Abstract
Since its publication in 1849, Carex ignota has been overwhelmingly ignored or treated as a synonym of C. striatula. Here, I present results of statistical analyses that support C. ignota as morphologically distinct from C. striatula and designate a lectotype for C. ignota. I also present the geography of C. ignota and C. striatula from over 750 examined and georeferenced specimens and results of maximum likelihood phylogenetic reconstruction of ddRAD loci. Carex ignota is distinguished from C. striatula by its narrower leaves on vegetative shoots, more loosely-flowered pistillate spikes, staminate spike with longer peduncles, and narrower perigynia. Based on strong morphologic and molecular evidence, C. ignota should be resurrected and treated as a distinct species. Carex ignota occurs on the Coastal Plain in the southeastern United States, as well as northern Alabama, the Ouachita in Arkansas, and the Piedmont in Georgia, where it grows in sandy to loamy soils in ravines and on slopes in mesic deciduous or mixed hardwood-conifer forests.
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Acknowledgements
I would like to extend special thanks to Charles Bryson and Robert Naczi for sharing their collections and for assistance during field work. I would like to thank: BRIT, GH, and PH for specimen loans, and also GH, NY, P, and PH for allowing me to visit their collections and search for type material; Donna Ford-Werntz, John Freudenstein, Rick Gardner, Jason Larson, Robert Naczi, Rich Rabeler, Mesfin Tadesse, Jim Vanderhorst, and Michael Vincent for sending me photographs of herbarium specimens or sharing their data for records of Carex striatula; Kanchi Gandhi for very helpful discussions on nomenclature and typification; Damon Little for helping extensively in designing and testing the protocol for building ddRAD libraries, and for help analyzing the resulting data; Jing Leong, Dave Lohman, and Y-Lan Nguyen for help accessing and operating the Pippin Prep at City College (New York, NY); Walter Kittridge for imaging and barcoding the lectotype of C. ignota at GH; the staff at the Gray Herbarium Library for granting permission to publish a photo of the C. ignota lectotype; Chick Gaddy for sharing a collection of C. ignota; and Jordan Hoffman and Simon Verlynde for assistance photographing and editing images of C. ignota and C. striatula. Field work in the southeastern United States was supported by funding from The Commodore Matthew Perry Graduate Studies Program at the New York Botanical Garden and the Earl Core Student Research Award from Southern Appalachian Botanical Society. I would like to acknowledge The Lewis B. and Dorothy Cullman Program for Molecular Systematics Studies (the New York Botanical Garden) for providing funding for ddRAD sequencing and the CUNY Doctoral Student Research Grant program for additional funding for ddRAD sequencing and for travel to GH and PH to search for type material. The Alabama Department of Natural Resources, Alabama State Parks Division, Kentucky Department of Parks, Mississippi Natural Heritage Program, Pisgah Civitan Club, and United States Forest Service granted permission to collect on their properties for this project. Thank you to the following individuals for assistance with permits and collection permission: Frank Allen, Wayne Barger, Forrest Bailey, Ralph Edmonds, Gary Kauffman, Heather Sullivan, David Taylor, and Ron Vanover. Finally, thank you to Robert Naczi and three anonymous reviewers for providing comments and edits that helped improve this manuscript.
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Appendices
Appendix 1
Specimens measured.
Those marked by * are voucher specimens for the phylogenetic reconstruction from ddRAD sequencing data presented in Fig. 3. Those marked by † are voucher specimens for the photos in Fig. 4. Coordinates given in square brackets are estimated from the label locality data.
