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Saharan wheats: before they disappear

  • Notes on Neglected and Underutilized Crops
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Abstract

Wheat landraces cultivated in the Saharan oases have been submitted during centuries to drought, heat and salinity and are expected to have developed tolerance to these constraints. The different forms present in the oases of Libya, Algeria and Morocco have been collected and described by several authors who highlighted the peculiarity and diversity of this germplasm, on the basis of spike and grain morphology. The origin, date and way of introduction of wheat in the Saharan oases are however largely unknown. Most landraces may have been introduced from Egypt, possibly during wet climatic episodes. Spike morphological traits of some landraces suggest an Asiatic origin (Iran, Turkestan and Afghanistan). More recent introductions have also probably occurred. This lack of information as well as the potential interest of this germplasm for wheat improvement would merit further studies. Marker-based diversity analysis of Saharan wheats and comparison with North-African and Asian wheats might permit to precise their classification and confirm their origin. Evaluation for tolerance to abiotic constraints would allow a better understanding of their adaptation to the oases environments and identification of potential progenitors to be used by wheat breeders. Such studies are urgent considering the quick genetic erosion of this germplasm due to drastic ecological and socio-economic changes affecting Saharan oases and their cropping systems.

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Notes

  1. See in “Appendix” an index of geographical names cited in the text.

References

  • Adams R (1810) A narrative of Robert Adams, a sailor who was wrecked on the western coast of Africa in the year 1810. John Murray, London

  • Adams J (1998) Africa during the last 150,000 years http://www.esd.ornl.gov/projects/qen/nercAFRICA.html. Accessed 3 Nov 2013

  • Al Alazzeh A, Hammer K, Lehmann CO, Perrino P (1982) Report on a travel to the Socialist People’s Libyan Arab Jamahiriya 1981 for the collection of indigenous taxa of cultivated plants. Kulturpflanze 30:191–202

    Article  Google Scholar 

  • Al Khanjari SS (2005) Exploration and estimation of morphological and genetic diversity of wheat (Triticum ssp.) landraces in Oman. Kassel University Press, Kassel

  • Ammar K, Gharbi MS, Deghaïs M (2011) Wheat in Tunisia. In: Bonjean A, Angus WJ, van Ginkel M (eds) The world wheat book. A history of wheat breeding, vol 2. Lavoisier Publishing, Paris, pp 443–465

    Google Scholar 

  • Ballouche A, Marinval P (2003) Données palynologiques et carpologiques sur la domestication des plantes et l’agriculture dans le Néolithique ancien du Maroc septentrional (Site de Kaf Taht El-Ghar). Revue d’Archéométrie 27:49–54 (in French)

    Google Scholar 

  • Barbaza M (2005) Le Sahel des “siècles obscurs”. Données croisées de l’art rupestre, de l’archéologie, des chroniques et des traditions orales. Société Préhistorique Ariège-Pyrénées 60:61–102 (in French)

    Google Scholar 

  • Bellil I, Chekara Bouziani M, Khelifi D (2012) Genetic diversity of high and low molecular weight glutenin subunits in Saharan bread and durum wheats from Algerian oases. Czech J Genet Plant Breed 48:23–32

    CAS  Google Scholar 

  • Benlaghlid M, Bouattoura N, Monneveux P, Borries C (1990) Etude de la diversité génétique et de la physiologie de l’adaptation au milieu. Options Méditerranéennes 11:171–194 (in French)

    Google Scholar 

  • Broodbank C (2006) The origins and early development of Mediterranean maritime activity. J Mediterr Archaeol 19:199–230

    Google Scholar 

  • Brooks NPJ, Drake NA, McLaren SJ, White K (2003) Studies in geography, geomorphology, environment and climate. In: Mattingly DJ (ed) The Archaeology of Fazzān, volume 1, synthesis. The Society for Libyan Studies/Tripoli, The Department of Antiquities, London, pp 37–74

  • Bushuk W, Zillman RR (1978) Wheat cultivar identification by gliadin electrophoregrams. 1. Apparatus, method and nomenclature. Can J Plant Sci 58:505–515

    Article  Google Scholar 

  • Campano F (1996) Agriculture development in sub-Saharan Africa. Medit 4:29–33

    Google Scholar 

  • Chevalier A (1932) Ressources végétales du Sahara et de ses confins nord et sud. Muséum d’Histoire Naturelle, Paris, Laboratoire d’Agronomie Coloniale (in French)

