The fluvial deposits of the Manzanares and Jarama rivers present one of the largest concentrations of lithic and faunal remains of Pleistocene sites in Europe. In the Manzanares River close to the confluence of the Jarama River, the stepped terrace system disappears and gives way to the Complex Terrace of Butarque (CTB), where the sites of Santa Elena and Oxígeno are located. Different numerical dates obtained from the visible CTB’s bottom suggest that it was deposited during the MIS 6 or even MIS 7. This paper provides the first taphonomic and palaeoecological interpretation of both collections. A total of 445 fossil elements have been recorded in Oxígeno. The most represented are cranial fragments of Elephas sp. About Santa Elena, 130 fossil elements have been recorded. The best represented are fragments of Bos/Bison sp. horns and Elephas sp. tusks. The taphonomic processes identified at both sites bring to light a complex taphonomic history. The incidence of processes related to fluvial transport is higher in Oxígeno than in Santa Elena. Besides, dismemberment cut marks have been identified exclusively on the humerus of Bos primigenius in Santa Elena. Several tooth marks from carnivores have been recorded: small pits and scores at Oxígeno as well as pits and furrowing on Bos primigenius and Megaloceros sp. bones from Santa Elena. The faunal association depict an open landscape of wet meadows with herbaceous vegetation adjacent to riparian wood habitats associated with the Manzanares alluvial plain, where hominids and predators competed to some unknown extent.
The lithic assemblages associated with the Acheulean technocomplex in the Iberian Peninsula have been recorded almost entirely in fluvial deposits where no faunal remains have been preserved, which hinders the study of an important part of the behaviour of the human groups that developed this technology in Europe.
The fluvial deposits of the rivers Manzanares and Jarama, located in the centre of the sedimentary basin of Madrid, in the Tagus Middle Basin—Province of Madrid—have sealed one of the largest concentrations of Acheulean sites in Europe, with lithic and faunal assemblages assigned to a chronological frame spanning from the Middle to the Late Pleistocene (Sesé and Soto 2000, 2002; Rubio-Jara et al. 2002, 2016; Rubio-Jara and Panera 2019; Yravedra et al. 2019b). The Acheulean of the Iberian Peninsula is characterised by a significant presence of macrotools, often knapped on large flakes, as well as by the preferential use of quartzite as a raw material, except in the Manzanares valley, where flint is virtually exclusively used in the lithic industry.
The Oxígeno and Santa Elena sites were discovered in the so-called Complex Terrace of Butarque (CTB), in the valley of the river Manzanares, chronologically framed between the MIS 7 and MIS 6. They were retrieved from archeopaleontological interventions carried out during the motoring and control of the exploitation of the Manzanares sand quarries during the 1950s and the 1960s (Rus and Querol 1981; Rubio-Jara et al. 2016). The lithic assemblages retrieved from these sites fit within the last evidence of the Acheulean technocomplex in Europe. Additionally, further evidence from these sites suggests that this technocomplex coexisted with Early Middle Palaeolithic lithic industries (Santonja et al. 2016). The coexistence from MIS 9, and perhaps from MIS 10, of Acheulean industries together with others defined as Middle Ancient Palaeolithic (European Ancient Middle Palaeolithic EAMP) has been detected in the Iberian Peninsula. These last probably evolved from a technological substratum of flake industries and cores such as those recorded in Lower Pleistocene sites in Orce (Granada), Atapuerca-Sima Elefante and TD6 in Atapuerca (Burgos) and Cueva Negra de Quípar (Murcia) (Santonja 2020). Currently, a good number of deposits with EAMP industries are known in the Iberian Peninsula with MIS 8/MIS 9 chronologies both in the open air, for example, in Cuesta de la Bajada (Teruel), Ambrona (Soria) and Solana del Zamborino (Granada), as in cave, cases of Bolomor (Valencia), Gran Dolina-Atapuerca (Burgos) and Cueva de las Grajas (Málaga), which coexist with Acheulean assemblages clearly later than MIS 8, even up to MIS 6, such as Valdocarros (Madrid) or Torralba (Soria) (Rubio-Jara et al. 2016; Santonja et al. 2016). In the Manzanares valley, where there is evidence of Middle Palaeolithic industries during the MIS 5 and probably since the end of the Middle Pleistocene (Rubio-Jara 2011), sites such as Oxígeno and Santa Elena as well as other sites located in the interfluvium of the Manzanares and Jarama rivers (Bárez et al. 2010; Baena et al. 2015), with lithic assemblages that contain Acheulean elements such as handaxes, cleavers on flake and trihedral picks associated with a high percentage of retouched tools on flake, bifaces-tool support (Boëda 2001) and a stronger levallois presence, could be the result of the spatial–temporal relationship between the Acheulean and the EAMP in the region (Santonja et al. 2016; Santonja 2020).Together with thousands of lithic pieces, hundreds of faunal remains have been found in these sites, which enables the analysis of exploitation and processing of macromammals carried out by human groups with Acheulean technology. This type of analyses has been possible only in a limited number of open-air sites in the Iberian Peninsula (Villa et al. 2005; Yravedra 2010; Yravedra et al. 2010, 2012, 2014, 2019a, b, c, 2021; Baena-Escudero et al. 2014; Panera et al. 2015; Domínguez-Rodrígo et al. 2015; Rubio-Jara et al. 2016; Pineda et al. 2017; Pineda and Saladié 2019).
The nature of the deposits where faunal remains and lithic industries have been preserved has had a direct influence on the taphonomic processes that have affected the origin, preservation and condition of both assemblages. Thus, the taphonomic analysis contributes to the understanding of the biological, physical and anthropic agents involved in their formation (Yravedra et al. 2019b). The study of the faunal remains of Oxígeno and Santa Elena presented here constitutes the first taphonomic research for both collections on the basis of an exhaustive analysis of the cortical surfaces of the bone remains. Its aim was to identify any agents that may have left traces on them and to establish whether the remains were exploited, and to what extent, by human groups with Acheulean technology (Behrensmeyer 1978; Behrensmeyer and Hill 1980; Behrensmeyer et al. 1986; Lyman 1994a, b). This study also brings to light a complex taphonomic history when trying to understand these faunal accumulations associated with Acheulean industries. Preservation of cut marks on faunal remains linked to Acheulean assemblages is not frequent. Santa Elena provides new evidence of human intervention on megafauna, as dismemberment cut marks have been identified on a humerus of Bos primigenius. Together with other sites with similar evidence, this suggests that exploitation of megafauna by hominids in these valleys during the Middle and Late Pleistocene was a recurrent strategy for obtaining meat resources (animal protein). Additionally, several carnivore tooth marks have been recorded: small pits and scores at Oxígeno, as well as pits and furrowing on Bos primigenius and Megaloceros sp. bones at Santa Elena, suggesting that carnivores also interacted with the same fauna as the hominids. Both predators, carnivores and hominids, therefore, shared the same spaces in competition for animal resources. Evidence indicates that humans had primary access to the consumption of megafauna (Yravedra et al. 2010) and that their environment exploitation patterns could have been defined by itinerant routes in a familiar landscape and by the recurrent use of specific places, for example, for the consumption of megafauna meat (Yravedra et al. 2019b). In this regard, the species identified in Oxígeno and Santa Elena reflect a mixed environment of open spaces, of wet meadows and of wooded riverside areas, associated with a fluvial environment, as is deduced from the associations of pollens, small mammals and herpetofauna identified in different sites of the river Manzanares area (Álvarez Catalán et al. 2009; Sesé et al. 2011a, b; Blain et al. 2013, 2019; Laplana et al. 2015; Gil-García et al. 2019). At certain times of the year, mainly during drier seasons, those conditions could have favoured the concentration of fauna in this environment (Panera et al. 2015; Yravedra et al. 2019b).
Geomorphological and chronological framework
The sites of Santa Elena and Oxígeno were located in SE Madrid (Villaverde), approximately 1.5 km from each other (Rus and Querol 1981; Gamazo 1982, 2002; Baena et al. 2002). The river Manzanares valley is located in the so-called Tagus Basin, in the South Sub-plateau of the Iberian Peninsula (Pérez-González 1994). The Manzanares, tributary of the river Jarama, is a low sinuosity river embedded in an asymmetrical valley in which two terrace systems can be distinguished (Pérez-González and Uribelarrea 2002). Up to 13 stepped terraces can be identified in the high and middle course (between + 4–5 and + 95 m) (Pérez-González 1994), no thicker than 6–7 m. The terraces were constructed by isostatic rising, together with blocks adjustment and lithological structured controls (Alia 1960; Pérez-González 1971, 1980, 1994; Silva et al. 1988a, b), and date from the last third of the Lower Pleistocene to the Final Late Pleistocene (Rubio-Jara and Panera 2019). The chronostratigraphy of the Manzanares valley has been established based on the sequence of terraces with geomorphological criteria that allow establishing equivalences with the terraces of the rivers of the nearby valleys (Jarama and Tagus), complemented with numerical dating and biostratigraphic criteria from the known faunal associations (Moreno et al. 2019). Thus, the highest terrace (+ 60 m) may date to the end of the Lower Pleistocene, as the Matuyama Chron (1st) limit was detected in equivalent terraces of the Jarama and Tagus rivers, or in other rivers in the interior of the peninsula such as the Arlanzón, in the Duero basin, where this limit has been set by ESR at least 0.780 Ma. However, there are no numerical dates or biostratigraphic data for the higher terraces of + 52–54 m, + 44–46 m and + 35–40 m, although they must be between MIS 19 and MIS 12, since the terrace average of + 25–30 m can be attributed from the faunal associations to the MIS 11/9. On the other hand, we have AAR and ESR numerical dates on the equivalent terrace of the Jarama that refer to this chronology and IRSL numerical dates on the equivalent terrace of the Tagus River that dates it at MIS 9 or early MIS 8. We also do not have numerical dating or biostratigraphic data for the + 18–20-m terrace, although dating has been obtained on the equivalent terrace of the Jarama River by AAR with dates between MIS 7 and MIS 5 and by ESR that date it in MIS 6. Finally, on the lower terrace of the Manzanares (+ 11–12 m), two dates have been obtained by TL (40 + 2.7 ka and 40.2 + 4.6 ka) and another two by OSL on the + 8 m terrace, which date these deposits between 23.8 and 29.6 ka (Rubio-Jara and Panera 2019). In the low stretch of the river, downstream from Villaverde, and until it meets the river Jarama, the terrace system is complex (Goy et al. 1989) due to the underlying karst of evaporitic rocks that create synsedimentary overlapped terraces and affects the terraces + 25–30 m, + 18 + 20 m and + 12–15 m. This phenomenon gives rise to a thickening of the alluvial deposits (Uribelarrea 2008) and is known as the Complex Terrace of Butarque (Goy et al. 1989). The geographic identification of the sites Santa Elena and Oxígeno (Baena et al. 2002; Gamazo 2002), together with the georeferencing of published maps (Gamazo 1982), and the overlaying of orthophotos and the MAGNA (National Geological Maps) scale 1:50,000 from Madrid and Getafe have enabled an accurate location of their geomorphological position at the CTB (Rubio-Jara and Panera 2019) (Fig. 1). The deposits of the CTB, near the mouth of the Manzanares into the Jarama River, seem to be related to successive fining cycle characteristic of meandering rivers. This pattern has been observed in different outcrops in the lower course of the river Manzanares, at PRERESA, some 9 km away from the mouth of the Manzanares River into the Jarama (Carrillo et al. 1978; Arche 1983), in Perales del Río (14 km away) (Silva 2003) and Los Estragales, also 14 km away (Pérez-González et al. 2008), where fluvial levels including channel gravel and sand deposits at the base, sealed with a layer of mud that represents flood plain facies, have been described. Most of these contain mammal fauna and lithic industry (Gaibar Puertas 1974). The CTB includes at least the T + 12–15 m and T + 18–20 m, and probably also T + 25–30 m of the river Manzanares, and in some places can reach up to 70 m in thickness (Uribelarrea 2008), although virtually only the top 15 m are accessible for visual inspection. In this stretch a timeframe ranging from MIS 7/6 to MIS 4 has been obtained with various techniques and procedures along the river Manzanares (Table 1). The series of numerical dates obtained, together with the association of micromammals identified in different sites, particularly the evolutionary stage of Microtus brecciensis, as well as the presence of Microtus arvalis (Laplana et al. 2015), suggests that the base of the CTB dates to the final MIS 8 or early MIS 7. Although this species turns up in the Middle Pleistocene in the northern area of the Iberian Peninsula, it only starts to be identified in the Manzanares and Jarama terraces records to the end of the Middle Pleistocene, as it happens in Arriaga (Sesé and López Martínez 2013) and H-02 (Laplana et al. 2015). This occurs simultaneously with a cold spell that would explain spreading to southernmost areas. This species lives currently in the northern half of the Peninsula, where it would have sheltered after the Holocene climatic improvement (Laplana et al. 2015). The TL dates are considered therefore to be a minimum age, when taking into account the AAR (Panera et al. 2011), the ESR dates (Moreno et al. 2019) and the Mediterranean warm and semi-humid climatic conditions with dry summers suggested by the study of the herpetofauna (Blain et al. 2012).