Carex ignota.—U.S.A. Alabama: Chilton County: ca. 4.5 mi NE of Verbena, along N side of route 22, 1.3 road mi W of Coosa County border, 32°47′3”N, 86°26′58”W, 23 Apr 2002, R. F. C. Naczi 9099 (NY)*†. Coosa County: ca. 3.5 mi SW of Unity, along W side of county route 55, 7.3 road mi E of eastern edge of bridge over Coosa River, [approx. 32°58′51”N, 86°24′26”W], 14 Apr 2000, R. F. C. Naczi 8373 (NY). Lawrence County: Bankhead National Forest, Thompson Creek crossing of Forestry Rd. 208, [approx. 34°20′51”N, 87°13′56”W], 8 May 1988, C. T. Bryson 7601 (NY); along an old logging road near Beech Creek, compartment 47 off FR-244 in the Bankhead National Forest, Warrior Basin district of the Cumberland Plateau, 34°18′9”N, 87°19′5”W, 19 Apr 2003, D. D. Spaulding 11,738 (NY). Marion County: along east branch of Dickenson Creek southwest of Brilliant and west of SR-129, Warrior Basin district of the Cumberland Plateau, 34°0′15”N, 87°47′48”W, 3 May 2003, D. D. Spaulding 11,818 (NY). Russell County: ca. 2.5 mi NW of Cottonton, N of county route 4, along E side of Sand Branch, [approx. 32°9′58”N, 85°6′42”W], 3 May 1996, R. F. C. Naczi 5198 (NY). Tuscaloosa County: ca. 11 mi SE of Windham Springs, just S jct. of Blue Creek and Watermelon Road, 33°27.001’N, 87°24.7851’W, 2 May 2005, C. T. Bryson 20,518 (NY). Winston County: ca 4.4 air mi SW of Addison, along W side of Brushy Creek, just S of route 278, [approx. 34°10′47”N, 87°13′55”W], 21 May 1996, R. F. C. Naczi 5435 (NY). Arkansas: Ashley County: South end of Georgia Pacific Lake, 8 miles northwest of Crossett, [approx. 33°12′57”N, 92°02′07”W], 28 Apr 1985, S. L. Hooks 109 (NY). Dallas County: Ark. 229 on south side of Bryant Creek about 10.5 miles south of Carthage and Ark. 48, [approx. 33°57′16”N, 92°29′22”W], 10 May 1996, R. D. Thomas 148,467 (NY). Nevada County: along Ark. 299 at Cypress Creek at county road 40 east of Morris, [approx. 33°40′21”N, 93°11′54”W], 14 Apr 2001, R. D. Thomas 169,313 (NY)*. Florida: Hamilton County: Holton Creek Wildlife Management Area, south of Road 4 and west of Road 26, north of the Suwannee River, east of Holton Creek, [approx. 30°25′54”N, 83°3′19”W], 3 Apr 2000, R. J. Abbott 13,391 (NY). Jackson County: N side Chipola River just N of Mariana where Fla. 167 crosses, [approx. 30°47′37”N, 85°13′18”W], 19 Mar 1976, R. Kral 57,393 (NY). Walton County: tributary to Oak Grove Branch, ca. 0.7 air mi SE of Douglas Crossroads, ca. 2.7 air mi NNW of Knox Hill, Ponce de Leon 7.5′ Quad, 30°40′48”N, 85°58′7”W, 8 Apr 1993, S. L. Orzell 21,412 (BRIT). Georgia: Baldwin County: Scottsboro, along both sides of route 441, 0.5 road mi NW of its junction with route 243, [approx. 33°1′41”N, 83°14′34”W], 21 May 1997, R. F. C. Naczi 6082 (NY). Lowndes County: Troupville Woods, just W of Valdosta, along S side of Hwy GA 94, E of Withlacoochee River, [30°46′59”N, 83°19′56”W], 13 Apr 1987, R. Carter 5367 (NY). Louisiana: Catahoula Parish: Sicily Island Hills south of Leland Lookout Tower, [approx. 31°52′10”N, 91°44′28”W], 3 Mar 1975, R. D. Thomas 42,716 (NY). Morehouse Parish: intersection of Cay Walker Road and Boney Trail south of La. 834 west of Jones, [approx. 32°58′26”N, 91°42′41”W], 30 Apr 1992, R. D. Thomas 128,312 (NY). Ouachita Parish: E side Selman Field, Monroe, [approx. 