  • Chevalier A (1938) Le Sahara, centre d’origine des plantes cultivées. In La vie dans la région désertique nord-tropicale de l’ancien monde. Société de Biogéographie VI, Paul Lechevalier ed, Paris, pp 307–322 (in French)

  • Chevalier A (1949) A quelle époque a pris naissance la protoculture au Sahara. Rev Bot Appl Agric Trop 29:418 (in French)

  • Chowdhury KA, Buth GM (1971) 4,500-year-old plant remains from Egyptian Nubia. Proc Indian Natn Sci Acad 38(1–2):55–71

    Google Scholar 

  • Ciferri R, Garavini M (1941) I cereali dell’Africa Italiana. III. I frumenti oasicoli del Sahara libico. Bibliotheca Agraria Coloniale 19, Regio Istituto Agronomico per l’Africa Italiana, Firenze (in Italian)

  • Cubadda R, Marconi E (2002) Spelt wheat. In: Belton P, Taylor J, John RN (eds) Pseudocereals and less common cereals: grain properties and utilization potential. Springer, London Ltd, pp 153–175

    Chapter  Google Scholar 

  • Damania AB, Hakim S, Moualla MY (1992) Evaluation of variation in T. dicoccum for wheat improvement in stress environment. Hereditas 116:163–166

    Article  Google Scholar 

  • Damidaux R, Autran JC, Grignac P, Feillet P (1978) Mise en évidence de relations applicables en sélection entre l’électrophorégramme des gliadines et les propriétés viscoélastiques du gluten de Triticum durum Desf. C R Acad Sci 287:701–704 (in French)

    Google Scholar 

  • Darbandi AI, Samadi BY, Boushehri AAS, Mohammadi M (2010) Allelic variations in Glu-1 and Glu3 loci of historical and modern Iranian bread wheat (Triticum aestivum L.) cultivars. J Genet 89:193–199

    Article  Google Scholar 

  • De Arana M (1934) El Instituto de cerealicultura y los nuevos tipos de trigo. Agricultura 67:437–448 (in Spanish)

    Google Scholar 

  • Desprez F (1927) Notes et observations sur les nouveaux blés “hybrides” de M. Schribaux. CR Acad d’Agric de France Paris 13:939–948 (in French)

    Google Scholar 

  • Dorofeev VF, Filatenko AA, Migushova EF, Udachin RA, Jakubziner MM (1979) Wheat. In: Dorofeev VF, Korovina ON (eds) Flora of cultivated plants, vol 1. Kolos, Leningrad, p 346

    Google Scholar 

  • Ducellier L (1909) Culture et vente des céréales en Algérie. Bull Soc Agric Alger, no 373 (in French)

  • Ducellier L (1920) Les blés du Sahara. Bibliothèque du colon de l’Afrique du Nord no 4, Alger (in French)

  • Ducellier L (1929) Céréales recueillies par le Dr. R. Maire au cours de sa mission au Hoggar. Bull Soc Hist Nat Afrique du Nord 20:221–226 (in French)

    Google Scholar 

  • Ducellier L (1930) Espèces et variétés de céréales cultivées en Algérie. In: Les céréales d’Algérie. Imprimerie Algérienne, Alger (in French)

  • Dvorak J, Luo MC (2001) Evolution of free-threshing and hulled forms of Triticum aestivum: old problems and new tools. In: Caligari PDS, Brandham PE (eds) The Linnean, special issue no 3. Wheat taxonomy: the legacy of John Percival. Academic Press, London, pp 127–136

  • Dvorak J, Luo M, Yang Z (1998) Restriction fragment length polymorphism and divergence in the genomic regions of high and low recombination in self-fertilizing and cross-fertilizing Aegilops species. Genetics 148:423–434

    CAS  PubMed Central  PubMed  Google Scholar 

  • Dvorak J, Deal KR, Luo MC, You FM, von Borstel K, Dehghani H (2012) The origin of spelt and free-threshing hexaploid wheat. J Heredity 103:426–441

    Article  CAS  Google Scholar 

  • Erroux J (1952) Les blés des oasis. Bull Soc des Agriculteurs d’Algérie no 567 (in French)

