Abstract
Continuation of archaeobotanical and palaeoecological research on three semi-artificial lake islands, Nowy Dworek, Chycina and Lubniewice, has provided new information on the history, development and use of grasslands in Ziemia Lubuska (Lubusz land) in western Poland during the early Middle Ages. Pollen analysis reveals that the reduction in woodland and opening up of the regional landscape and the appearance of grassland communities there began around the 7th century ad, which preceded the construction of the islands in the lakes. The analysis of plant macroremains collected from the settlement layers on these islands, of pollen from cores in the lakes and studies of the phytosociological plant communities of recent vegetation have helped to describe the botanical composition of these past meadows and pastures. The results show strong similarities between the three sites, which suggests that the local populations were using the same grassland types, which were on soils ranging from damp to rather dry and sandy. The main grassland in the vicinity of the islands included the Molinio-Arrhenatheretea syntaxonomical class of communities, although Festuco-Brometea grasslands were also present. The presence of plant remains associated with various other types of meadows and pastures shows the diversity of habitats represented by the plant remains.
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Introduction
The geographical, historical and cultural conditions in the Ziemia Lubuska (Lubusz land, western Poland), defined as an early medieval border area, make this microregion a very important place to study past settlement development. This is demonstrated by numerous publications on archaeological sites which provide data on the functioning of past local communities (Kurnatowska and Łosińska 1996; Kobyliński 2014; Kaczmarek 2019). Particularly interesting are some semi-artificial lake islands, which resemble crannogs, prehistoric island sites in the British Isles (Morrison 1985; Menotti 2012). In Poland, an interdisciplinary research project has studied the role of these unusual structures in the early medieval landscape of Ziemia Lubuska revealed by three islands in the lakes of Paklicko Wielkie, Lubiąż and Jezioro Długie (Fig. 1). Archaeological results have made it possible to define them as “central places” which were important centres of human activity for management, transport and even religious beliefs (Bleile 2010). One of the elements of this project involved recognising the environmental setting of these sites, with an emphasis on the use of plants on the islands. However, the issue of the botanical diversity of plant communities on the land around the lakes had not yet been studied in detail. The long list of taxa identified from the plant macroremains from the islands includes numerous plants which were components of natural and semi-natural plant communities as well as those connected with human activities, including meadows and pastures. The development and use of ancient grasslands poses interesting questions for both botany and archaeology (Greig 1988; Behre and Jacomet 1991; Hodgson et al. 1999; Pini et al. 2021).
Grasslands are generally strongly connected with human activity. Farming, including grazing and mowing, has the greatest impact on their appearance in the landscape, development and botanical composition (Pärtel et al. 2005; Hejcman et al. 2013; Boch et al. 2020). They are floristically rich communities which include plants from various taxonomic groups. The explanation of the role of meadows and pastures in the human landscape is rather complicated in the absence of good archaeobotanical evidence (Küster and Keenleyside 2009). The general history of meadows and pastures in Europe is known thanks to numerous pollen studies (such as Hjelle 1998; Ralska-Jasiewiczowa et al. 2006; Hejcman et al. 2013; Kuneš et al. 2015; Qin et al. 2020). More detail has been provided by studies of ancient grasslands based on plant macroremains which have been done in Switzerland (Akeret et al. 2018), Germany (Knörzer 1975; Kreuz and Schäfer 2011) and the Czech Republic (Pokorný et al. 2006; Hájková et al. 2018). In Poland, the origins of these important plant communities have not yet been discussed in much detail. Most often, grasslands are mentioned when other habitat groups are discussed, when they have been identified, for example in urban deposits (Wasylikowa 1978; Święta-Musznicka and Latałowa 2016), medieval ports (Latałowa 1999) or strongholds (Koszałka 2014). An outline of this topic was also presented in the study of materials from Żółte, another early medieval semi-artificial lake island in Pomorze Zachodnie (West Pomerania), Poland (Noryśkiewicz 2014; Pińska and Latałowa 2014).
The main objective of this article is to use the archaeobotanical data from the three islands to reconstruct the composition of the early medieval meadows and pastures which were located around the sites, and then to discuss the role of grassland communities in the local settlement system, also using the pollen data from the lake sediment.
Site description, materials and methods
The sites
The early medieval sites of Nowy Dworek, Lubniewice 10 (called Lubniewice in the following text) and Chycina represent rare small semi-artificial islands in lakes located in Ziemia Lubuska (Lubusz land) in western Poland (Fig. 1). They were elements of the local settlements. The wooden structures of the islands were built in natural shallows in the lakes. Wooden bridges and a system of piers linked the islands with the mainland. The phases of use at the individual sites have been determined from the archaeological results, dendrochronology and radiocarbon dates. They represent successive stages of use, from construction to abandonment. Near every lake and island there were contemporary settlements on dry land (Chudziak and Kaźmierczak 2020a).
The site of Chycina is situated on a lake, Jezioro Długie (95.5 ha). It is a small area of 0.3 ha raised up to 0.4–0.5 m above the present lake level, now covered with shrubs and trees, with some rushes on the shore. There were three major phases of use. Phase I (from the last quarter of the 8th to the mid 9th century) includes remains of a quay and the central part of the island. Phase II (from the mid 9th to the 1st quarter of the 10th century) covers intensive use of the interior as well as temporary flooding of the island, and Phase III (from the 2nd quarter of the 10th to the early 11th century) represents the final period of use. Samples were collected from material connected with Phases I, II, II/III and III for our archaeobotanical studies (Table 1). Most of the waterlogged organic material was collected in the samples from Phase I. Samples from the later phases contained large amounts of mineral material.