Material and methods
Although the lithic series from Oxígeno (9432 pieces) is barely published (Rus and Querol 1981) and Santa Elena (2496 pieces) is unpublished, their industries have been described as Large Flake Acheulean on flint, and possibly Middle Palaeolithic industries (Rubio-Jara et al. 2016). In this paper, a first approach to both lithic assemblages, based on the inventories preserved in the Museo Arqueológico Nacional (hereinafter MAN) and the data collected in published papers, is offered (Santonja and Querol 1980; Rus and Querol 1981).
Part of the findings retrieved from the archaeological works undertaken by different researchers in the Manzanares valley from the end of the nineteenth century and throughout the twentieth century (Quero 2002; Santonja and Vega 2002, 2008; Rubio-Jara et al. 2002) was incorporated into the MAN collections (Cacho and Martos 2002). Thus, the fauna and lithic industries of the Oxígeno and Santa Elena sites, located in fluvial deposits of the Manzanares, entered the MAN in 1973 from the archaeological works carried out at the beginning of the 1950s by Julio Martínez Santa Olalla (Rus and Querol 1981; Quero 2002) (Goy et al. 1989; Santonja and Vega 2002). The urban growth of the city of Madrid since the middle of the twentieth century has eliminated or buried a large part of the deposits from the river Manzanares, which is why the collections deposited in museums such as the MAN are a fundamental element for studying human behaviour during the Pleistocene in the Manzanares valley. Recent works have enabled the discovery of the Oxígeno and Santa Elena sites, with fauna and lithic industries that remain largely unpublished, within the sequence of Pleistocene terraces of the river Manzanares (Rubio-Jara and Panera 2019). Its georeferencing, supported by biostratigraphic contextualization based on the faunal series, has provided a chronological context that allows its integration into current models of interpretation of the Acheulean technocomplex, both from a regional perspective, in the middle Tagus Basin, and from a larger scale within the framework of the Iberian Peninsula (Santonja et al. 2001, 2011; Rubio-Jara et al. 2002, 2016; Rubio-Jara 2011; Santonja and Vega 2008; Rubio-Jara and Panera 2019).
In this paper, we have analysed a sample of 575 bone remains belonging to the Middle Pleistocene deposits of the Santa Elena (n = 130) and Oxígeno (n = 445) sites that are currently deposited in the MAN. The bone remains have been recorded according to the NISP (number of identified specimens); age patterns have also been analysed based on the tooth-wear patterns and the degree of epiphyseal fusion and tooth replacement, according to Barone (1976). After determining the species, the taphonomic analysis of the remains has been carried out using 10–20 × hand magnifiers, following the criteria of Blumenschine (1986). From the analysis of the bone surfaces, cut marks have been observed and identified according to Binford (1981), Shipman (1981) and Yravedra (2006). The identification of tooth marks has been carried out according to Binford (1981), Shipman (1981) and Blumenschine (1988). With regard to other alterations, weathering has been analysed according to Behrensmeyer (1978), taking into consideration hydric alterations, rounding and polishing of the bones, as well as chemical alterations, according to Fernández-Jalvo and Andrews (2017).
In both series, the presence of flint is dominant (over 90%). The presence of quartzite, in which large cutting tools (LCTs) have generally been configured, except for the trihedrals, which are all made from flint, is residual. The most represented technological group in both series is that of flakes and cores (representing between 80 and 85% in each series), with a token presence of flakes and levallois cores (Table 2). Among the retouched tools, the best represented are the retouched flakes and scrapers and, to a lesser extent, denticulates. Simple scrapers are better represented than transverse, convergent, bifacial and double types.
In the group of LCTs, handaxes dominate, with a significant presence of cleavers on flake, while trihedral picks are practically non-existent from Santa Elena (Fig. 2).
In Oxígeno we have the study of a sample of LCTs from the collection deposited in the MAN (219 handaxes, 38 trihedrons and 21 cleavers) (Rus and Querol 1981). The handaxes, cleavers on flake and trihedral picks analysed do not show signs of intense fluvial alteration (except for one trihedral, one cleaver on flake and approximately 10% of the handaxes, all flint, which show abraded edges). Most of the handaxes were made from flint (95%) and rarely from quartzite. The most used support is flake on flint (n = 127; 58%) and occasionally quartzite (n = 7; 5.5% of the handaxes on flake).
Thick handaxes dominate (n = 195; 89%) with lanceolate-Micoquian, amygdaloid and protolimande silhouettes (according to Bordes 1961), while among the flat handaxes cordiform, discoid and oval types stand out. In general, the series consists of handaxes with bilateral and bifacial configuration, which occasionally show retouch and edges (9 handaxes were categorized as handaxe-scraper) conformed with soft hammer (use of soft hammer was identified on one third of the sample). The trihedrals, all flint, were also knapped mainly on flake (68%) and had thick and cortical butts. Soft hammer is rarely used for knapping trihedrals (this is obvious only in two instances). The use of soft hammer on the cleavers studied is not frequent either, although they tend to show very regular and symmetrical silhouettes, with type V being the dominant one over II, 0 and I (classification Tixier 1956; Inizan et al. 1999: pp. 55–57).
The analysed faunal assemblages consist of 445 remains from Oxígeno (Fig. 3) and 130 remains from Santa Elena (Fig. 4). The following species have been identified in Oxígeno: Bison priscus, Bos primigenius, Bos/Bison sp., Cervus elaphus, Equus ferus, Equus hydruntinus, Elephas sp., Mammuthus sp., Megaloceros matritensis, Palaeoloxodon antiquus and Stephanorhinus sp. (Table 3). The best represented species are proboscideans, exemplified by Palaeoloxodon antiquus and Mammuthus sp., followed by Bos primigenius and Equus. In this faunal representation, the presence of the two types of proboscideans along with Megaloceros matritensis (Van der Made 2019) is outstanding, as this is one of the last few existing references to this species.
All the remains analysed have been ascribed to adult individuals. The following species have been identified from Santa Elena: Bos primigenius, Bos/Bison sp., Cervus sp. Equus ferus, Equus sp., Mammuthus sp., Megaloceros sp., Palaeoloxodon antiquus and Stephanorhinus hemitoechus (Table 3). Although the species representation is similar to that of Oxígeno, the aurochs slightly outnumber the proboscideans, and the horses are less represented. As in Oxígeno, all individuals were adults.
The faunal association of both sites corresponds to the Middle Pleistocene, defined, above all, by the species Bos primigenius, Megaloceros matritensis, Palaeoloxodon antiquus and Stephanorhinus hemitoechus. The fauna is the characteristic of temperate climates, where open steppe-type environments predominated and developed in intercalations of closed and open areas, as suggested by the presence of rhinoceros, horse, Equus hydruntinus and proboscideans. Skeletal profiles are highly skewed due to cranial elements predominating in all taxa, which present worse preservation conditions than the appendicular elements, since some elements such as tusks are highly fragmented. Among the postcranial elements, the lower appendicular elements stand out, such as the metapodials, which appear complete in both sites, although other long bones such as humeri are also present.
Despite the small size of the bone samples from Oxígeno (Fig. 5) and Santa Elena (Fig. 6), the incidence of various taphonomic processes has been observed at both sites. The state of bone surfaces shows regular conditions, with moderate or severe weathering, defined by the presence of cracking on the cortical surface. In Santa Elena, transverse and longitudinal marks overlapping the bone axis, with an open and superficial morphology, have been observed. This has been interpreted as weathering—the bones seem to show more severe weathering than those of Oxígeno, which would suggest a slower sedimentation process in Santa Elena. Along with weathering, other alterations have been recorded, such as trampling, which can easily be confused with cut marks. Among the different taphonomic processes that can be recorded on the bones, trampling marks are easily mistaken with cut marks because both have similar characteristics. Both are elongated marks with variable orientations and lengths, both can appear anywhere on the bone and both can be found in groups or isolated. Traditionally, cut marks were said to be V-shaped, and deeper than trampling marks can sometimes be V-shaped and can be deep. Although this is not usual, sometimes uncertain marks appear that can only be differentiated by analysing the marks in detail, paying attention to elements such as the continuity/brokenness of the line or the presence/absence of micro-striations. This topic has generated an extensive bibliography for 40 years. There are the works of Potts and Shipman (1981), Fiorillo (1984), Oliver (1984), Andrews and Cook (1985); Behrensmeyer et al. (1986) and Fisher (1995) who already raise the problem of trampling marks and how they affect the interpretation of cut marks and sites. In some cases, important debates have been generated about the correct identification of cut marks, since sometimes the authors discuss whether the marks are from cuts or trampling (Domínguez Rodrigo et al. 2011; Domínguez Rodrigo and Alcalá 2016 vs McPherron et al. 2010). Therefore, given the difficulty of correctly differentiating among cut or trampling marks, some authors have tried to generate methodological protocols that allow the two types of alterations to be correctly differentiated (Domínguez Rodrigo et al. 2009; Pineda et al. 2014; Courtenay et al. 2019, 2020).