  • Erroux J (1954) Les blés du Fezzan. Bull Soc Hist Nat Afrique du Nord 45:302–317 (in French)

    Google Scholar 

  • Erroux J (1958) Note sur les blés des oasis du Sahara Algérien. Bull Soc Hist Nat Afrique du Nord no 49 (in French)

  • Erroux J (1962) Les blés des oasis Sahariennes. Mémoire no 7, Université d’Alger, Institut de Recherches Sahariennes (in French)

  • Erroux J (1991) Blé. In: Camps G (ed) Encyclopédie Berbère. Edisud, Aubenas, pp 1526–1536 (in French)

    Google Scholar 

  • Feldman M, Kislev ME (2007) Domestication of emmer wheat and evolution of free-threshing tetraploid wheat. Israel J Plant Sci 55:207–221

    Article  Google Scholar 

  • Filatenko AA, Hammer K (2014) Wheat landraces from Oman: a botanical analysis. Emir J Food Agric 26:119–136

    Google Scholar 

  • Flaksberger C (1930) Ursprungszentrum und geographische Verbreitung des Spelzes (Triticum spelta L.). Angewandte Botanik, Band 12(2):86–99 (in German)

  • Follie M (1792) Voyage dans les déserts du Sahara. Imprimerie du Cercle Social, Paris, France (in French)

  • Goncharov NP (2012) Sravnitelnaya genetika pshenitz i ikh sorodichey (Comparative genetics of wheats and their related species). Academic Publishing House Geo, Novosibirsk, Russia (in Russian with English Summary)

  • Grillot G (1948) Les blés du Maroc. La Terre Marocaine 224:173–184 (in French)

    Google Scholar 

  • Gsell S (1908) Histoire ancienne de l’Afrique du Nord, tome 1 (Les conditions du développement historique, les temps primitifs, la colonisation phénicienne et l’empire de Carthage). Hachette, Paris (in French)

    Google Scholar 

  • Guarino L, Chadja H, Mokkadem A (1991) Wheat collecting in South Algeria. Rachis 10:23–25

    Google Scholar 

  • Hammer K, Perrino P (1985) A check-list of the cultivated plants of the Ghāt oases. Die Kulturpflanze 33:269–286

    Article  Google Scholar 

  • Hammer K, Filatenko AA, Pistrick K (2011) Taxonomic remarks on Triticum L. and ×Triticosecale Wittm. Genet Resour Crop Evol 58:3–10

  • Harlan JR (1970) Evolution of cultivated plants. In: Frankel OH, Bennett E (eds) Genetic resources in plants. Their exploration and conservation. Aldine, Chicago, pp 19–32

    Google Scholar 

  • Hidore JJ, Albokhair Y (1982) Sand encroachment in Al Hasa Oasis, Saudi Arabia. Geogr Rev 72:350–356

    Article  Google Scholar 

  • Jaaska V (1978) NADP-dependent aromatic alcohol dehydrogenase in polyploid wheats. Theor Appl Genet 53:209–217

    Article  CAS  PubMed  Google Scholar 

  • Jasny N (1944) The wheats of classical antiquity. Johns Hopkins Press, Baltimore

    Google Scholar 

  • Jlibene M, Nsarellah N (2011) Wheat breeding in Morocco, a historical perspective. In: Bonjean A, Angus WJ, van Ginkel M (eds) The World Wheat Book, vol 2. A history of wheat breeding. Lavoisier Publishing, Paris, pp 425–442

    Google Scholar 

  • Kerber ER, Rowland GG (1974) Origin of the free-threshing character in hexaploid wheat. Can J Gen Cytol 16:145–154

    Google Scholar 

  • Kipfer BA (2000) Encyclopedic dictionary of archeology. Kluwer Academic Press, New York

    Book  Google Scholar 

  • Kobrehel K, Feillet P (1975) Identification of genomes and chromosomes involved in peroxydase synthesis of wheat seeds. Can J Bot 53:2336–2344

    Article  CAS  Google Scholar 

  • Lethielleux J (1943) Le Fezzan, ses jardins, ses palmiers. Publication de l’Institut des belles Lettres Arabes. Bascone et Muscat, Tunis (in French)