32°31′12”N, 92°3′25”W], 5 Apr 1959, R. Kral 8406 (BRIT). Rapides Parish: Beaver pond north of Mill Creek Road, southwest of Pecan Road, and east of Lee Bridge Road, [approx. 31°24′57”N, 92°17′17”W], 25 Mar 1994, C. Slaughter 4613 (NY). Mississippi: Amite County: ca. 14 mi ENE of Crosby, Homochitto National Forest, E of Forest Service Road 107L, 31°20′12”N, 90°50′38”W, 4 May 2004, R. F. C. Naczi 10,204 (NY). Carroll County: ca. 2.2 mi W Carrollton, S of US Hwy 82, [approx. 33°30′9”N, 89°57′60”W], 22 Apr 1987, C. T. Bryson 5405 (BRIT). Forrest County: ca 14 mi SE of Hattiesburg, Ragland Hills, adjacent to Camp Shelby (U.S. National Guard), [approx. 31°12′22”N, 89°10′4”W], 30 Apr 1996, R. F. C. Naczi 5098 (NY); Ragland Hills, SE of Hattiesburg, [approx. 31°12′30”N, 89°10′54”W], 9 Apr 1969, K. Rogers 2133-B (BRIT). Franklin County: ca. 5 air mi S of Roxie, W jct. of Hwy MS 33 and White Apple Road, behind White Apple Church and cemetery in a ravine, [approx. 31°26′44”N, 91°6′19”W], 24 Apr 2007, C. T. Bryson 22,079 (NY); Homochitto National Forest, Okhissa Lake site at end of USFS 149 east of US 98 south of Bude, [approx. 31°26′44”N, 91°6′19”W], 11 Apr 2003, R. D. Thomas 173,716 (NY). Holmes County: 6.2 mi E Thornton, [approx. 33°4′21”N, 90°13′18”W], 14 Apr 1992, C. T. Bryson 11,220 (NY). Lincoln County: SE of Wesson, Lincoln State Park, S of Lake Lincoln, [approx. 31°40′48”N, 90°20′22”W], 16 Apr 2004, C. T. Bryson 20,044 (NY). Lowndes County: Columbus Air Base, W side, [approx. 33°38′45”N, 88°28′46”W], 19 May 1994, C. T. Bryson 13,721 (NY). Neshoba County: ca. 6 mi N of Philadelphia, Burnside Lake, N of Pearl River, 32°50′44”N, 89°6′14”W, 21 May 2004, C. T. Bryson 20,262 (NY). Pearl River County: ca. 3.2 mi S Marion-Pearl River Co. line, above Mill Creek, W of MS Hwy 43, [approx. 30°58′6”N, 89°41′45”W], 26 Apr 1991, C. T. Bryson 10,753 (NY). Perry County: ca.11 mi NE of Wiggins, NW of junction of forest road 318B-1 and Black Creek Trail, along Black Creek Trail, [approx. 30°59′38”N, 89°2′33”W], 30 Apr 1996, R. F. C. Naczi 5113 (NY). Simpson County: NW of Mendenhall, S of Hwy US 49 and Strong River, [approx. 31°59′2”N, 89°53′9”W], 9 May 2001, C. T. Bryson 18,619 (NY); Simpson Legion Lake, to N of Hwy US 49, [approx. 31°55′2”N, 89°47′29”W], 8 May 2002, C. T. Bryson 19,241 (BRIT). Wilkinson County: W of Pond, Miss., [approx. 31°4′14”N, 91°30′36”W], 28–30 Apr 1978, K. Rogers 45,129 (BRIT). Winston County: 11.8 km NE of Louisville, W side of MS Hwy 25, Tombigbee National Forest, 33.211850°N, 88.986150°W, 20 Apr 2015, J. E. Dorey 150.2 (NY)*; 6.7 km NE of Highpoint, along Pigeon Roost Rd., Tombigbee National Forest, 33.231300°N, 89.095966°W, 20 Apr 2015, J. E. Dorey 161 (NY)*. Yazoo County: ca. 4 air mi ESE center of Yazoo City, ca 2 mi E jct. of Hwy US 49 and MS 16, S of Hwy US 16 near Parkview Church of God, 32°49′52”N, 90°21′13”W, 18 Apr 2010, C. T. Bryson 23,396 (NY)*. South Carolina: Berkeley County: bluff of Huger Creek, [approx. 33°6′5”N, 79°49′30”W], 2 May 2016, L. L. Gaddy s.n. (NY)*. Horry County: 5 miles south of Myrtle Beach, [approx. 33°39′2”N, 78°57′46”W], 2 Apr 1939, R. T. Clausen 3757 (NY); 2 miles south of Myrtle Beach, [approx. 33°32′56”N, 79°3′7”W], 18 Apr 1932, C. A. Weatherby 16,458 (NY).