Hydric alterations have been recorded on both sites; however, the incidence is less pronounced in Santa Elena, where episodes of rounding have not occurred. On the contrary, in Oxígeno, the water action is more pronounced, giving rise to rounding and alterations of the bone surfaces. As for alterations of biological origin, the marks of biochemical origin produced by bacteria, fungi or insects, as well as those caused by vermiculation by roots, have been observed in both sites.
Regarding alterations of anthropogenic origin, only Santa Elena has preserved cut marks on several remains specifically on a humerus of Bos primigenius. This shows the connection between the lithic industry recorded on the site to the fauna. However, the sample is small, and the scarcity of postcranial elements has prevented more cut and percussion marks from being recorded. In the case of Oxígeno, on the one hand, the poor preservation of the bone surfaces and the rounding of the bones may have masked the anthropic traces. On the other hand, both in Oxígeno and in Santa Elena, tooth marks have been recorded, which suggests that carnivores could have been important competitors to the humans during the Middle Pleistocene in the valley of the river Manzanares. The recorded tooth marks are of furrowing type on an indeterminate bone in Oxígeno, as well as pits and scores on a proximal radius of Megaloceros and on some epiphyses at Santa Elena.
The sites of Santa Elena and Oxígeno are located on fluvial deposits of the Complex Terrace of Butarque (CTB) of the river Manzanares, whose base has been chronologically framed between MIS 7 and MIS 6 (Rubio-Jara and Panera 2019). The collections, retrieved in the early 1950s (Rus and Querol 1981), have provided lithic assemblages with a very high number of elements associated with fauna, an exceptional circumstance in the whole of the Middle Pleistocene archaeo-paleontological record of the Iberian Peninsula. The materials from the Santa Elena and Oxígeno sites have remained practically unpublished and lacked a precise geomorphological context until now. Its location, based on the georeferentiation and the biostratigraphic contextualization of its faunal series, in the so-called CTB, within the sequence of the fluvial terraces of the river Manzanares, has allowed the integration of both sites into the current interpretation model of the Acheulean in the Middle Tagus Basin, in the interior of the Iberian Peninsula. In the current geochronological frame, the earliest Acheulean evidence in the region could date from sometime between MIS 15 and MIS 13. Although no excavated site has been dated earlier than MIS 13, infrequent finds of small assemblages of lithic industry have been located in some middle and high terraces of the river Tagus (Santonja and Pérez-González 1997, 2000–2001; Rodríguez de Tembleque et al. 2010). Additionally, lithic industry within a stratigraphic context and showing clear Acheulean features has been recorded in different locations in terrace levels of the river Tagus. They probably date from MIS 15/MIS 12, for example, Hornaguera and Paridera (Santonja and Pérez-González 1997; Rodríguez de Tembleque 2008), Buenavista inferior and Salchicha inferior (Alférez 1977; Santonja 1981) or Gébalo (Rodríguez de Tembleque 2008). In all cases, quartzite is the predominant material used where, together with flakes and cores, some LCTs appear, such as choppers, cleavers, trihedrals and handaxes. Puente Pino, located in an intermediate position between two middle terraces of the river Tagus, could actually be the oldest Acheulean site excavated in the Tagus Basin (Rodríguez de Tembleque 2004, 2005, 2008). It can be dated to just before the expansion of the Acheulean technocomplex in the area, around MIS 11/MIS 9, in accordance with the numerous lithic assemblages found in stratigraphic contexts and the sites excavated throughout the Tagus Basin. Sites such as Pinedo (Querol and Santonja 1979), Cañete Bajo (Santonja and Pérez-González 1997, 2000–2001; Rodríguez de Tembleque 2008), Cañete Alto (Rodríguez de Tembleque 2008), Cien Fanegas (Baena et al. 2010) in the Tagus valley, Áridos 1 and 2 (Santonja et al. 1980) in the Jarama valley, San Isidro (Santonja 1977) and Transfesa/Tafesa (Baena and Baquedano 2010) in the Manzanares valley have provided industries with significant percentages of macrotools, among which the use of large flakes for shaping bifaces is common, together with retouched tools barely elaborated and scant levallois presence. Handaxes, often made on flake, are the most common bifaces and tend to be poorly elaborated, with non-regularized or non-retouched edges, while trihedral picks are scarcely represented. Cleavers on flake (0 and I Tixier’s types mainly) are also present. The latest evidence of the Acheulean technocomplex in the Tagus Basin dates as late as the end of the Middle Pleistocene MIS 8 to MIS 6 in sites such as Las Acacias (Santonja and Querol 1977; Santonja 1981) and Valdocarros (Blain et al. 2012) in the Jarama valley, Santa Elena and Oxígeno (Rus and Querol 1981) in the Manzanares valley or Los Ahijones and El Cañaveral, found on the dividing platform along the Manzanares-Jarama river valleys (Bárez et al. 2010; Baena et al. 2015). A significant presence of more elaborately made handaxes than those of previous dates has been detected among the macrotools of these deposits, often with edges regularized by retouching, as well as complex retouched tools on flake. In Santa Elena and Oxígeno also, levallois cores and débitage have been described. These assemblages may correspond to the interrelation between Acheulean and Early Middle Palaeolithic (Santonja et al. 2016).
Therefore, the sites Santa Elena and Oxígeno fit into the last beats of the Acheulean in the Iberian Peninsula. Fluvial loads from the rivers Manzanares and Jarama have been influential in the choice of raw materials for manufacturing these industries, but not decisive. For example, in the Tagus and Jarama valleys, where there is plenty of quartzite, this raw material prevails for knapping macrotools, whereas in the Manzanares valley, flint predominates and quartzite is absent. Nevertheless, in this last valley, manufacturing of bifaces on flake is frequent, and in the Jarama valley, flint pieces are more frequent as retouched tools, despite being a rare raw material. The lithic industries of Santa Elena and Oxígeno do not show signs of intense fluvial alterations, due to the presence of flint hegemonic. This last issue is an anomaly in the Acheulean technocomplex in the Iberian Peninsula, where the macrotooling is represented mainly in quartzite. The almost exclusive use of flint in the Manzanares valley to produce macrotools, mainly from the configuration of large flakes, could initially be related to availability of the raw material in the environment, but other factors such as functionality could have played a part. The fluvial loads of the river Manzanares in the area where the sites Santa Elena and Oxígeno were located consist mainly of quartz, which has sporadic representation in the lithic assemblages of both sites, apart from flint in gravels and pebbles bars, whose diameters are seldom over 7–8 cm in length (Goy et al. 1989). The flint used in the elaboration of the macrotools of these sites could come from the Miocene outcrops located in the interfluve of the Manzanares and Jarama rivers (Rus and Enamorado 1991; Baena et al. 2015; Bárez et al. 2016; Yravedra et al. 2019b), situated some 10 km to the northeast, where several Acheulean sites associated with flint exploitation, production of large flake blanks and preconfiguration of bifacial products in contexts related to MIS 7/MIS 6 have been excavated (Baena et al. 2015; Bárez et al. 2016). In the same way, the quartzite, although residual, since it was preferably used for macrotools, would have been introduced from outside the Manzanares valley, perhaps from the Jarama River valley, to which the Manzanares is a tributary, since at the fluvial loads, quartzite is abundant. Additionally, in the sites along the river Jarama, there is a greater use of flint in retouched tools, which is actually absent in the fluvial loads (Rubio Jara et al. 2016).
The presence of all technological categories in the Santa Elena and Oxígeno assemblages points to a complete set. The chaînes opératoires of the débitage dominate the lithic assemblages, with flakes and cores better represented than retouched tools and LCTs. Handaxes with bilateral and bifacial configuration, which occasionally show retouch and edges shaped with soft hammer, and among which amygdaloidal silhouettes predominate, are outstanding in both Santa Elena and Oxígeno. Cleavers on flake, also present in both series, are mostly shaped with bifacial retouch (type V), whereas trihedrals are sporadic in the Santa Elena series. Retouched elements are more common in Oxígeno than in Santa Elena, of which scrapers are the best represented. The existence of levallois elements—flakes and cores—and the presence of handaxes with silhouettes regularized by retouch or retouched edges, containing biface-tool support (according to Boëda 2001), suggest a scenario of coexistence of Acheulean industries.
This lasted up to the end of the Middle Pleistocene, together with Early Middle Palaeolithic type industries in the middle Tagus Basin, which has already been suggested by other researchers in different regions of the Iberian Peninsula (Santonja et al. 2016; Santonja 2020; Ferreira et al. 2021).
The finds from Santa Elena and Oxígeno have not been retrieved from a systematic excavation, and therefore, the available information is limited. Nevertheless, both sites are part of the short list of Middle Pleistocene open air sites in the Iberian Peninsula which have fauna associated with lithic industries, among which sites in the Manzanares (Panera et al. 2015; Rubio-Jara and Panera 2019; Yravedra et al. 2019c) and Jarama valleys are included (Rubio-Jara et al. 2016), the Orce region (Espigares et al. 2013; Yravedra et al. 2021), and to a lesser extent the Pleistocene terraces of the river Guadalquivir (Caro et al. 2004; Baena-Escudero and Guerrero 2011; Baena-Escudero et al. 2013, 2014) in the south of the Iberian Peninsula, as well as some sites scattered in the peninsular interior, such as Torralba and Ambrona (Villa et al. 2005; Domínguez-Rodrígo et al. 2015; Rubio-Jara et al. 2016; Santonja et al. 2014b; Pineda et al. 2017; Santonja et al. 2018; Pineda and Saladié 2019). It is even rarer to record a relationship between lithic industry and fauna associated with anthropogenic alterations, in the form of green fractures, percussion or cut marks on bones, as in the case of Santa Elena. Without these signs of anthropic alteration, it is difficult to discriminate whether there is a direct relationship and what degree of association is present or if they belong to completely independent events (Bunn et al. 2010; Yravedra et al. 2014; Haynes 2022; Haynes et al. 2020). In the Iberian Peninsula, cut and percussion marks or green fractures have been recorded in different open air sites, such as Torralba and Ambrona (Shipman and Rose 1983; Villa 1990; Villa et al. 2005; Pineda and Saladié 2019), Solana del Zamborino (Díez 1992) or Pinedo (Yravedra personal observation) and also in the Middle Tagus Basin in Áridos (Santonja et al. 1980; Villa 1990; Yravedra et al. 2010), Tafesa (Yravedra 2010) and Valdocarros (Yravedra in preparation) in the Jarama valley and ETB-H02 (Yravedra et al. 2019b), PRERESA (Yravedra et al. 2012) and Arriaga (Yravedra et al. 2019b) in the Manzanares valley. Cuts and percussion marks and green fractures have also been recorded in cave sites such as Galería (Diez et al. 1999) and TD10 (Rodríguez Hidalgo et al. 2017) at the karst complex of Atapuerca, and in southern areas of the Iberian Peninsula, as in Cueva del Ángel (García Solano 2014).