  • Linstadter J (2008) The Epipalaeolithic-Neolithic-Transition in the Mediterranean region of the Northwest Africa. Quartär 55:41–62

    Google Scholar 

  • Luo MC, Young ZL, Kawahara T, You F, Waines JG, Dvorak J (2007) The structure of wild and domesticated emmer wheat populations, gene flow between them, and the site of emmer domestication. Theor Appl Genet 114:947–959

    Article  PubMed  Google Scholar 

  • MacKey J (2005) Wheat, its concept, evolution and taxonomy. In: Royo C, Nachit M, Di Fonzo N, Araus JL, Pfeiffer WH, Slafer GA (eds) Durum wheat breeding: current approaches and future strategies. Howorth Press, New York, pp 3–62

    Google Scholar 

  • Maire R (1940) Etude sur la flore et la végétation du Sahara Central. Mem Soc Hist Nat Afr du Nord, vol 2. Alger (in French)

  • Mattingly DJ, Sterry M (2013) The first towns in the central Sahara. Antiquity 87:503–518

    Google Scholar 

  • Mattingly DJ, Brooks N, Cole F, Dore J, Drake N, Leone A, Hay S, McLaren S, Newson P, Parton H, Pelling R, Preston J, Reynolds T, Schrüfer-Kolb I, Thomas D, Tindall A, Townsend A, White K (2001) The Fezzan Project 2001: preliminary report on the fifth season of work. Libyan Stud 32:133–153

    Google Scholar 

  • Mattingly DJ, McLaren S, Savage E, Al-Fasatwi Y, Gadgood K (2006) The Libyan desert: natural resources and cultural heritage. Society for Libyan Studies, Monograph no 6

  • McFadden ES, Sears ER (1946) The origin of Triticum spelta and its free-threshing hexaploid relatives. J Hered 37:81–89

    PubMed  Google Scholar 

  • Mehra KL (2003) The origin, domestication and selection of crops for specific Yemeni environments. In: Al-Hakimi A, Pelat F (eds) Indigenous knowledge and sustainable agriculture in Yemen. Centre Français d’Archéologie et de Sciences Sociales de Sana’a, Les Cahiers du CEFAS, pp 9–14 (in French)

  • Miège E (1924) Sur les divers Triticum cultivés au Maroc. Bull de la Soc des Sc Nat du Maroc 4:5–6 (in French)

    Google Scholar 

  • Monneveux P, Reynolds MP, Trethowan R, González-Santoyo H, Peña RJ, Zapata F (2005) Relationship between grain yield and carbon isotope discrimination in bread wheat under four water regimes. Euro J Agron 22:231–242

    Article  CAS  Google Scholar 

  • Nesbitt M, Samuel D (1996) From staple crop to extinction? The archaeology and history of the hulled wheat. In: Padulosi S, Hammer K, Heller J (eds) Hulled wheats, promoting the conservation and used of underutilized and neglected crops. IPGRI, Rome, pp 40–99

    Google Scholar 

  • Ohtsuka I (1998) Origin of central European wheat. In: Slinkard AE (ed) 9th International Wheat Genetics Symposium. University of Saskatchewan Extension Press, Saskatoon, pp 303–305

    Google Scholar 

  • Oliveira HR, Campana MG, Jones H, Hunt HV, Leigh F, Redhouse DI, Lister DL, Jones MK (2012) Tetraploid wheat landraces in the Mediterranean Basin: taxonomy, evolution and genetic diversity. PLoS ONE 7:1–13

    Google Scholar 

  • Passager P (1957) Ouargla (Sahara Constantinois). Etude historique, géographique et médicale. Archives de l’Institut Pasteur d’Algérie 35:2 (in French)

  • Passager P, Barbançon S (1956) Taghit (Sahara Oranais). Etude historique, géographique et médicale. Archives de l’Institut Pasteur d’Algérie 34:3 (in French)

  • Payne PI, Corfield KG, Holt LM, Blackman JA (1981) Correlation between the inheritance of certain high molecular weight subunits of glutenin and bread-making quality in progenis of six crosses of bread wheat. J Sci Food Agr 32:51–60

    Article  CAS  Google Scholar 

  • Pelling R (2005) Garamantian agriculture and its significance in a wider North African context: the evidence of the plant remains from the Fazzan Project. J N Afr Stud 10:397–411

