigusheva, V. and Titov, V., Large deer from the Villafranchian of Eastern Europe (Sea of Azov Region): Evolution and paleoecology, Quatern. Int., 2013, vol. 284, pp. 110–122.

    Article  Google Scholar 

  • Blumenbach, J., Handbuch der Naturgeschichte, Gottingen: Johann Christian Dieterich, 1799, vol. 16.

    Google Scholar 

  • Brookes, J., A Catalogue of the Anatomical and Zoological Museum, London: Joshua Brookes, 1828, part 1.

    Google Scholar 

  • Croitor, R., Stefaniak, K., Pawlowska, K., et al., Giant deer Megaloceros giganteus Blumenbach, 1799 (Cervidae, Mammalia) from Palaeolithic of Eastern Europe, Quatern. Int., 2014, vols. 326–327, pp. 91–104.

    Google Scholar 

  • Dietrich, W.O., Neue Riesenhirschreste aus dem schwabischen Diluvium, Jahrb. Ver. Vaterland. Naturk. Württemberg, 1909, vol. 65, pp. 132–161.

    Google Scholar 

  • Garutt, W.E., A skeleton of the southern elephant, Archidiskodon meridionalis (Nesti, 1825), from a sand-pit near Georgievsk, northern Caucasus, Russia, Cranium, 1990, vol. 15, no. 1, pp. 33–38.

    Google Scholar 

  • Garutt, V.E. and Safronov, I.N., A find of a skeleton of the southern elephant Archidiskodon meridionalis (Nesti) near the town of Georgievsk (northern Caucasus), Byull. Kom. Izuch. Chetvert. Per., 1965, no. 30, pp. 79–88.

    Google Scholar 

  • Gould, S.J., The origin and function of “bizarre” structures: Antler size and skull size in the “Irish "Elk”, Megaloceros giganteus, Evolution, 1974, vol. 28, pp. 191–220.

    Article  Google Scholar 

  • Gromova, V.I., Kratkii obzor chetvertichnykh mlekopitayushchikh Evropy (A brief review of Quaternary mammals of Europe), Moscow: Nauka, 1965.

    Google Scholar 

  • Hart, J., Description of the Skeleton of the Fossil Deer of Ireland (Cervus megaceros) in the Museum of the Royal Dublin Society, Dublin: Graisberry, 1825.

    Google Scholar 

  • Heintz, E., Les cervides villafranchiens de France et d’Espagne, Mém. Mus. Nat. Hist. Natur. Ser. C, 1970, vol. 1, pp. 1–303.

    Google Scholar 

  • Hu, Ch. and Qi, T., Gongwangling Pleistocene mammalian fauna of Lantian, Shaanxi, Palaeontol. Sin. Ser. C, 1978, vol. 155, no. 21, pp. 1–64.

    Google Scholar 

  • Hughes, S., Nayden, Th.J., Douady, Ch.J., et al., Molecular phylogeny of the extinct giant deer Megaloceros giganteus, Mol. Phylog. Evol., 2006, vol. 40, no. 1, pp. 285–291.

    Article  Google Scholar 

  • Khomenko, I.P., Materials of the paleontology of Tertiary and post-Tertiary mammals of the northern Caucasus, Tr. Stavropol. Ob-va Izuch. Severokavkaz. Kraya, 1914, vol. 1, no. 1, pp. 1–39.

    Google Scholar 

  • Kuehn, R., Ludt, Ch.J., Schroeder, W., and Rottmann, O., Molecular phylogeny of Megaloceros giganteus—the giant deer or just a giant red deer?, Zool. Sci., 2005, vol. 22, no. 9, pp. 1031–1044.

    Article  Google Scholar 

  • Lebedeva, N.A., Korrelyatsiya antropogenovykh tolshch Ponto-Kaspiya (Correlation of the Anthropogene Strata of the Pontian–Caspian Sea), Moscow: Nauka, 1978.

    Google Scholar 

  • Lister, A.M., Megaceros or Megaloceros? The nomenclature of the giant deer, Quatern. Newslet., 1987, vol. 52, pp. 14–16.

