Abstract
A trace fossil associated with its assumed in situ maker, a holaspid specimen of the trilobite Agraulos ceticephalus (Barrande in Notice préliminaire sur le Systême Silurien et les trilobites de Bohême, Leipzig, 1846), is reported from the middle Cambrian Buchava Formation (Drumian Stage) of the Skryje-Týřovice Basin, Czech Republic. The ichnofossil is preserved on the surface of a mudstone, behind the posterior part of the intact trilobite exoskeleton; this natural association is interpreted as mortichnia. Possible mode of life and feeding strategy of the trilobite genus Agraulos are discussed. For the association of a fodichnion with its producer preserved in situ (atop, in, or at the end of its trace fossil), the designation fodichnial association is proposed.
Kurzfassung
Aus der Buchava Formation des Skryje-Týřovice Beckens (Drumium, Tschechische Republik) wird ein mit seinem Erzeuger, einem holaspiden Agraulos ceticephalus (Barrande in Notice préliminaire sur le Systême Silurien et les trilobites de Bohême, Leipzig, 1846), in situ assoziiertes Spurenfossil bekannt gemacht. Das Spurenfossil ist auf der Schichtfläche eines Tonsteins hinter dem posterioren Teil des intakten Trilobiten-Exoskeletts erhalten; diese natürliche Assoziation wird als Mortichnium interpretiert. Die mögliche Lebens- und Ernährungsweise von Agraulos wird diskutiert. Für die Assoziation von fodichnion mit in situ erhaltenem Erzeuger (oberhalb, drin, oder am Ende der Spur) wird das Terminus ‘fodichnial association’ vorgeschlagen.
References
Barrande, J. 1846. Notice préliminaire sur le Systême Silurien et les trilobites de Bohême. vi+ 1–97. Leipzig.
Barrande, J. 1852. Systême Silurien du centre de la Bohême. lère partie. Recherches paléontologiques, vol. I. Crustacès, Trilobites, xxx+ 935 pp. Prague and Paris.
Barthel, W. 1990. Solnhofen: a study in Mesozoic palaeontology. Cambridge: Cambridge University Press.
Cherns, L., J.R. Wheeley, and L. Karis. 2006. Tunneling trilobites: habitual infaunalism in an Ordovician carbonate seafloor. Geology 34(8): 657–660.
Chlupáč, I. 1993. Geology of the barrandian. A field trip guide. Senckenberg-Buch 69. Frankfurt am Main: Verlag Waldemar Kramer.
Chlupáč, I., and R. Mikuláš. 1995. Amanitichnus omittus ichnogen. et ichnosp. nov. from the Middle Cambrian, Barrandian area, Czech Republic. Ichnos 3: 273–279.
Collette, J.H., J.W. Hagadorn, and M.A. Lacelle. 2010. Dead in their tracks–Cambrian arthropods and their traces from intertidal sandstones of Quebec and Wisconsin. Palaios 25: 475–486.
English, A.M., and L.E. Babcock. 2007. Feeding behaviour of two Ordovician trilobites inferred from trace fossils and non-biomineralised anatomy, Ohio and Kentucky, USA. Memoirs of the Association of Australasian Palaeontologists 34: 537–544.
Fatka, O. 1990. Das Kambrium von Skryje und Týřovice. In Klassische Fundstellen der Paläontologie, Band 2, ed. K.H. Weidert, 12–17. Korb: Goldschneck.
Fatka, O. 2004. Association of fossils and history of research at the Týřovice-Pod hruškou locality (Middle Cambrian, Skryje-Týřovice Basin, Barrandian area). Journal of the Czech Geological Society 49: 107–117.
Fatka, O., V. Micka, M. Szabad, V. Vokáč, and T. Vorel. 2011a. Cambrian stratigraphy of the Skryje-Týřovice Basin. Bulletin of Geosciences (in press).
Fatka, O., P. Kraft, and M. Szabad. 2011b. Wiwaxia Walcott, 1911 from the Middle Cambrian of the Barrandian area (Czech Republic). Acta Palaeontologica Polonica 56 (in press).
Fatka, O., R. Mikuláš, M. Szabad, V. Micka, and M. Valent. 2011c. Arachnostega Bertling, 1992 in Cambrian of the Barrandian area (Czech Republic). Acta Geologica Polonica (in press).
Fortey, R.A., and B.D.E. Chatterton. 1988. Classification of the trilobite suborder Asaphida. Palaeontology 31: 165–222.