    Article  Google Scholar 

  • Perrino P (1991) Plant germplasm collecting activities in Africa of the National Research Council (CNR) of Italy. In: Attere F, Zedan H, Ng NQ, Perrino P (eds) Crop genetic resources of Africa. Proceedings of an international conference, Ibadan. Ebenezer Baylis, The Trinity Press, UK, pp 61–68

  • Perrino P, Porceddu E (1990). Wheat genetic resources in Ethiopia and the Mediterranean region. In: Srivastava JP, Damania AB (eds) Wheat genetic resources: meeting diverse needs. Wiley, Chichester, pp 161–178

  • Perrino P, Polignano GB, Porceddu E (1976) Frumenti del Nord-Africa. II. Risultati di una missione nel Nord-Ovest Algerino e nelle oasi. Annali della Facoltà di Agraria dell’Università di Bari 28:13–39 (in Italian)

    Google Scholar 

  • Perrino P, Polignano GB, Hammer K, Lehmann CO (1984) Wheat and barley collected in Libya. FAO/IPGRI Plant Genet Resourc Newslett 58:39–41

    Google Scholar 

  • Porceddu E, Perrino P (1973) Wheat in Ethiopia: preliminary report of a collection mission. Plant Genet Resourc Newslett 30:33–36

    Google Scholar 

  • Porceddu E, Perrino P, Olita G (1973) Preliminary information on an Ethiopian wheat germplasm collecting mission. In: Scarascia Mugnozza GT (ed) Proceedings of the symposium on genetics and breeding of durum wheat, Bari, Italy. University of Bari, Italy, pp 181–200

  • Renfrew JM (1969) The archaeological evidence for the domestication of plants: methods and problems. In: Uoko PJ, Dimbley GW (eds) The domestication and exploitation of plants and animals. Gerald Duckworth and Co Ltd, London

    Google Scholar 

  • Rodriguez T (1932) Notas zootecnicas. La Semana. Veterinaria 801:300–301 (in Spanish)

    Google Scholar 

  • Salamini F, Özkan H, Brandolini A, Schäfer-Pregl R, Martin W (2002) Genetics and geography of wild cereal domestication in the Near East. Nat Rev Genet 3:429–441

    CAS  PubMed  Google Scholar 

  • Schulz A (1916) Der Emmer des alten Aegyptens. Berichte der Deut Bot Gesell XXXV (in German)

  • Szabó AT, Hammer K (1996) Notes on the taxonomy of farro: Triticum monococcum, T. dicoccon and T. spelta. In: Padulosi S, Hammer K, Heller J (eds) Hulled wheats, promoting the conservation and used of underutilized and neglected crops. IPGRI, Rome, pp 2–40

    Google Scholar 

  • Talbert LE, Smith LY, Blake MK (1998) More than one origin of hexaploid wheat is indicated by sequence comparison of low-copy DNA. Genome 41:402–407

    Article  CAS  Google Scholar 

  • Tardieu F (2012) Any trait or trait-related allele can confer drought tolerance: just design the right drought scenario. J Exp Bot 63:25–31

    Article  CAS  PubMed  Google Scholar 

  • Tesfaye T, Jamal M (1982) Review of wheat breeding in Ethiopia. Eth J Agric Sci 4:11–24

    Google Scholar 

  • Tourte R (2005) Histoire de la recherche agricole en Afrique tropicale francophone. I. Aux sources de l’agriculture africaine: de la préhistoire au moyen âge. FAO, Rome (in French)

  • Toutain G (1977) Elements d’agronomie saharienne. De la recherche au développement. INRA-GRET, Paris (in French)

  • van der Veen M (1992) Garamantian agriculture: the plant remains from Zinchecra, Fezzan. Libyan Stud 23:7–39

    Google Scholar 

  • van Slageren MW (1994) Wild wheats: a monograph of Aegilops L. and Amblyopyrum (Jaub. & Spach) Eig (Poaceae). Wageningen Agricultural University, Wageningen, the Netherlands and International Center for Agricultural Research in Dry Areas (ICARDA), Aleppo, Syria

  • Vavilov NI (1964) Mirovye resursy sortov khlebnykh zlakov, zernovykh bobovykh, l’na i ikh ispolzovanie v selektzii (World resources of cereals, leguminous seed crops and flax, and their utilization in breeding). Nauka press, Moskow and Leningrad (in Russian)