A great deal of these anthropogenic alterations on bones is related to megafauna, mainly proboscideans. At the Barranc de la Boella site, located to the northeast of the Iberian Peninsula, and dated between 0.78 and 0.99 Ma, possible cut marks on two rib diaphyses of Mammuthus meridionalis have been identified (Pineda et al. 2017). In Ambrona, located in the Northern Meseta of the Iberian Peninsula, and dated to MIS 11/MIS 9, as well as in Torralba, located near Ambrona and dated to MIS 7, butchering marks produced by hominids on elephant bones (Palaeoloxodon antiquus) have been recorded, although they are rare in the whole assemblage of the recorded fauna (Shipman and Rose 1983; Villa 1990; Pineda and Saladié 2019). In the Middle Basin of the river Tagus, in Áridos 2, at the Jarama valley, MIS 11/MIS 9, cut marks (defleshing and evisceration) were identified on a scapular blade and another on the ventral side of one rib of an old Palaeoloxodon antiquus bull. They were probably made with a handaxe (Yravedra et al. 2010). In the Tafesa site, also in the Jarama valley, dated to MIS 11-MIS 12, cut marks were recorded on long bone shafts and axial fragments of Palaeoloxodon antiquus (Yravedra 2010). In the river Manzanares valley, at the CTB, dating from MIS 6 to MIS 5, cut marks and green fractures (for extraction of marrow) on Palaeoloxodon antiquus bones were identified in Arriaga IIa (Rus and Vega 1984; Panera et al. 2015), PRERESA (Yravedra et al. 2012) and also on a mammoth (Mammuthus intermedius or Mammuthus primigenius) in EDAR Culebro 1 (Yravedra et al. 2014).
Evidence of anthropic intervention on other species is scarcer. In Vallparadís, to the northeast of the Iberian Peninsula and dated to over 0.8 Ma, remains of rhinoceros and hippopotamus with cut marks have been described (Martínez et al. 2010); and in Cuesta de la Bajada, also in the northeast and dated to MIS 9-MIS 8, the same marks were recorded on equid and cervid bones (Santonja et al. 2014a; Domínguez-Rodrigo et al. 2015). In the Middle Basin of the river Tagus, cut and/or percussion marks have been recorded on the remains of rhino, large bovine, deer, horse and even beaver (for example, in Áridos 1, MIS 11-MIS 9) (Yravedra personal comment). In Tafesa, MIS 11-MIS 9, some cut and percussion marks were identified on cervid and bovine bones (Yravedra 2010). Cut marks have also been observed on horse and rhino bones in Pedro Jaro (Yravedra et al. 2019c), located in a middle terrace of the river Manzanares, dated to MIS 11-MIS 9. Some cut marks and green fractures were identified on the bones of a large bovine, probably auroch, in PRERESA, in the CTB of the Manzanares (Yravedra et al. 2012), and rare cut marks in ETH-H02 (also in the CTB) on equids, bovines and cervids associated with defleshing and disarticulation processes; also skinning marks were identified on a skull (Yravedra et al. 2019a). Now we have to add to this collection the marks from Santa Elena, where dismemberment cut marks on a humerus of Bos primigenius were identified. These are the only cut marks identified in the fauna of Santa Elena, and although no cut marks were identified in Oxígeno, human activity cannot be ruled out, as the scarcity of postcranial elements and the poor preservation of the bone surfaces may have made anthropic alterations invisible on the bones of both sites. In this sense, the large presence of remains of proboscideans in Santa Elena and Oxígeno and the well-documented exploitation of these animals in the area through green fractures and cut and percussion marks on proboscidean bones in the sites of Áridos 2 (Villa 1990; Yravedra et al. 2010), PRERESA (Yravedra et al. 2012), Arriaga (Panera et al. 2015; Yravedra et al. 2019b), EDAR Culebro 1 (Yravedra et al. 2014) and Tafesa (Yravedra 2010) would imply that similar exploitation was carried out in Santa Elena and Oxígeno. Cut marks recorded on the bones of other taxa in different sites of the Jarama and Manzanares valleys (Panera et al. 2015; Rubio-Jara et al. 2016; Yravedra et al. 2019b), such as Valdocarros (Yravedra personal observation), Tafesa (Yravedra 2010), ETB-H02 (Yravedra et al. 2019b), Pedro Jaro and also Santa Elena, where rhinoceros, aurochs, bison, deer and horse show evidence of anthropic exploitation, together with other small animals like the beaver from Áridos 1 (Yravedra et al. 2010), suggest a broader exploitation of taxa in Santa Elena and Oxígeno, such as that already recorded on the large bovine in Santa Elena. Additionally, Santa Elena and Oxígeno also provide evidence of carnivore intervention on the fauna. Examples of sites where the alternating activity of carnivores and humans on the same assemblages can be recorded are not frequent in the fossil record of the Iberian Peninsula. Scenarios of competition for carcasses have been identified in Fuente Nueva-3, a site located to the southeast of the Iberian Peninsula dated to 1.07–0.98 Ma, where hyenas (Pachycrocuta brevirostris) scavenged on the remains of the carcass after hominids had dealt with it (Espigares et al. 2013; Yravedra et al. 2021), and also, in Barranc de la Boella, in the northeast of the Iberian Peninsula, where cut and carnivore marks have been recorded on the same fauna (Pineda et al. 2017). In the Pleistocene sites of the Jarama and Manzanares valleys, in ETB-H02, some cut marks have also been recorded on the bones of equids, bovines and cervids associated with defleshing and disarticulation processes, along with others produced by small carnivores. This has been interpreted that humans and carnivores succeeded each other in different events in the occupation of the site space (Yravedra et al. 2019a). In EDAR Culebro 1, where anthropic marks have been recorded on mammoth bones, tooth marks on bones of rhinoceros were also recorded (Yravedra et al. 2019b). Similarly, in Tafesa, bones modified by small carnivores were identified within a basically anthropic fossil accumulation (Yravedra 2010). In Áridos 2, on the same elephant remains in which cut marks associated with defleshing and evisceration were recorded, tooth marks, intense furrowing on a distal epiphysis of the humerus and chewing on one rib end were caused by hyena (Yravedra et al. 2010). The sites of Santa Elena and Oxígeno provide new evidence of carnivore intervention on fauna accumulations in which there has also been human intervention. From the presence of human and carnivorous activity on the same type of prey detected in these sites, relevant conclusions are derived, since they may have conditioned the behaviour of human populations of the Middle Pleistocene. First, carnivores and humans shared the same spaces, succeeding each other in their wanderings through the landscape without interacting with each other. Second, according to the evidence recorded in some sites such as ETB-H02 (Yravedra et al. 2019a), Tafesa (Yravedra 2010), Áridos 2 (Yravedra et al. 2010), PRERESA (Yravedra et al. 2019c) and Valdocarros (Yravedra et al. in preparation), humans gained access to meat resources prior to carnivores. Third, the alternation between humans and carnivores in the same spaces indicates that the former did not remain for long periods of time in the places where they took advantage of these resources, although they probably visited them on a recurring basis as can be deduced from the variety of species represented in Oxígeno, Santa Elena (this paper), in ETB-H02 (Yravedra et al. 2019a), in Tafesa (Yravedra 2010) and in Valdocarros (Yravedra et al. in preparation).
All this evidence, considered in its regional context, suggests that Middle Pleistocene hominids carried out recurrent activity in the fluvial environments of the Manzanares and Jarama valleys. In this sense, the palaeoecological implications derived from the fossil faunas of Santa Elena and Oxígeno are also relevant. The species Bison priscus, Bos primigenius, Cervus elaphus, Equus ferus, Equus hydruntinus, Mammuthus sp., Megaloceros matritensis, Palaeoloxodon antiquus and Stephanorhinus hemitoechus are usually identified as species of open areas (steppes, tundra, meadows, etc.), although the deer is more linked to closed environments (forests). Remains of Bison priscus have been repeatedly found in meadow-type environments, in plains or river valleys; and equids are directly related to open environments, mainly grassland vegetation. Similar characteristics have been observed for Megaloceros matritensis and Stephanorhinus hemitoechus, which, unlike other species of this same genus, are identified as examples of steppes and/or tundra. Megaloceros matritensis, the last descendant of the genus Megaloceros, is a species recently identified in sites of the Manzanares valley. It is described as a browsing herbivore, probably adapted to the consumption of gypsophilous vegetation, which is predominant in the CTB environment (Van der Made 2019). Proboscideans (elephants and mammoths) are usually associated with open environments; however, the species Palaeoloxodon antiquus lived in more wooded environments and ate a diet of leaves, branches and soft vegetation (Garutt 1972, Garutt 1986a, b; Garutt and Vangengeim 1982; Pushkina 2007). Lastly, Cervus elaphus is commonly associated with denser forest type and more humid environments, although several authors have discussed whether this species could also be found in open forests or forests associated with steppes (Murie 1951; Flerov 1960). Generally, the association of bovines, equids, rhinos and proboscideans in the same group is usually related to cold events, although macroinvertebrates are a generalist group with a great capacity to adapt to different environments (Sala 1987). The presence of Mammuthus in Santa Elena and Oxígeno, and of Mammuthus primigenius in EDAR Culebro (Álvarez-Lao and García 2011), also in the CTB, would suggest cold conditions, although this probably relates to the last phase of this deposit, dated to MIS 6. The environmental variability inferred from the Santa Elena and Oxígeno faunas suggests a mixed and/or alternating environment, with a predominance of open areas of herbaceous vegetation, together with more closed and isolated areas. In the case of Oxígeno and Santa Elena, this mixed landscape may respond to the proximity of the river Manzanares, with areas of humid meadow next to a riparian forest that is nourished by the direct action of the river. This hypothesis is supported by the presence of different species that predominate in different environments or that can alternate between open and closed environments, and vice versa. This is the type of landscape suggested by the pollen analyses carried out in different points of the CTB, pine forest and local forest with Mediterranean taxa (evergreen type Quercus and Oleaceae), Mesophilic (Betula, Castanea, Corylus and evergreen type Quercus), riverside woodland (Alnus, Salix and Ulmus), several types of thicket (Ericaceae, Rosaceae and Cistaceae), herbaceous (Asteraceae, Poaceae and Chenopodiaceae) and continuous presence—although in low percentages—of aquatic taxa (Cyperaceae, Epilobium, Juncaceae, Myryophillum, Nymphaceae, Ranunculaceade and Typha) (Gil-García et al. 2019), which fit in the palaeoenvironmental demands considered typical of the species identified among the small mammals, Erinaceus europaeus, Crocidura russula, Rhinolophus ferrumequinum, Eliomys quercinus, Apodemus sp., Cricetulus (Allocricetus) bursae, Arvicola aff. sapidus, Microtus cabrerae and Microtus duodecimcostatus (Sesé et al. 2011a, b) and the herpetofauna, with toads (Pelobates cultripes, Bufo bufo, Bufo calamita), frogs (Pelophylax perezi), turtles, lacertid (Timon lepidus, Psamamodromus cf. algirus), scincid (Chalcides striatus, Chalcides sp.), lizards and snakes (Natrix maura, Coronella girondica) (Blain et al. 2012, 2013, 2019). A mostly open landscape was dominated by wet meadows with herbaceous or shrubby vegetation, interspersed with drier and shrubby areas, and also some forest; a mixed habitat, mainly open and partially wooded (riverside forests), was favoured by humid conditions due to its proximity to a river but also typical of a Mediterranean-type climate with alternating periods of aridity during the summer until the beginning of the autumn, identical to or even more temperate than the current one, and with abundant rains in winter and spring, characterised by less continentality and lower winter temperatures than nowadays. In this scenario, the riverside ecosystems, such as that of the Manzanares valley, could have played an important role, not only for the development of plants that could provide food for the Pleistocene hominids, but also because during the dry summer months, they could have acted as an ecological corridor in a landscape where the saline substrate would have limited the development of plant species (Panera et al. 2015), although we have pointed out that Megaloceros matritensis present in Santa Elena and Oxígeno shows a dentition adapted to xerophytic vegetation. The existence of these corridors would favour the concentration of fauna along the riverbank of the Manzanares and would facilitate hominids access to hunt or scavenge the fauna recorded from the sites.