Fortey, R.A., and R.M. Owens. 1999. Feeding habits in trilobites. Palaeontology 42: 429–465.
Fortey, R.A., and A. Seilacher. 1997. The trace fossil Cruziana semiplicata and the trilobite that made it. Lethaia 30: 105–112.
Geyer, G., O. Elicki, O. Fatka, and A. Zylińska. 2008. Cambrian. In The geology of central Europe, vol. 1, ed. T. McCann, 155–202. London: Geological Society of London.
Havlíček, V. 1971. Stratigraphy of the Cambrian of Central Bohemia. Sborník geologických Věd, Geologie 20: 7–52.
Hawle, I., and A.J.C. Corda. 1847. Prodrom einer Monographie der böhmischen Trilobiten. Abhandlungen der königlichen böhmischen Gesellschaft der Wissenschaften 5(5): 1–176.
Knaust, D. 2010. Remarkably preserved benthic organisms and their traces from a Middle Triassic (Muschelkalk) mud flat. Lethaia 43: 344–356.
Lu, Y.-H. 1940. On the ontogeny and phylogeny of Redlichia intermedia Lu (sp. nov.) (in Chinese with English summary). Geological Society of China Bulletin 20 (3–4): 333–342.
Maletz, J., M. Steiner, and O. Fatka. 2005. Middle Cambrian pterobranchs and the question: what is a graptolite? Lethaia 38: 73–85.
Mašek, J., J. Straka, P. Hrazdíra, P. Pálenský, P. Štěpánek, and P. Hůla. 1997. Geological and nature conservation map. Protected landscape area and biosphere reserve Křivoklátsko. Prague: Czech Geological Survey.
Mikuláš, R. 1990. The ophiuroid Taeniasteras a tracemaker of Asteriacites, Ordovician of Czechoslovakia. Ichnos 1: 133–137.
Mikuláš, R. 2000. Trace fossils from the Middle Cambrian of the Barrandian area (Central Bohemia, Czech Republic). Special Papers, 12. Prague: Czech Geological Survey.
Mikuláš, R. 2006. První nález stop po lezení trilobitů v příbramsko-jinecké pánvi (střední kambrium) [First find of trilobite crawling trace in the Příbram-Jince Basin (middle Cambrian)]. Zpravodaj České geologické společnosti 2: 20–21.
Mikuláš, R., O. Fatka, and M. Szabad. 2011. Palaeoecologic consequences of ichnofossils adjacent to slightly skeletonized body fossils, middle Cambrian of the Barrandian area, Czech Republic. Ichnos (submitted).
Osgood, R.G. 1970. Trace fossils of the Cincinnati area. Palaeontographica Americana 6: 281–444.
Ou, Q., D.G. Shu, J. Han, X.L. Zhang, Z.F. Zhang, and J.N. Liu. 2009. A juvenile redlichiid trilobite caught on the move: evidence from the Cambrian (Series 2) Chengjiang Lagerstätte, southwestern China. Palaios 24: 473–477.
Seilacher, A. 2007. Trace fossil analysis. Berlin: Springer.
Seilacher, A. 2008. Biomats, biofilms, and bioglue as preservational agents for arthropod trackways. Palaeogeography, Palaeoclimatology, Palaeoecology 270: 252–257.
Seilacher, A., and J.W. Hagadorn. 2010. Early molluscan evolution: evidence from the trace fossil record. Palaios 25: 565–575.
Šnajdr, M. 1958. Trilobiti českého středního kambria (Bohemian Middle Cambrian trilobites). Rozpravy Ústředního Ústavu Geologického 24: 1–280.
Whittington, H.B. 1988. Hypostomes and ventral cephalic sutures in Cambrian trilobites. Palaeontology 31: 577–609.
Acknowledgments
This study was supported by MSM 0021620855 (Material flow mechanisms in the upper spheres of the Earth), the Czech Science Foundation through Project No. 205/09/1521 (Feeding strategies in Cambrian to Middle Ordovician of the Barrandian Region).
Author information
Authors and Affiliations
Corresponding author
Rights and permissions
About this article
Cite this article
Fatka, O., Szabad, M. Burrowing trilobite caught in the act. Paläontol Z 85, 465–470 (2011). https://doi.org/10.1007/s12542-011-0102-4
Received:
Accepted:
Published:
Issue Date:
DOI: https://doi.org/10.1007/s12542-011-0102-4