  • Vavilov NI (1987) Proizhojdenie i geografia kulturnyh rasteniy (Origin and geography of cultivated plants). Nauka, Leningrad (in Russian)

    Google Scholar 

  • White K (2007) Monitoring oasis degradation using coarse resolution remotely sensed vegetation index data. In: Challenges for earth observation: scientific, technical and commercial remote sensing and photogrammetry society. The University of Nottingham, Nottingham, UK, pp 263–268

  • Wilkinson JC (1978) Problems of oasis development. School of Geography Research Paper 20, University of Oxford

  • Wilson A (2012) Saharan trade in the Roman period: short-, medium- and long-distance trade networks. Azania Archaeol Res Afr 47:409–449

    Article  Google Scholar 

  • Zaharieva M, Ayana NG, Al Hakimi A, Misra SC, Monneveux P (2010) Cultivated emmer wheat (Triticum dicoccon Schrank), an old crop with promising future: a review. Genet Resour Crop Evol 57:937–962

    Article  Google Scholar 

  • Zapata L, Peña-Chocarro L, Pérez-Jordá G, Stika HP (2004) Early neolithic agriculture in the Iberian Peninsula. J World Prehist 18:283–325

    Article  Google Scholar 

  • Zhukovsky PM (1964) Kulturnye rasteniya i ikh sorodichi (Cultivated plants and their relatives). Kolos, Leningrad (in Russian)

    Google Scholar 

Download references

Acknowledgments

The authors thank Dr. Moshe Feldman for his useful comments on the manuscript, and Dr. Miloud Hafsi for his translation in English of the meaning of vernacular names of landraces.

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Correspondence to Maria Zaharieva.

Appendix

Appendix

Index of geographical names cited in the text

Adrar: capital of Adrar Province (Algeria) based around an oasis in the Touat region, at an elevation of 258 m above sea level.

Aoulef: town in the Tidikelt region (Algeria).

Bechar: capital of Bechar Province (Algeria). Bechar lies at an elevation of 747 m, on the banks of Oued Bechar. To the East are the rocky highlands of the Djebel Bechar (1,206 m) and to the North-East the Djebel Antar (1,953 m). Main cultivated crop in Bechar are dates, vegetables, figs, cereals and almonds.

Beni-Abbes: capital of the Beni-Abbes district, on the South-West of the Bechar province (Algeria).

Biskra (Beskra): capital city of Biskra province (Algeria), located on the northern edge of the Sahara, at 87 m above sea level. It is the centre of the Zab (Ziban) group of oases south of a wide, open depression between the Aures Massif and the Tell Atlas Mountains. Darb el-Arbain: trade route running North–South between Middle Egypt and the Sudan, used from as early as the Old Kingdom of Egypt for the transport and trade of gold, ivory, spices, wheat and animals.

Djanet: oasis and capital of Djanet District, in Illizi Province, South-East Algeria. It is located 412 km South of the provincial capital, Illizi. Djanet lies at an altitude of 1,035 m in a valley carved by the Oued Idjeriou through the South-West edge of the Tassili n’Ajjer mountain range.

El Golea (or El Menia): oasis town located at the eastern border of the Grand Erg Occidental at an elevation of 380 m, capital of El Menia District, Ghardaia Province (Algeria). El Golea oasis grows many agricultural products, particularly palm trees.

Fezzan: desert at the South-West of Libya, broken by mountains, uplands, and dry river valleys (wadis) in the North. Fezzan is crossed in the North by the Ash-Shati Valley (Wadi Al Shatii) and in the West by the wadi Irawan. These two areas as well as portions of the Tibesti Mountains crossing the Chadian border and a sprinkling of oases are the only parts of the Fezzan able to support settled populations. Ergs cover most of the remaining land.

Ghardaia: oasis located within the Sahara Desert (North-Central Algeria), founded by the Mozabites (a Muslim Ibadi sect of non-Arabic Muslims) a thousand years ago in the M’Zab valley.

Ghat: oasis in the Fezzan (Libya). In historical times, Ghat was a major terminal point on the Trans-Saharan trade route.