Carex striatula.—U.S.A. Alabama: Barbour County: ca. 9 air mi SSW of Eufala, along E side of route 431, just S of its crossing of White Oak Creek, [approx. 31°46′16”N, 85°11′5”W], 4 May 1996, R. F. C. Naczi 5218 (NY). Coosa County: 1.8 mi S of Marble Valley, along N side of Paint Creek, ca. 0.4 mi W of Little Tom Road, 33°1′1”N, 86°27′11”W, 29 Apr 2000, R. F. C. Naczi 8538 (NY). Elmore County: ca. 1.8 mi SE of Titus, along E side of Weoka Creek, ca. 0.1 mi S of its crossing by county route 432, 32°41′54”N, 86°17′29”W, 23 Apr 2002, R. F. C. Naczi 9120 (NY). Jackson County: ca. 3 mi W of Carns, flank of Cumberland Plateau, along county route 33, 2.7 road mi S of junction of county route 33 and county route 39, 34°48′56”N, 86°1′36”W, 22 Apr 2002, R. F. C. Naczi 9036 (NY). Madison County: 2.9 mi E of center of Huntsville, Monte Sano Mountain, 0.1 mi E of Monte Sano Boulevard SE along Natural Well Trail, 34.72242°N, 86.53577°W, 26 Apr 2015, R. F. C. Naczi 15,759 (NY). Pickens County: NE of Ethelsville, ca. 0.2 mi W jct. Of Hwy US 82 and Pickens County Road 75, N of Hwy US 61, ca. 6.1 mi E of Alabama-Mississippi state line, [approx. 33°26′11”N, 88°11′45”W], 22 Apr 2006, C. T. Bryson 21,283 (NY). Delaware: Kent County: ca. 0.5 mi NW of Clayton, ca. 0.1 mi S of Providence Creek, ca. 0.5 mi W of its crossing by route 15, just N of cemetery of Saint Joseph’s Industrial School, 39°17′44”N, 75°38′48”W, 15 May 2002, R. F. C. Naczi 9282 (NY). Georgia: Chattahoochee County: ca. 9 air mi W of Cusseta, along E side of Oswichee Creek, S of crossing by Sunshine Road, ca. 0.4 mi N of Chattahoochee River, [approx. 32°17′5”N, 84°52′53”W], 23 May 1997, R. F. C. Naczi 6136 (NY). Columbia County: 5 mi NNE of Appling, along stream crossed by route 221, W of route 221, 0.9 mi S of junction of routes 221 and 104, 33°36′33”N, 82°16′50”W, 7 May 2003, R. F. C. Naczi 9526 (NY)*. Monroe County: ca. 2.5 mi SW of Russellville, N of route 74, 3.9 mi E of junction of routes 74 and 341, 32°52′52”N, 84°1′55”W, 25 Apr 2000, R. F. C. Naczi 8404 (NY). Upson County: ca. 3 mi SW of Thomaston, S of route 36 at shoals of Potato Creek, along W side of creek, 32°52′32”N, 84°22′27”W, 25 Apr 2000, R. F. C. Naczi 8421 (NY)*. Kentucky: Bell County: ca. 1.5 mi SW of Pineville, Pine Mountain State Nature Preserve, along park road, ca. 0.2 mi NE of campground, [approx. 36°44′56”N, 83°42′48“W], 26 Jun 1997, R. F. C. Naczi 6732 (NY). McCreary County: 3.9 mi SW of community of Hill Top, along E side of Beech Grove Road, 36°40’13”N, 84°33′41“W, 24 Apr 2010, R. F. C. Naczi 12869 (NY)*. Maryland: Dorchester County: Le Compte Wildlife Management Area, along Steels Neek Road, 1.5 mi south of New Bridge Road, 38°27’39”N, 75°53′29“W, 14 May 2001, C. lea 2344 (NY); North of the town of Vienna, along Chicone Road east of Chicone Creek, 38°31’3”N, 75°49′13“W, 13 May 2007, W. D. Longbottom 8782 (NY)*. Mississippi: Grenada County: ca. 2 mi E of Gore Springs, along S side of route 8, 33°45’34”N, 89°35′55“W, 29 May 2003, R. F. C. Naczi 9694 (NY). Lauderdale County: ca. 2 mi SW of Suqualena, [approx. 