    Google Scholar 

  • Lister, A.M., The evolution of the giant deer, Megaloceros giganteus (Blumenbach), Zool. J. Linn. Soc., 1994, vol. 112, pp. 65–100.

    Article  Google Scholar 

  • Lister, A.M., Edwards, C.J., Nock, D.A.W., et al., The phylogenetic position of the ‘giant deer’ Megaloceros giganteus, Nature, 2005, vol. 438, no. 8, pp. 850–853.

    Article  Google Scholar 

  • Lönnberg, E., Which is the taxonomic position of the Irish Giant Deer and allied races?, Arkiv Zool., 1906, vol. 3, no. 14, pp. 1–8.

    Google Scholar 

  • Made, J., van der, The evolution and biogeography of the Pleistocene giant deer Megaloceros giganteus (Cervidae, Mammalia), Cour. Forsch. Senckenb, 2006, vol. 256, pp. 117–129.

    Google Scholar 

  • Made, J. and Tong, H.W., Phylogeny of the giant deer with palmate brow tines Megaloceros from West and Sinomegaceros from East Eurasia, Quatern. Int., 2008, vol. 179, no. 1, pp. 135–162.

    Article  Google Scholar 

  • McKenna, M.C. and Bell, S.K., Classification of Mammals above the Species Level, New York: Columbia Univ. Press, 1997.

    Google Scholar 

  • Osnovy paleontologii (Fundamentals of Paleontology), Moscow: Gos. Nauch.-tekh. Izd-vo Lit. Geol. Okhr. Nedr, 1962. 421.

  • Owen, R., Report on the British fossil Mammalia, Brit. Ass. Advanc. Sci., 1844, pp. 208–241.

    Google Scholar 

  • Pavlow, M., Les selenodontes posttertiares de la Russie, Mém. Acad. Sci. Petersburg. Ser. 8, 1906, vol. 20, no. 1, pp. 20–39.

    Google Scholar 

  • Pfeiffer, T., The first complete skeleton of Megaloceros verticornis (Dawkins, 1868) (Cervidae, Mammalia) from Bilshausen (Lower Saxony, Germany): Description and phylogenetic implications, Mitt. Mus. Naturk. Berl. Geowiss., 2002, vol. 5, pp. 289–308.

    Google Scholar 

  • Pidoplichko, I.G. and Flerov, K.K., A new deer taxon from the Pliocene of southern Ukraine, Dokl. Akad. Nauk SSSR, 1952, vol. 84, no. 6, pp. 1239–1242.

    Google Scholar 

  • Pohlig, H., Die Cerviden des thuringischen Diluvial–Travertines mit Beitragen uber andere diluviale und uber recente Hirschformen, Palaeontograph, 1892, vol. 39, pp. 215–263.

    Google Scholar 

  • Radulesco, C. and Samson, P., Sur un nouveau cerf megacerin du Pleistocène moyen de la depression de Brasov (Roumanie), Geol. Rom., 1967, vol. 6, pp. 317–344.

    Google Scholar 

  • Reynolds, S.H., The British Pleistocene Mammalia: The giant deer, Palaeontogr. Soc. London Monogr., 1929, vol. 81, pp. 1–61.

    Google Scholar 

  • Rütimeyer, L., Beiträge zu einer natürlichen Geschichte der Hirsche, Abh. Schweiz. Paläontol. Ges., 1883, vol. 10, pp. 1–120.

    Google Scholar 

  • Shcheglova, V.V., On the taxonomic position and developmental history of the giant deer, Dokl. Akad. Nauk SSSR, 1950, vol. 73, no. 6, pp. 813–816.

    Google Scholar 

  • Shikama, T., Discovery of a giant fallow deer from the Pleistocene of Japan, Jap. J. Geol. Geogr., 1938, vol. 16, nos. 1–2, pp. 115–122.