This study reveals the great potential of the finds preserved in museums such as the MAN, which come from excavations not carried out with current scientific parameters. Despite these limitations, they contribute to the knowledge of the behavioural dynamics of hominids during the Pleistocene, especially if, as in the case of Santa Elena and Oxígeno, they come from deposits that have already disappeared or are inaccessible due to the urban development of the city of Madrid. Santa Elena and Oxígeno add to the rare group of sites with lithic industries associated with Middle Pleistocene fauna. The presence of some cut and other marks related to the activity of carnivores on the fauna of Santa Elena and Oxígeno allows both collections to be related to the activities of hominids and carnivores, with some degree of competition, and in the same landscape. The existence of this type of alternation carried out by hominids and carnivores in different sites of the valleys of the Manzanares and Jarama reinforces the theory that the consumption of megafauna (proboscideans, rhinoceros, large bovines) was more than a marginal strategy of protein obtainment by hominids, and that it was a frequent strategy over time, maintained throughout the Middle and Late Middle Pleistocene, with carnivores playing a secondary role in accessing carcasses previously exploited by hominids. The association of taxa defined in Santa Elena and Oxígeno, together with the pollen data and the palaeoecological requirements of small mammals and herpetofauna collected in different sites of the CTB, suggests a mixed landscape of open spaces of more or less humid meadows and wooded areas of riparian forest within a Mediterranean climate similar to the current one with variation of temperatures and a seasonal rainfall regime. These conditions and the geological characteristics of the Manzanares valley in the surroundings of the Santa Elena and Oxígeno sites could have conditioned the alluvial plain to become an ecological corridor that favoured the concentration of fauna and then exploited by the hominids. Then again, the synsedimentary process that affected the formation of the CTB, and gave rise to the thickening of the terrace, could have preserved fauna and industries from different times (from the Late Middle Pleistocene to the Early Late Pleistocene) together in the same deposits. It is in this sense that the presence in both sites of remains of Mammuthus sp. together with Palaeoloxodon antiquus and lithic industries attributed to the Acheulean and Early Middle Palaeolithic technocomplexes should be assessed.
Alférez F (1977) Estudio del sistema de terrazas del río Tajo al W de Toledo. Estud Geol 33:223–250
Alia M (1960) Sobre la tectónica profunda de la fosa del Tajo. Notas y Comunicaciones Del Instituto Tecnológico y Geominero De España 58:125–162
Álvarez Catalán V, de los Arcos Fernández S, Gallego Lletjos N, Gil Ortiz C, González García I, Herráez Igualador E, Ruiz Zapata B, Yravedra J (2009) Yacimiento Paleolítico del Estanque de Tormentas de Butarque. In: Santonja M (eds) Actas de las Cuartas Jornadas de Patrimonio Arqueológico en la Comunidad de Madrid. Los primeros pobladores: Arqueología del Pleistoceno, Alcalá de Henares, pp 333–337
Álvarez Lao DJ, García N (2011) Southern dispersal and Palaeoecological implications of woolly (Coelodonta antiquitatis): review of the Iberian occurrences. Quatern Sci Rev 30:2002–2017. https://doi.org/10.1016/j.quascirev.2011.05.005
Andrews P, Cook J (1985) Natural modifications to bones in a temperate setting. Man 20(4):675–691
Arche A (1983) Coarse-grained meander lobe deposits in the Jarama River, Madrid. Spain. In: Collinson J.D, Lewin J.L (eds) Meander and ancient fluvial systems. International Association Sedimentology Special Publication 6, pp 313–321
Baena J, Baquedano I (2010) Las huellas de nuestro pasado. Estudio del yacimiento de pleistoceno madrileño de Tafesa (Antigua Transfesa). Zona Arqueológica 14. Museo Arqueológico Regional. Alcalá de Henares.
Baena J, Conde Ruiz C, Gamazo M, Sesé C, Soto E (2002) Repertorio de yacimientos paleolíticos del Manzanares y del Jarama. In: Panera J, Rubio-Jara S (eds) Bifaces y elefantes. La investigación del Paleolítico Inferior en Madrid. Museo Arqueológico Regional de la Comunidad de Madrid. Zona Arqueológica 1, pp 461–491
Baena J, Ortiz I, Torres C, Bárez S (2015) Recycling in abundance: re-use and recycling processes in the Lower and Middle Paleolithic contexts of the central Iberian Peninsula. Quatern Int 361:142–154. https://doi.org/10.1016/j.quaint.2014.07.007
Baena J, Rus I, Cuartero F, Martín D, Rubio D, Roca M (2010) Estudio tecnológico de la industria lítica del yacimiento de Las Cien Fanegas (Madrid) en el contexto Achelense de la Meseta. Cuaternario y Arqueología: Homenaje a Francisco Giles Pacheco. Diputación Provincial de Cádiz, Cádiz: 71–83
Baena-Escudero R, Fernández-Caro J.J, Guerrero I, Posada J.C (2014) La Terraza Compleja del río Guadalquivir en “Las Jarillas” (La Rinconada, Sevilla. SW de España): cronoestratigrafía, industria lítica y macro-fauna asociada. Cuaternario y Geomorfología 28, 3–4: 107–125
Baena-Escudero R, Guerrero I (2011) Aproximación a la magnetoestratigrafía de la última terraza media del Guadalquivir (T12) en “Las Jarillas” (La Rinconada, Sevilla). Resúmenes XIII Reunión Nacional de Cuaternario, Andorra 2011. El Cuaternario En España y Áreas Afines, Avances En 2011:191–195
Baena-Escudero R, Guerrero I, Fernández-Caro J.J, Espigares M.P, Ros S, Posada J.C (2013) Restos de macromamíferos en la terraza T12 del Guadalquivir en la Rinconada (Sevilla): presencia de Hippopotamus amphibius Linneo, 1758. In: Recorridos cuaternarios: terrazas del Guadalquivir en el sector Carmona-Sevilla y paleosismología de terremotos en Baelo-Claudia (Tarifa, Cádiz). VIII Reunión de Cuaternario Ibérico, Sevilla: AEQUA, GTPEQ, pp 278–284
Bárez S, Baena J, Pérez-González A, Torres C, Pérez IR, de Miguel JV (2016) Acheulian flint quarries in the Madrid Tertiary basin, central Iberian Peninsula: first data obtained from geoarchaeological studies. Quatern Int 411:329–348. https://doi.org/10.1016/j.quaint.2016.01.041
Bárez S, Rus I, Pérez-González A, Vega de Miguel J (2010) Los yacimientos achelenses de “Los Ahijones”, metodología geoarqueológica y resultados preliminares de la intervención. Actas de las V jornadas de patrimonio arqueológico de la Comunidad de Madrid. Museo Arqueológico Regional de la Comunidad de Madrid 12–14 noviembre 2008. Alcalá de Henares, Madrid: 173–189
Barone R (1976) Anatomie comparée des mammifères domestiques. Tome I: Ostéologie (2 fascicules). Vigot. Paris
Behrensmeyer AK (1978) Taphonomic and ecologic information from bone weathering. Paleobiology 4(2):150–162
Behrensmeyer AK, Hill AP (1980) Fossils in the making: vertebrate taphonomy and paleoecology. University of Chicago Press, Chicago
Behrensmeyer AK, Gordon KD, Yanagi GT (1986) Trampling as a cause of bone surface damage and pseudo-cutmarks. Nature 319:768–771
Binford LR (1981) Bones: ancient men and modern myths. Academic Press Inc, New York
Blain HA, Bisbal-Chinesta JF, Martínez-Monzón A, Panera J, Rubio-Jara S, Uribelarrea D, Yravedra J, Pérez-González A (2019) The Middle to Late Pleistocene herpetofaunal assemblages from the Jarama and Manzanares valleys (Madrid, central Spain): an ecological synthesis. Quatern Int 520:49–63. https://doi.org/10.1016/j.quaint.2018.03.004
Blain H.A, Sesé C, Panera J, Rubio-Jara S, Uribelarrea D, Pérez-González A (2012) Paleoclimatic and paleoenvironmental proxies to the Marine Isotope Stage 7e (Middle Pleistocene) in central Spain (Valdocarros II, Madrid) by means of the small-vertebrate assemblages. In International Conference. European Middle Palaeolithic during MIS 8-MIS 3. Cultures-environment-chronology. Wolbrom, Poland. September 25th-28th: 77–78. http://hdl.handle.net/10261/66620
Blain HA, Sesé C, Panera J, Rubio-Jara S, Uribelarrea D, Pérez-González A (2013) Reconstitution paléoenvironnementale et paléoclimatique du Pléistocène supérieur ancien (MIS 5a) dans le centre de l’Espagne: les petits vertébrés (amphibia, reptilia & mammalia) des gisements de Hat et PRERESA (sud-est de Madrid). Quaternaire 24(2):191–205. https://doi.org/10.4000/quaternaire.6508
Blumenschine R.J (1986) Early hominid scavenging opportunities. Implications of carcass availability in the Serengeti and Ngorongoro ecosystems. BAR International Series 283
Blumenschine RJ (1988) An experimental model of the timing of hominid and carnivore influence on archaeological bone assemblages. J Archaeol Sci 15:483–502. https://doi.org/10.1016/0305-4403(88)90078-7
Boëda E (2001) Determination des unités tecno-fonctinelles de pièces bifaciales provenant de la couche acheuléenne C03 Base du site de Barbas I. In: Cliquet D (ed) Les industries à outils bifaciaux du Paléolithique moyen d’Europe occidentale. ERAUL 98: 51–75
Bordes F (1961) Typologie du Paléolithique ancien et moyen. Imprimeries Delmas, Bordeaux
Bunn HT, Mabulla AZP, Domínguez-Rodrigo M, Ashley GM, Barba R, Diez-Martín F, Remer K, Yravedra J, Baquedano E (2010) Was FLK North levels 1–2 a classic “living floor” of Oldowan hominins or a taphonomically complex palimpsest dominated by large carnivore feeding behavior? Quatern Res 74(3):355–362. https://doi.org/10.1016/j.yqres.2010.06.004