32°25’32”N, 88°50′31“W], 17 Apr 1985, K. L. Gordon 3098. (NY). Lee County: 5.1 km NE of Plantersville, E side of Group Camp Dr., Tombigbee State Park, 34.235500°N, 88.616533°W, 21 Apr 2015, J. E. Dorey 182 (NY). Simpson County: Simpson Legion Lake, to N of Hwy US 49, [approx. 31°55’2”N, 89°47′29“W], 8 May 2002, C. T. Bryson 19241 (NY). Tishomingo County: ca. 1.3 mi S Mingo, to W of Tishomingo Co. Rd. 85, [approx. 34°36’18”N, 88°8′59“W], 17 May 1992, C. T. Bryson 11686 (NY). Winston County: 6.0 km NE of Highpoint, Tombigbee National Forest, vicinity of Little Rock Cemetery, N of Noxubee Crest, 33.220917°N, 89.095767°W, 20 Apr 2015, J. E. Dorey 160 (NY). Yalobusha County: S side of Lake Tillatoba, Holly Springs National Forest Recreation Area, [approx. 33°58’39”N, 89°49′8“W], 16 April 1994, C. T. Bryson 13381 (NY). New Jersey: Ocean County: Point Pleasant, [approx. 40°5’14”N, 74°4′15“W], 30 May 1898, E. P. Bicknell s.n. (NY). North Carolina: Alleghany County: Cherry Lane Township, Miles/Roaring Gap, located along Lyons Rd. (SR 1102), 0.6 mi northwest of the Oklahoma Rd. (SR1100) jct, Stone Mountain State Park property, 36°23’25.6”N, 80°59′59.2“W, 19 May 2010, D. Poindexter 10-203 (NY). Durham County: back of Physics Building, Duke Univ., [approx. 36°0’13”N, 78°56′32“W], 18 May 1949, H. L. Blomquist 14505 (NY). Jackson County: 7.9 km SE of Highlands, 0.05 km N of Chattooga River, N side of FS road 1178 (Bull Pen Rd.), 35.015205°N, 83.122619°W, 11 Jun 2015, J. E. Dorey 1183 (NY). South Carolina: Berkeley County: Old Santee Canal State Park, [approx. 33°11’32”N, 79°58′10“W], 18 Apr 2004, C. Frye 5269 (NY)*. Edgefield County: On W side of Horn Creek, just N of FR 634, about 9 mi SW of Edgefield, [approx. 33°42’43”N, 82°1′19“W], 24 Apr 1997, Nelson 18139 (NY). Greenwood County: 2 mi W of Kirksey, along tributary of Cuffeytown Creek, 34°2’8”N, 82°5′3“W, 26 Apr 2002, R. F. C. Naczi 9197 (NY). Lancaster County: S side of Flat Creek, 1.1 air mi SE of jct US 601 and Sec Hwy 27, 1.65 air mi NE of Flat Creek School (jct US 601 and SC 903), 34.6445°N, 80.5009°W, 10 May 2005, J. B. Nelson 25368 (NY)*†. McCormick County: overlooking Little River, [approx. 33°53’7”N, 78°36′53“W], 28 May 1950, W. H. Duncan 10993 (NY). Orangeburg County: Santee Cooper WMA hardwood regeneration area, 33.425°N, 80.2867°W, 12 Apr 2001, A. B. Pittman 4120123 (NY); Santee State Park, 33.517°N, 80.4844°W, 9 May 2001, A. B. Pittman 5090129 (NY). York County: ca. 4 mi NW of Bethany, Kings Mountain State Park, W of campsite #53, [approx. 35°8’54”N, 81°27′25”W], 17 May 1996, R. F. C. Naczi 5311 (NY).