Gourara: region of Algeria on the North of Touat that contains oases with date palms irrigated by foggaras. The largest town, Timimoun, is a little oasis, 162 km North-East of Adrar, located at the edge of the plateau of Tadmaït.

Hoggar (or Ahaggar): highland region in central Sahara (Southern Algeria), with an average altitude of more than 900 m. The highest peak is at 3,003 m (Mount Tahat). The main city nearby the Hoggar is Tamanrasset, built in a desert valley or wadi. The main oases are Silet, Ideles, Abelassa and In Amguel.

Illizi: town situated in the south-eastern part of Algeria. Also name of the sedimentary basin to the North of Tassili n’Ajjer.

In Salah (Ain Salah): oasis town in central Algeria and important trade link of the trans-Saharan caravan route. Main crops are date palms, fruits and vegetables.

Murzuk (also Murzuq): oasis and capital of the Murzuk District in the Fezzan region of South-West Libya. Murzuk lies on the northern edge of the Murzuk Desert, a desert of ergs or great sand dunes. Murzuk was an important stop on the North–South trade route across the Sahara.

Ouargla (also Wargren or Wargla): town in southern Algeria, surrounded by five small oases: Beni Thour, Sidi Khouiled, Chott, Rouissat, Said Otba and N’goussa.

Saoura: valley in South-West Algeria. The Oued Saoura is formed from the confluence of the Oued Guir and Oued Zouzfana at Igli.

Tafilalt (also Tafilet): most important oasis of the Moroccan Sahara and considered one of the largest oases in the world, the oasis is entirely located along the Ziz River. In Ifli, the central portion, formerly existed the town of Sijilmasa, founded by Miknasa Berbers in 757. It was on the direct caravan route from the Niger to Tangier.

Taghit: oasis in the Bechar Province, on the eastern bank of the Oued Zouzfana, in West Algeria.

Tamanrasset (also Tamanghasset or Tamenghest): oasis town and capital of Tamanrasset Province in South Algeria, in the Hoggar Mountains, located at an altitude of 1,320 m. Main cultivated crops are citrus fruits, apricots, dates, almonds, cereals, corn, and figs.

Tassili n’Ajjer: vast plateau in South-East Algeria at the borders of Libya and Niger. The highest point is Adrar Afao, 2,158 m, and the nearest town is Djanet, about 10 km South-West of the range.

Tell: Arabic word (height) referring to the entire Maghreb fringe ‘useful’ reliefs near the coast, in opposition to the Sahara and the High Plains steppe.

Tibesti: mountain range in the central Sahara, primarily located in the extreme North of Chad, with a small extension into southern Libya.

Tidikelt: region in the center of Sahara. Its boundaries are to the West, the Touat and the valley of Saoura, to the East, Tassili n’Ajjer, to the North the Tademait and to the South the foothills of the Mouydir and Ahnet mountains. From the North to the South can be distinguished a sandy depression that houses Saharan villages and the southern plains crossed by wadis. The main cities are In Salah and Aoulef.

Timimoun: little oasis town and capital of Timimoun District in Adrar Province, Gourara region (Algeria), located at the edge of the plateau of Tademait, at an elevation of around 288 m above sea level.

Touat (also Tuwat and Tuat): a desert region of central Algeria, located to the South of the Grand Erg Occidental, to the East of the Erg Chech and to the South-East of the Tademait Plateau. It contains a string of small oases distributed along the eastern edge of the Wadi Messaoud, a continuation of the Wadi Saoura. The oases extend over a distance of 160 km from the district of Bouda in the North to Reggane in the South. The largest town in the region is Adrar, 20 km South-East of Bouda.

Tsabit: capital of Tsabit District, Adrar Province (Algeria) located near a string of oases, at the southern end of the Gourara region, at an elevation of around 257 m above sea level.

Wadi al Shatii (also ash Shati): one of the districts of Libya in the central-western part of the country. To the West, Wadi al Shatii borders the Illizi Province of Algeria. Wadi al Shatii District is named after the depression Wadi ash-Shati.

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Zaharieva, M., Bonjean, A. & Monneveux, P. Saharan wheats: before they disappear. Genet Resour Crop Evol 61, 1065–1084 (2014). https://doi.org/10.1007/s10722-014-0122-5

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  • DOI: https://doi.org/10.1007/s10722-014-0122-5

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