    Google Scholar 

  • Shpansky, A.V., Skeleton of the giant deer Megaloceros giganteus giganteus (Blumenbach, 1803) (Mammalia, Artiodactyla) from the Irtysh Region near Pavlodar, Paleontol. Zh., 2014, no. 5, pp. 81–98.

    Google Scholar 

  • Simpson, G.G., The principles of classification and a classification of mammals, Bull. Am. Mus. Natur. Hist., 1945, vol. 85, pp. 1–350.

    Google Scholar 

  • Smith, C.H., Supplement to the order Ruminantia, in The Class Mammalia Arranged by the Baron Cuvier, with Specific Description, Griffith, E., Smith, C.H., and Pidgeon, E., Eds., London, 1827.

    Google Scholar 

  • Teilhard de Chardin, P., Fossil mammals from locality 9 of Choukoutien, Palaentol. Sin., 1936, vol. 7, no. 4, pp. 5–70.

    Google Scholar 

  • Tesakov, A.S., Biostratigrafiya srednego pliotsena-eopleistotsena Vostochnoi Evropy (Biostratigraphy of the Middle Pliocene–Eopleistocene of Eastern Europe), Moscow: Nauka, 2004.

    Google Scholar 

  • Titov, V.V., Krupnye mlekopitayushchie pozdnego pliotsena Severo-Vostochnogo Priazov’ya (Late Pliocene Large Mammals of the Northeastern Sea of Azov Region), Rostov-onDon: YuNTs Ross. Akad. Nauk, 2008.

    Google Scholar 

  • Vereshchagin, N.K., Remains of mammals from the Lower Quaternary beds of the Taman Peninsula, Tr. Zool. Inst. Akad. Nauk SSSR, 1957, vol. 22, pp. 9–49.

    Google Scholar 

  • Vereshchagin, N.K., Mlekopitayushchie Kavkaza (Mammals of the Caucasus), Moscow: Akad. Nauk SSSR, 1959.

    Google Scholar 

  • Viret, J., Artiodactyla, Traité de Paléontologie, Paris: Masson et Cie, 1961, vol. 6, no. 1, pp. 887–1084.

    Google Scholar 

  • Vislobokova, I.A., Fossil deer of Eurasia, Tr. Paleontol. Inst. Akad. Nauk SSSR, 1990, vol. 240, pp. 1–208.

    Google Scholar 

  • Vislobokova, I.A., History of the giant deers (Megacerini, Cervidae, Artiodactyla), Tr. Paleontol. Inst. Ross. Akad. Nauk, 2012a, vol. 293, pp. 1–103.

    Google Scholar 

  • Vislobokova, I.A., Giant deer: origin, evolution, role in the biosphere, Paleontol. J., 2012b, vol. 46, no. 7, pp. 643–775.

    Article  Google Scholar 

  • Vislobokova, I.A., Morphology, taxonomy, and phylogeny of megacerines (Megacerini, Cervidae, Artiodactyla), Paleontol. J., 2013, vol. 47, no. 8, pp. 833–950.

    Article  Google Scholar 

  • Young, Ch.-Ch., On the Artiodactyla from the Sinanthropus site at Chouk’outien, Palaentol. Sin., 1932, vol. 8, no 2, pp. 1–100.

    Google Scholar 

Download references

Author information

Authors and Affiliations

Authors

Corresponding author

Correspondence to V. V. Titov.

Additional information

Original Russian Text © V.V. Titov, A.K. Shvyreva, 2016, published in Paleontologicheskii Zhurnal, 2016, No. 1, pp. 82–90.

Rights and permissions

Reprints and permissions

About this article

Check for updates. Verify currency and authenticity via CrossMark

Cite this article

Titov, V.V., Shvyreva, A.K. Deer of the genus Megaloceros (Mammalia, Cervidae) from the Early Pleistocene of Ciscaucasia. Paleontol. J. 50, 87–95 (2016). https://doi.org/10.1134/S0031030116010111

Download citation

  • Received:

  • Published:

  • Issue Date:

  • DOI: https://doi.org/10.1134/S0031030116010111

Keywords

Navigation