Appendix 2. Characters measured.
Character abbreviation | Description |
LfWV | Widest leaf on vegetative shoots in millimeters. |
TspPd | Longest peduncle of the staminate spike in millimeters. |
RNpgPs | Ratio of the number of perigynia to spike length (mm) on the second-most distal pistillate spike. This spike was chosen to specifically exclude the distal-most pistillate spike, as it is sometimes bears only a few perigynia. |
RBSBB | Ratio of the bract sheath length to the bract blade length on the second-most distal pistillate spike. This spike was chosen to specifically exclude the distal-most pistillate spike, as plants sometimes display an aberrant morphology where perigynia are borne at the base of the terminal spike. |
PgL | Length from the base to the apex of the perigynium in millimeters. |
PgW | Width of the perigynium in the sagittal plane in millimeters. |
Appendix 3
Additional selected representative specimens.
Carex ignota.—U.S.A. Tennessee: Benton County: Mark Carroll Forest, [approx. 36°09′19”N, 88°03′50”W], 11 May 1993, DeSelm s.n. (EKY barcode n. 31234100062155). Henderson County: Natchez Trace State Forest on Shiloh Rd. near Coffee Trail, [approx. 35°48′21”N, 88°12′30”W], 27 May 1993, DeSelm s.n. (TENN barcode n. TENN-V-0050847). Texas: Cass County: Ca. 1 mi SE of Zion Hill Church, on tributary of Wilburn Creek, ca 3 mi E of US 59 at Lanier, 32°55′20”N, 94°19′00”W, 27 Mar 1987, Orzell 4934 (MICH barcode n. 1388018). Harrison County: Along Cypress Bayou, near Jefferson, [approx. 32°45′29”N, 94°21′34”W], 23 May 1958, Correll 18,784, (LL barcode n. 00342032). Houston County: 0.1 of a mile NW on FR 227 from its jct with Davy Crockett (NF) Road 526, [approx. 31°29′25”N, 95°13′32”W], 10 Apr 1990, Jones 4310 (MICH barcode n. 1388014). Jasper County: Between Martin Dies Jr. State Park Road 48 N and 48 S on US 190, E of Woodville, [approx. 30°51′11”N, 94°10′10”W], 24 Mar 1991, Jones 6266 (MICH barcode n. 1388097). Marion County: Along & just E of Pine Bluff, E of N arm of Caddo Lake, N of Potters Point, 32°43′07”N, 94°05′03”W, 29 Mar 1987, Orzell 4946 (MICH barcode n. 1388016). Polk County: Livingston, [approx. 30°42′40”N, 94°55′59”W], 14 Apr 1914, E. J. Palmer 5247 (F). Rusk County: E of Naco. Co. Rd. #173 ext., ca. 2.1 mi N of int. FM 1087 (at Freewill Baptist Church); ENE of Sandy Creek crossing, 31°52′03”N, 94°34′26”W, 14 Apr 1988, Orzell 6264 (TEX barcode n. 00293353). Sabine County: Near Red Hills Lake. Sabine National Forest, Compartment 72, Ca. 3 miles N. of Milam, [approx. 31°28′29”N, 93°49′57”W], 8 Mar 1996, MacRoberts 2985 (TEX barcode n. TEX00342034). San Augustine County: N & NE of FS Rd. 184 crossing of Crooked Creek, ca 2 air mi SSE of jct FM 353 & FM 2785 at White Rock, ca 5 air mi E of San Augustine, 31°31′57”N, 94°01′19”W, 12 Apr 1989, Orzell 9198 (MICH barcode n. 1388017). San Jacinto County: Along Big Creek, N of FS Rd. 217, ca 1.7 mi W of TX 150 at a point 4.6 mi NW of Shepherd, 30°30′40”N, 95°05′25”W, 21 Apr 1988, Orzell 6439 (MICH barcode n. 1388015). Shelby County: E of FM 2261, ca 0.3 air mi ESE of jct FS Rd. 106 in East Hamilton, 31°36′10”N, 93°50′38”W, 12 Apr 1989, Orzell 9264 (MICH barcode n. 1388012).