Cacho C, Martos J.A (2002) Colecciones paleolíticas de Madrid en el Museo Arqueológico Nacional. In: Panera J, Rubio-Jara S (eds) Bifaces y elefantes. La investigación del Paleolítico Inferior en Madrid. Museo Arqueológico Regional de la Comunidad de Madrid. Zona Arqueológica 1, pp 382–407
Caro J.A, Díaz del Olmo F, Risquez J (2004) Las industrias paleolíticas del yacimiento de Tarazona (La Rinconada, Sevilla): Achelense y Musteriense clásico en las terrazas medias del Guadalquivir. XI Reunión Nacional de Cuaternario, Oviedo (Asturias), 2–4 julio 2003: 175–182
Carrillo L, Gisbert J, Arche A (1978) Modelo de sedimentación de la terraza baja (+18-20 m) del río Manzanares. Estud Geol 34:549–552
Courtenay LA, Yravedra J, Huguet R, Ollé A, Aramendi J, Maté-González MÁ, González-Aguilera D (2019) New taphonomic advances in 3D digital microscopy: a morphological characterisation of trampling marks. Quatern Int 517:55–66
Courtenay LA, Huguet R, Gonzalez Aguilera D, Yravedra J (2020) A hybrid geometric morphometric deep learning approach for cut and trampling mark classification. Appl Sci 10(1):150. https://doi.org/10.3390/app10010150
Díez C (1992) Zooarqueología de Atapuerca (Burgos), e implicaciones paleoeconómicas del estudio tafonómico de yacimientos del Pleistoceno Medio. Universidad Complutense de Madrid. Doctoral Thesis
Díez C, Moreno V, Rodríguez J, Rosell J, Cáceres I (1999) Estudio arqueozoológico de los restos de macrovertebrados de la Unidad III de Galería. In: Carbonell E, Rosas González A, Díez J.C (eds) Atapuerca. Ocupaciones humanas y paleoecológicas del Yacimiento de Galería. Arqueología de Castilla y León, pp 265–281
Domínguez-Alonso R.M, De los Arcos Fernández S, Ruiz Zapata B, Gil García M.J (2009) Nuevos datos sobre la Terraza Compleja de Butarque en Villaverde Bajo. IV Jornadas sobre Patrimonio Arqueológico de la Comunidad de Madrid. Celebradas en el Museo Arqueológico Regional de la Comunidad de Madrid en 2007. Alcalá de Henares. Madrid: 339–343. ISBN 978–84–451–3259–3
Domínguez-Rodrigo M, de Juana S, Galán AB, Rodríguez M (2009) A new protocol to differentiate trampling marks from butchery marks. J Archaeol Sci 36(12):2643–2654. https://doi.org/10.1016/j.jas.2009.07.017
Domínguez-Rodrigo M, Pickering TR, Bunn HT (2011) Experimental study of cut marks made with rocks unmodified by human flaking and its bearing on claims of ∼3.4-million-year-old butchery evidence from Dikika. Ethiopia Journal of Archaeological Science 39:205–214. https://doi.org/10.1016/j.jas.2011.03.010
Domínguez-Rodrigo M, Barba R, Soto E, Sesé E, Santonja M, Pérez-González A, Yravedra J, Galán AB (2015) Another window to the subsistence of Middle Pleistocene himinins in Europe: a taphonomic study of Cuesta de la Bajada (Teruel, Spain). Quatern Sci Rev 126:67–95. https://doi.org/10.1016/j.quascirev.2015.08.020
Domínguez-Rodrigo M, Alcalá L (2016) 3.3-Million-Year-Old stone tools and butchery traces? More evidence needed. PaleoAnthropology. https://doi.org/10.4207/PA.2016.ART99
Espigares MP, Martínez-Navarro B, Palmquist P, Ros-Montoya S, Toro I, Agustí J, Sala R (2013) Homo vs Pachycrocuta: earliest evidence of competition for an elephant carcass between scavengers at Fuente Nueva-3 (Orce, Spain). Quatern Int 295:113–125. https://doi.org/10.1016/j.quaint.2012.09.032
Fernández-Jalvo Y, Andrews P (2017) Atlas of taphonomic identifications. Springer, Vertebrate Paleobiology and Paleoanthropology Series
Ferreira C, Cunha-Ribeiro J.P, Méndez-Quintas E (2021) O tecno-complexo Acheulense em Portugal: contribuiçao para um balanço dos conhecimientos. Ophiussa 5: 5–29. https://doi.org/10.51679/ophiussa.2021.80
Fiorillo A.R (1984) An experimental study of trampling: implications for the fossil record. In: Bonnichsen R, Sorg M.H (eds) Bone Modification. University of Maine Press, Maine, pp 73–98
Fisher J.W (1995) Bone surface modification in zooarchaeology. Journal of Archaeological Method and Theory2 (1): pp 7–68
Flerov K (1960) Musk deer and deer (Fauna of USSR, Mammals, Vol. I, No. 2). Jerusalem (trans. by A. Biron & Z. Cole).
Gaibar Puertas C (1974) Descubrimiento de la terraza wüermiense en la margen izquierda del río Manzanares: aportaciones paleoclimáticas, nuevos restos y testimonios del madrileño hombre prehistórico y protohistórico. Estudios Geológicos XXX 2–3:235–252
Gamazo M (1982) Prospecciones en las terrazas de la margen derecha del río Manzanares (Getafe y Rivas-Vaciamadrid). Noticiario Arqueológico Hispánico 14:7–148
Gamazo M (2002) Las colecciones paleolíticas del Manzanares y del Jarama del Museo de San Isidro. In: Panera J, Rubio-Jara S (eds) Bifaces y elefantes. La investigación del Paleolítico Inferior en Madrid. Museo Arqueológico Regional de la Comunidad de Madrid. Zona Arqueológica 1, pp 358–381
García Solano J.A (2014) La persistencia de las estrategias de subsistencia de los grupos humanos del Pleistoceno medio ibérico a partir del registro fósil de la cueva del Ángel (Lucena, Córdoba). Universidad de Granada. Doctoral Thesis
Garutt V.E (1972) On a finding of the Palaeoloxodon elephant in Eastern Urals. In: Yakhimovich V.L (ed) Questions of Stratigraphy and Correlation of the Pliocene and Pleistocene Deposits of Northern and Southern Parts of Eastern Urals. AN SSSR, Geology Institute, Ufa, Bashkirija, Russia, pp 19–26
Garutt V.E (1986a) Origin of elephants, Elephantidae and their phylogeny. In: Garrutt V.E (ed) Mammals of the Quaternary Fauna of the USSR. Proceedings of the Zoological Institute, 149, pp 15–32
Garutt VE (1986b) The origin of elephants and ways of their phylogeny. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR 149:15–32
Garutt V.E, Vangengeim E.A (1982) Order Proboscidea. In: Shantser E.V (ed) Stratigraphy of the USSR, Quaternary System, Nedra, Moscow, Russia, pp. 311–317
Gil-García MJ, Ruiz-Zapata MA, Rubio-Jara S, Panera J (2019) Landscape evolution during the Middle and Late Pleistocene in the Madrid basin (Spain): vegetation dynamics and human activity in the Jarama-Manzanares Rivers (Madrid) during the Pleistocene. Quatern Int 520:39–48. https://doi.org/10.1016/j.quaint.2018.02.034
Goy JL, Pérez-González A, Zazo C (1989) Cartografía y Memoria del Cuaternario y Geomorfología, Hoja de Madrid (559). Mapa Geológico de España. Escala 1:50.000. 2ª Serie (MAGNA). IGME. Servicio de Publicaciones del Ministerio de Industria. Madrid
Haynes G (2022) Late Quaternary proboscidean sites in Africa and Eurasia with possible or probable evidence for hominin involvement. Quaternary 5(1):18. https://doi.org/10.3390/quat5010018
Haynes G, Krasinski K, Wojtal P (2020) Elephant bone breakage and surface marks made by trampling elephants: Implications for interpretations of marked and broken Mammuthus spp. bones. Journal of Archaeology Science, 33: 102491. https://doi.org/10.1016/j.jasrep.2020.102491
Inizan ML, Reduron-Ballinger M, Roche H, Tixier J (1999) Technology and terminology of knapped stone (Préhistoire de la Pierre taillé 5). Nanterre: Cercle de Recherches et d’Études Préhistoriques: 55–57
Laplana C, Herráez E, Yravedra J, Bárez S, Rubio-Jara S, Panera J, Pérez-González A (2015) Biocronología de la Terraza Compleja de Butarque en el río Manzanares en el Estanque de Tormentas al sur de Madrid. Estud Geol 71(1):e028. https://doi.org/10.3989/egeol.41808.338
Lyman RL (1994a) Chapter 8: butchering, bone fracturing and bone tools. In: Lyman R.L (ed) Vertebrate Taphonomy. Cambridge Manuals in Archaeology, pp 294–353
Lyman RL (1994b) Chapter 9: other biostratinomic factors. In: Lyman R.L (ed) Vertebrat Taphonomy. Cambridge Manuals in Archaeology, pp 354–403
Manzano L, Expósito A, Pérez-González A, Soto E, Sesé C, Yravedra J, Ruiz Zapata B, Millán A, Benéitez P, Torres T, Mondéjar J.A, Zarco E, Sánchez H, Citores A, Ramos M, Rodríguez A (2010) El yacimiento arqueopaleontológico de E.D.A.R. CULEBRO 1 (Estación Depuradora de Aguas Residuales de la Cuenca Baja del Arroyo Culebro. Ministerio de Medio Ambiente. Confederación Hidrográfica del Tajo. Actas de las V jornadas de patrimonio arqueológico en la Comunidad de Madrid. Celebradas en el Museo Arqueológico Regional de la Comunidad de Madrid del 12–14 de noviembre de 2008. Alcalá de Henares. Madrid: 203–214. http://hdl.handle.net/10261/40992
Martínez K, García J, Carbonell E, Agustí J, Bahain JJ, Blain HA, Burjachs F, Cáceres I, Duval M, Falguères C, Garcés M, Gómez M, Huguet R (2010) A new Lower Pleistocene archaeological site in Europe (Vallparadís, Barcelona, Spain). Proc Natl Acad Sci USA 107(3):5762–5767. https://doi.org/10.1073/pnas.091385610
McPherron S, Alemseged Z, Marean CW, Wynn JG, Reed D, Geraads D, Bobe R, Béarat HA (2010) Evidence for stone-tool-assisted consumption of animal tissues before 3.39 million years ago at Dikika. Ethiopia Nature 466:857–860. https://doi.org/10.1038/nature09248
Moreno D, Duval M, Rubio-Jara S, Panera J, Bahain JJ, Shao Q, Pérez-González A, Falguères C (2019) ESR dating of several Middle to Late Pleistocene archaeopaleontological sites from the Manzanares and Jarama river valleys (Madrid basin, Spain). Quatern Int 520:23–38. https://doi.org/10.1016/j.quaint.2017.09.003
Murie O (1951) The Elk of North America. D. C, Washington