Carex striatula.—U.S.A. Connecticut: New London County: Lime Kiln, Preston [Records of a lime-kiln in NW corner of North Stonington near Preston boundary], [approx. 41°31′38”N, 71°57′56”W], 29 May 1902, Graves s.n. (CBS barcode n. 30588). District of Columbia: Broad Branch, [approx. 38°57′02”N, 77°03′09”W], 31 May 1897, Pieters s.n. (MICH barcode n. 1388072). Florida: Holmes County: on McDuffie Rd., ca. 0.9 mi NW of FL 2A, ca. 0.1 mi S of Alabama state line, 30°59′30”N, 85°57′30”W, 8 Apr 1993, Orzell 21,398 (FLAS barcode n. 195391). Jackson County: E of Pittman Hill Rd., ca 10 air mi SSW of Marianna, 30.60909°N, 85.25063°W, 19 Mar 2012, Anderson 26,132 (NY barcode n. 2305757). New York: [Queens County]: Long Island. Between 1st and 2nd reservoirs [Ridgewood Reservoir?], [approx. 40°41′14”N, 73°53′16”W], 30 May 1903, Bicknell 3119 (NYS, NY barcode n. 02335973). Suffolk County: Cutchogue, [approx. 41°00′36”N, 72°29′13”W], 10 Jul 1924, Latham 2196 (GA barcode n. 113294, NY barcode n. 02335964); Montauk, Long Island, [approx. 41°02′08”N, 71°57′21”W], 19 Jun 1927, Latham 4077 (NY barcode n. 02335965). Ohio: Jackson County: Liberty Twp., [approx. 39°04′58”N, 82°43′04”W], 9 Jun 1935, Bartley s.n., (OS accession n. 439,541). Vinton County: Vinton Furnace Experimental Forest, Arch Rock Study Area, [approx. 39°11′26”N, 82°22′18”W], 1 Jun 1999, Hines 64 (BHO barcode n. 0004215). Pennsylvania: Berks County: 0.75 mi SW of McKnights Gap, [approx. 40°21′42”N, 75°54′02”W], 12 June 1941, Berkheimer 2556 (PH barcode n. 00184688). Centre County: Edge of Spring Creek 3.5 mi N of Lemont, [approx. 40°51′44”N, 77°48′18”W], 2 Jun 1950, Wahl 8775 (PAC barcode n. 0007957). Lancaster County: Near Haines Station, [39°44′21”N, 76°12′45”W], 5 June 1901, Heller 4843 (NY barcode n. 02335966). Tennessee: Marion County: West on highway 27 towards Powells Cross-roads. 1.2 miles W on Kellys Ferry Road, [approx. 35°07′16”N, 85°24′39”W], 15 Apr 1998, Beck 1505 (UCHT barcode n. UCHT004289). Roxane County: Oak Ridge Reservation, Natural Area 51, just N of Melton Hill Lake, 35°53′20.4”N, 84°18′57.6”W, 28 Apr 2010, Durr 10–004-1 (TENN barcode n. TENN-V-0050860). Virginia: Pittsylvania County: Boyd Mtn., ca. 3 km NE of Leaksville Junction on Southern Railroad, [approx. 36°33′44”N, 79°37′12”W], 28 Apr 1993, Wieboldt 8456 (NY). Prince William County: Telegraph Road Picnic Area, Prince William Forest Park, [approx. 38°33′31”N, 77°20′33”W], 14 May 1988, Brunton 7902 (MICH barcode n. 1388058). West Virginia: Fayette County: New Haven Twp. 2.4 mi S of Nicholas Co. line, at junction of Rt. 19 and Miller Ridge Rd., [approx. 38°07′32”N, 80°58′09”W], 22 May 1991, Vincent 4717 (MU barcode n. MU-V-000171073).
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Dorey, J.E. Resurrection of Carex ignota (Cyperaceae, section Laxiflorae), a long overlooked sedge of the southern U.S.A.. Brittonia 71, 253–267 (2019). https://doi.org/10.1007/s12228-019-09581-0
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DOI: https://doi.org/10.1007/s12228-019-09581-0