Oliver JS (1984) Analogues and site context: bone damages from shield trap cave (24CB91), carbon county, Montana, USA. In: Bonnischen R, Sorg M.H (eds) Bone Modification. University of Maine Press, Maine, pp. 61–72
Panera J, Rubio-Jara S, Yravedra J, Blain HA, Sesé C, Pérez-González A (2015) Manzanares valley (Madrid, Spain): a good country for proboscideans and Neanderthals. Quatern Int 326–327:329–343. https://doi.org/10.1016/j.quaint.2013.09.009
Panera J, Torres T, Pérez-González A, Ortiz JE, Rubio-Jara S, Uribelarrea D (2011) Geocronología de la Terraza Compleja de Arganda en el valle del río Jarama (Madrid, España). Estud Geol 67(2):495–504. https://doi.org/10.3989/egeol.40550.204
Pérez-González A (1971) Estudio de los procesos de hundimiento en el valle del río Jarama y sus terrazas (nota preliminar). Estudios Geológicos XXVII 4:317–324
Pérez-González A (1980) Geología y estratigrafía de los yacimientos de Áridos en la llanura aluvial de Arganda (Madrid). In: Santonja M, López-Martínez N, Pérez-González A (eds) Ocupaciones achelenses en el valle del Jarama. Arqueología y Paleontología I. Diputación Provincial de Madrid, pp 49–61
Pérez-González A (1994) Depresión del Tajo. In: Gutierrez E (ed) Geomorfología de España. Editorial Rueda, Madrid, pp 389–436
Pérez-González A, Rubio-Jara S, Panera J, Uribelarrea D (2008) Geocronología de la sucesión arqueoestratigráfica de Los Estragales en la Terraza Compleja de Butarque (Valle del río Manzanares, Madrid). Geogaceta, 45: 39–42. http://hdl.handle.net/10272/8190
Pérez-González A, Uribelarrea D (2002) Geología del Cuaternario de los valles fluviales del Jarama y Manzanares en las proximidades de Madrid. In: Panera, J, Rubio-Jara S (eds) Bifaces y elefantes. La investigación del Paleolítico Inferior en Madrid. Museo Arqueológico Regional de la Comunidad de Madrid. Zona Arqueológica 1, pp 303–317
Pineda A, Saladié P, Vergés JM, Huguet R, Cáceres I, Vallverdú J (2014) Trampling versus cut marks on chemically altered surfaces: an experimental approach and archaeological application at the Barranc de la Boella site (La Canonja, Tarragona, Spain). J Archaeol Sci 50:84–93
Pineda A, Saladié P (2019) The Middle Pleistocene site of Torralba (Soria, Spain): a taphonomic view of the Marquis of Cerralbo and Howell faunal collections. Anthropological and Archaeological Sciences 11:2539–2556. https://doi.org/10.1007/s12520-018-0686-7
Pineda A, Saladié P, Huguet R, Cáceres I, Rosas A, Estalrrich A, García-Tabernero A, Vallverdú J (2017) Changing competition dynamics among predators at the late Early Pleistocene site Barranc de la Boella (Tarragona, Spain). Palaeogeography Paleoclimatology Palaeocology 477:10–26. https://doi.org/10.1016/j.palaeo.2017.03.030
Potts R, Shipman P (1981) Cutmarks made by stone tools on bones from Olduvai Gorge, Tanzania. Nature 291:577–1981
Pushkina D (2007) The Pleistocene Easternmost Distribution in Eurasia of the Species Associated with the Eemian Palaeoloxodon Antiquus Assemblage 37(3):224–245. https://doi.org/10.1111/j.1365-2907.2007.00109.x
Quero S (2002) Investigación del Paleolítico en Madrid durante el franquismo (1936–1971). In: Panera J, Rubio-Jara S (eds) Bifaces y elefantes. La investigación del Paleolítico Inferior en Madrid. Museo Arqueológico Regional de la Comunidad de Madrid. Zona Arqueológica 1, pp 168–193
Querol MA, Santonja M (1979) El yacimiento achelense de Pinedo (Toledo). Excavaciones Arqueológicas en España 106. Ministerio de Cultura. Madrid: 181
Rodríguez de Tembleque JM (2004) Yacimiento de Puente Pino: Nuevas perspectivas para el estudio del Paleolítico inferior en la Península Ibérica. In: Baquedano E, Rubio-Jara S (eds) Miscelánea en Homenaje a Emiliano Aguirre, IV. Zona Arqueológica 4, pp 441–450
Rodríguez de Tembleque J.M (2005) El Paleolítico inferior en la cuenca del Tajo (Castilla-La Mancha). In: Rodríguez de Tembleque J.M (ed) Los Primeros Pobladores de Castilla-La Mancha. Fundación Cultura y Deporte de JCCM y Caja de Castilla-La Mancha, pp 112–139
Rodríguez de Tembleque J.M (2008) Primeras ocupaciones humana en la Meseta española: Estudio geoarqueológico de depósitos fluviales en la cuenca media del Tajo. Universidad Autónoma de Madrid, Madrid (España). Servicio de Publicaciones de la UAM. (Doctoral Thesis): 843
Rodríguez de Tembleque JM, Pérez-González A, Santoja M (2010) Indicios de presencia humana en la Meseta Ibérica durante la formación de la Terraza de +75–80 m del Río Tajo. In: Rodríguez de Tembleque J.M, Pérez-González A, Santoja M (eds) Cuaternario y Arqueología: Homenaje a Francisco Giles Pacheco. Asociación Profesional del Patrimonio Histórico-Arqueológico de Cádiz. Servicio de Publicaciones de la Diputación Provincial de Cádiz, pp 45–54
Rodríguez Hidalgo A, Saladié P, Ollé A, Arsuaga JL, Bermúdez de Castro JM, Carbonell E (2017) Human predatory behavior and the social implications of communal hunting based on evidence from the TD10.2 bison bone bed at Gran Dolina (Atapuerca). J Hum Evol 105:89–122. https://doi.org/10.1016/j.jhevol.2017.01.007
Rubio-Jara S (2011) El paleolítico en el valle del río Manzanares (Madrid). Caracterización geoarqueológica de depósitos pleistocenos y estudio tecnoeconómico de la industria lítica. Universidad Nacional de Educación a Distancia. (Unpublished Doctoral Thesis).
Rubio-Jara S, Panera J (2019) Unravelling an essential archive for the European Pleistocene. The human occupation in the Manzanares valley (Madrid, Spain) throughout nearly 800,000 years. Quatern Int 520:5–22. https://doi.org/10.1016/j.quaint.2018.08.007
Rubio-Jara S, Panera J, Martos J.A, Santonja M, Pérez-González A (2002) Revisión crítica y síntesis del Paleolítico de los valles del Manzanares y Jarama. In: Panera J, Rubio-Jara S (eds) Bifaces y elefantes. La investigación del Paleolítico Inferior en Madrid. Museo Arqueológico Regional de la Comunidad de Madrid. Zona Arqueológica 1, pp. 338–355
Rubio-Jara S, Panera J, Rodríguez de Tembleque J, Santonja M, Pérez-González A (2016) Large flake Acheulean in the middle of Tagus basin (Spain): middle stretch of the river Tagus valley and lower stretches of the rivers Jarama and Manzanares valleys. Quatern Int 411:349–366. https://doi.org/10.1016/j.quaint.2015.12.023
Rus I, Enamorado J (1991) Flint supply in the Manzanares valley: the acheulian site of Arriaga (Madrid, Spain). VI Flint International Symposium (Granada): 569–576
Rus I, Querol MA (1981) El arenero de Oxígeno: bifaces, hendedores y triedros conservados en el Museo Arqueológico Nacional. Trab Prehist 38:39–67
Rus I, Vega LG (1984) El yacimiento de Arriaga II: problemas de una definición actual de los suelos de ocupación. Primeras Jornadas de Metodología e Investigación Prehistórica (Soria, 1981). Ministerio de Cultura: 387–404
Sala B (1987) Bison schoetensacki Freud. from Isernia la Pineta (early Mid-Pleistocene-Italy) and revision of the european species of bison. Palaeontographia Italica Pisa 74:113–170
Santonja M (1977) Los bifaces del Cerro de San Isidro conservados en el Museo Arqueológico Nacional. Revista De Archivos, Bibliotecas y Museos 80(1):147–182
Santonja M (1981) El Paleolítico inferior de la Meseta central española. Universidad Complutense de Madrid (Doctoral Thesis)
Santonja M (2020) Achelense y Paleolítico medio antiguo en la Península Ibérica. Evolución tecnológica o diferentes orígenes? Actualidad de la Investigación Arqueológica en España I (2018–2019). Conferencias impartidas en el Museo Arqueológico Nacional. Ministerio de Cultura y Deporte. Madrid, pp 137–157
Santonja M, Querol MA (1977) La gravera cuaternaria de las Acacias en el río Jarama (Mejorada del Campo). In: Actas del XIV Congreso de Arqueología Nacional (Vitoria 1975), Zaragoza, pp 49–56
Santonja M, Querol MA (1980) Las industrias achelenses en la región de Madrid. In: Santonja M, López N, Pérez-González A (eds) Ocupaciones achelenses en el valle del Jarama. Arqueología y Paleontología I. Diputación Provincial de Madrid, pp 29–48
Santonja M, Pérez-González A (1997) Los yacimientos achelenses en terrazas fluviales de la Meseta Central española. In: Rodríguez Vidal J (ed) Cuadernillo Ibérico: 224–234
Santonja M, Pérez-González A (2000–2001) El Paleolítico inferior en el interior de la Península Ibérica. Un punto de vista desde la Geoarqueología. Zephyrus 53–54: 27–77. http://hdl.handle.net/10366/71572
Santonja M, Vega LG (2002) La investigación del valle del Manzanares (1862-1975) en el contexto del Paleolítico español. In: Panera J, Rubio-Jara S (eds) Bifaces y elefantes. La investigación del Paleolítico Inferior en Madrid. Museo ArqueológicoRegional de la Comunidad de Madrid. Zona Arqueológica 1, pp 243–275
Santonja M, Vega LG (2008) Pérez de Barradas en la investigación del Paleolítico del Valle del Manzanares. In: Arqueología, América, Antropología, José Pérez de Barradas 1897-1981. Museo de los Orígenes, Casa de San Isidro, Madrid, pp 45–61
Santonja M, López Martínez N, Pérez González A (1980) Ocupaciones achelenses del valle del Jarama (Arganda, Madrid), Arqueología Y Paleoecología, 1. Servicios de Extensión cultural y Divulgación de la Diputación de Madrid, Madrid, pp 352
Santonja M, Pérez-González A, Vega G, Rus I (2001) Elephants and stone artifacts in the Middle Pleistocene terraces of the Manzanares river (Madrid, Spain). The World of Elephants. Proceedings of the International Congress (Roma), pp 597–601
Santonja M, Pérez-González A, Vega L.G, Uribelarrea D (2011) La evolución de las ideas sobre el paleolítico en Madrid. In: Pérez R, Pastor F.J, Rodríguez R (eds) Actas de las quintas jornadas de Patrimonio Arqueológico en la Comunidad de Madrid, Comunidad de Madrid, Dirección General de Patrimonio Histórico, pp 27–60
Santonja M, Pérez-González A, Domínguez-Rodrigo M, Panera J, Rubio-Jara S, Sesé C, Soto E, Lee James A, Mathieu D, Demuro M, Ortiz JE, Torres T, Mercier N, Barba R, Yravedra J (2014a) The Middle Paleoltihic site of Cuesta de la Bajada (Teruel, Spain): a perspective on the Acheulean and Middle Paleolithic technocomplex in Europe. J Archaeol Sci 49:556–571
Santonja M, Pérez-González A, Panera J, Rubio-Jara S, Sesé C, Soto E, Sánchez-Romero L (2014b) Ambrona and Torralba archaeological and paleontological sites. In: Sala R (ed) Pleistocene and Holocene Hunter-Gatherers in Iberia and the Gibraltar Strait. The Current Archaeological Record. Universidad de Burgos & Fundación Atapuerca, Burgos, pp 517–527
Santonja M, Pérez-González A, Panera J, Rubio-Jara S, Méndez-Quintas E (2016) The coexistence of Acheulean and Ancient Middle Palaeolithic technocomplexes in the Middle Pleistocene of the Iberian Peninsula. Quaternary International 411, B: 367–377. https://doi.org/10.1016/j.quaint.2015.04.056
Santonja M, Rubio-Jara S, Panera J, Sánchez-Romero L, Tarriño A, Pérez-González A (2018) Ambrona revisited: The Acheulean lithic industry in the Lower Stratigraphic Complex. Quat Int 480:95–117. https://doi.org/10.1016/j.quaint.2017.01.021
Sesé C, López-Martínez N (2013) Nuevos datos paleontológicos del Pleistoceno en el Valle del Manzanares (Madrid, España): Los micromamíferos del yacimiento del Arenero de Arriaga. Estud Geol 69(2):271–282
Sesé C, Panera J, Rubio-Jara S, Pérez-González A (2011a) Micromamímeros del Pleistoceno Medio y Pleistoceno Superior en el Valle del Jarama: yacimimientos de Valdocarros y HAT (Madrid). Estud Geol 67(2):131–151
Sesé C, Rubio-Jara S, Panera J, Pérez-González A (2011b) Micromamíferos del Pleistoceno Superior del yacimiento de PRERESA en el valle del Manzanares y su contribución a la reconstrucción paleoambiental de la cuenca de Madrid durante el Pleistoceno. Estud Geol 67(2):471–494
Sesé C, Soto E (2000) Vertebrados del Pleistoceno de Madrid. In J. Morales (Coord.) Patrimonio Paleontológico de la Comunidad de Madrid. Consejería de Educación de la Comunidad de Madrid. 216–243
Sesé C, Soto E (2002) Catálogo de los yacimientos de vertebrados del Pleistoceno en las terrazas de los ríos Jarama y Manzanares. In: Panera, J., Rubio-Jara, S. (Eds.), Bifaces y elefantes. La investigación del Paleolítico Inferior en Madrid. Museo Arqueológico Regional de la Comunidad de Madrid. Zona Arqueológica 1, pp. 430–457
Shipman P (1981) Life history of a fossil. Harvard University Press, Cambridge, Massachusetts and London, England, An introduction to taphonomy and paleoecology
Shipman P, Rose J (1983) Early hominid hunting, butchering and carcass-processing behaviours: approaches to that fossil record. J Anthropol Archaeol 2:57–98
Silva PG (2003) El valle inferior del Manzanares (Cuenca de Madrid, España). Volumen en Homenaje a D. Manuel Hoyos Estudios Geológicos 59:107–131
Silva PG, Goy JL, Zazo C (1988a) Neotectónica del sector centro-meridional de la Cuenca de Madrid. Estud Geol 2:133–136
Silva PG, Goy JL, Zazo C, Hoyos M, Alberdi MT (1988b) In: El valle del Manzanares y su relación con la depresión Prados-Guaten durante el Pleistoceno inferior (Madrid, España). II Congreso geológico de España. Comunicaciones, vol. 1 (Granada, 1988b), pp. 403–406
Silva PG, López-Recio M, Tapias F, Roquero E, Morín J, Rus I, Carrasco-García P, Giner-Robles JL, Rodríguez-Pascua MA, Pérez-López R (2013) Stratigraphy of the Arriaga Palaeolithic sites. Implications for the geomorphological evolution recorded by thickened fluvial sequences within the Manzanares River valley (Madrid Neogene Basin, Central Spain). Geomorphology 196:138–161. https://doi.org/10.1016/j.geomorph.2012.10.019
Tixier J (1956) Le hachereau dans l’Acheuléen Nord-Africain. Notes typologiques. Congrès Préhistorique de France, XVème sesión (Poitiers), pp. 914–923
Uribelarrea D (2008) Dinámica y evolución de las llanuras aluviales de los ríos Manzanares, Jarama y Tajo, entre las ciudades de Madrid y Toledo. Tesis Doctoral. Universidad Complutense de Madrid. Facultad de Ciencias Geológicas. Departamento de Geodinámica. p. 396
Van der Made J (2019) The dwarfed “giant deer” Megaloceros matritensis n.sp. from the Middle Pleistocene of Madrid – a descendant of M. savini and contemporary to M. giganteu. Quatern Int 520:110–1139. https://doi.org/10.1016/j.quaint.2018.06.006
Vegas R, Pérez-González A, Miguez F (1975) Cartografía y memoria geológica del Mapa Geológico de España a escala 1:50.000, Serie MAGNA, nº de Hoja 582 (Getafe). IGME. Madrid
Villa P (1990) Torralba and Aridos: elephant exploitation in Middle Pleistocene Spain. J Hum Evol 19(3):299–309
Villa P, Soto E, Santonja M, Pérez-González A, Mora R, Parcerisas J, Sesé C (2005) Nuevos datos sobre Ambrona: cerrando el debate caza versus carroñeo. In Los yacimietos paleolíticos de Ambrona y Torralba (Soria). Un siglo de investigaciones arqueológicas. Zona Arqueológica 5:352–380
Yravedra J (2006) Tafonomía Aplicada a Zooarqueología. Aula Abierta. Universidad Nacional de Educación a Distancia (UNED), Madrid
Yravedra J (2010) Estudio tafonómico y zooarqueológico de los macromamíferos del yacimiento arqueológico de Tafesa (Villaverde Bajo, Mabrid). In Las huellas de nuestro pasado. Estudio del Yacimiento del pleistoceno madrileño de Tafesa (Antigua Transfesa). Zona Arqueológica 14:155–176
Yravedra J, Domínguez-Rodrigo M, Santoja M, Pérez-González J, Panera J, Rubio-Jara S, Baquedano E (2010) Cut marks on the Middle Pleistocene elephant carcass of Áridos 2 (Madrid, Spain). J Archaeol Sci 37:2469–2476. https://doi.org/10.1016/j.jas.2010.05.007
Yravedra J, Panera J, Rubio-Jara S, Manzano I, Expósito A, Pérez-González A, Soto E, López-Recio M (2014) Neanderthal and Mammuthus interactions at EDAR Culebro 1 (Madrid, Spain). Journal of Archaological Science 42:500–508
Yravedra J, Panera J, Rubio-Jara S, Pérez-González A, Gallego N, González I (2019a) Middle Pleistocene Human occupation in the interior of the Iberian Peninsula during cold climate conditions: Zooarchaeology and Taphonomy of ETB-H02 site in the Manzanares valley (Madrid, Spain). Quatern Int 520:99–109. https://doi.org/10.1016/j.quaint.2018.03.014
Yravedra J, Rubio-Jara S, Panera J, Martos JA (2019b) Hominis and proboscideans in the Lower and Middle Paleolithic in the Central Iberian Peninsula. Quatern Int 520:140–156. https://doi.org/10.1016/j.quaint.2017.12.002
Yravedra J, Rubio-Jara S, Panera J, Van der Mada J, Pérez-González A (2019c) Neanderthal diet in fluvial environments at the end of the Middle Pleistocene / early Late Plesitocene of PRERESA site in the Manzanares Valley (Madrid, Spain). Quatern Int 520:72–83. https://doi.org/10.1016/j.quaint.2018.01.030
Yravedra J, Rubio-Jara S, Panera J, Uribelarrea D, Pérez-González A (2012) Elephants and subsistence. Evidence of the human exploitation of extremely large mammal bones from the Middle Palaeolithic site of PRERESA (Madrid, Spain). J Archaeol Sci 39:1063–1071. https://doi.org/10.1016/j.jas.2011.12.004
Yravedra J, Solano JA, Lloyd AC, Saarinen J, Linares-Matás G, Luzón C, Serrano-Ramos A, Herranz-Rodrigo D, Cámara JM, Ruiz A, Titton S, Rodríguez-Alba JJ, Mielgo C, Blain HA, Agustí J, Sánchez-Bandera Ch, Montilla E, Toro-Moyano I, Fortelius M, Oms O, Barsky D, Jiménez-Arenas JM (2021) Use of meat resources in the Early Pleistocene assemblages from Fuente Nueva 3 (Orce, Granada, Spain). Archaeol Anthropol Sci 13:213. https://doi.org/10.1007/s12520-021-01461-7
Open Access funding provided thanks to the CRUE-CSIC agreement with Springer Nature. This work has been carried out in the framework of Grants PGC2018-093612-B-100, funded by MICIN/AEI/10.13039/501100011033 and ERDF; CEN154P20 and co-financed by the ERDF 728 (European Regional Development Fund) and the Junta de Castilla y León.
We would like to thank the National Archaeological Museum (MAN) for providing us with the facilities and technical equipment necessary for our study.
All authors agreed to participate in the Archaeological and Anthropological Sciences Journal and present a new study that has not been published before.
All authors read the journal policies of the Archaeological and Anthropological Sciences Journal about publishing and agreed to publish our article. All authors have worked together to reproduce a new-fangled work.
Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations.
Rights and permissions
Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.
About this article
Cite this article
Claver, I., Martos, J.A., Yravedra, J. et al. The interaction between large mammals and Acheulean tools during the Middle Pleistocene in the Manzanares valley (Madrid, Spain): new evidence for Santa Elena and Oxígeno sites. Archaeol Anthropol Sci 15, 5 (2023). https://doi.org/10.1007/s12520-022-01695-z
- Acheulean technocomplex
- Human carnivore interaction
- Middle Pleistocene landscape
- Cut marks
- Tooth marks