The site of Lubniewice (0.2 ha), is located in the southeastern part of Lubiąż lake (146 ha), ca. 135 m from the lake shore. Today, it is a flat area, reaching about 0.6 m above the water level. The island is covered with trees and shrubs, but there is no rush vegetation along the shore. The two main phases of occupation were distinguished by stratigraphy and archaeological finds. Phase I (the 4th quarter of the 8th to 1st half of the 9th century) corresponds to intensive use of the entire island, the building of a wooden wharf and a bridge abutment after a rise in water level. Phase II covers resettlement after a short break of settlement in the 2nd to 3rd quarters of the 10th century. Unfortunately, sufficient material for archaeobotanical analysis was preserved only in the levels associated with Phase I, which consisted mostly of waterlogged material mixed with various amounts of minerals, mainly sand (Table 1).
The smallest of the islands is Nowy Dworek (0.02 ha), rising only 0.4 m above the lake level of Paklicko Wielkie (196 ha). Today, there are only isolated trees on the island and a wide strip of rushes around the edge. There are three phases of site use according to archaeological evidence. Phase I (from the last quarter of the 8th to the mid 9th century) represents the time when the bridge, embedded log structure and shore reinforcement were built. Phase II (from the mid 9th to the 1st quarter of the 10th century) covers remains from repairs and the building of a great superstructure to raise the surface of the island. In Phase III (from the 2nd quarter of the 10th to the early 11th century), although the bridge crossing ceased to operate, the island was still intensively used. The archaeobotanical samples consisted of perfectly preserved waterlogged organic material with a few charred seeds and some sand. However, it should be noted that the cultural layers in Phase I were present only in the form of thin inclusions among the pieces of wood. Phase II provided much more organic material, which may be because typical culture layers associated with the most recent settlement phase III could have been washed away as a result of rising water levels in the following periods (Table 1). For Phase III, no suitable material for archaeobotanical analysis was found.
Materials and methods
This study is based on the results from 45 bulk samples which represent the culture layers of the three islands (Table 1). The samples represent the material used for their construction and residues from activities there. Most samples were collected from Lubniewice (19) and Nowy Dworek (17), while Chycina provided only nine. All samples contained significant numbers of macroremains of meadow and pasture plants (Fig. 2). Most of them were waterlogged, but occasional specimens were charred. Mineralized remains were found only from Chycina.
The same procedures for macrofossil sample collection strategy and laboratory preparation and then analysis were applied to all the samples. The samples were taken from trenches on the islands and in shallow water. From every sample, a 300 cm3 subsample of sediment was selected, which was soaked for 24 h in KOH solution and wet sieved with 2.0, 0.5 and 0.2 mm meshes. The remaining material was wet sieved only with a coarse sieve in order to obtain larger seeds which are normally under-represented in the main sample. Identification was made using the modern reference collection held at Laboratory of Palaeoecology and Archaeobotany, University of Gdańsk, as well as with reference to seed atlases and keys (Körber-Grohne 1991; Cappers et al. 2006). Information on the qualitative and quantitative content of the botanical material is archived in the ARCHBOT-UGDA DATABASE at the Laboratory of Palaeoecology and Archaeobotany, University of Gdańsk. The identified taxa are named according to Mirek et al. (2002) and the arrangement of plant communities uses the phytosociological classification of Matuszkiewicz (2008) and Zarzycki et al. (2002). Ecological and habitat features follow Zarzycki et al. (2002), Roo-Zielińska (2014) and Ellenberg et al. (1992). The identified species have been assigned to a particular phytosociological rank within the grassland communities, while taxa less closely associated with grassland or with their optimum niche in meadows or pastures, but without being characteristic of any of the particular plant communities (syntaxons) discussed here, have been classed as "others". The species within the rush communities of class Phragmitetea were not included in the list. Some of these taxa may represent such communities around the islands and not on the lake shores. To illustrate the relationship between the botanical composition of the samples and the individual sites, detrended components analysis (DCA) in CANOCO 4.5 for Windows was done with square-root transformation of data sets for Phase I.
Pollen analysis of the sediments of all three lakes was done in order to study the extent of open unwooded areas in the early Middle Ages, including grasslands near the individual sites. Each profile of sediments was taken at maximum water depth and as close as possible to the islands (Fig. 1). Pollen analysis was done on the upper parts of the cores, which cover the record of environmental changes from the Migration Period (ca ad 375–568) to the present day. A total of 175 samples were analysed from the lakes, 40 from Jezioro Długie, 77 from Lubiąż and 58 from Paklicko Wielkie. Standard laboratory treatment was used for every sample (Berglund and Ralska-Jasiewiczowa 1986). During pollen analysis more than 500 AP or 1000 AP + NAP grains were counted per sample. Identification was made using the reference collection and the reference atlases (Beug 2004; Punt et al. 1976–2003) at the Institute of Archaeology, Nicolaus Copernicus University in Toruń. The resulting pollen counts were calculated and pollen diagrams drawn up and zoned using Polpal v. 2009 and later modifications (Walanus and Nalepka 1999). For every diagram, local pollen assemblage zones and the phases of more intensive levels of human activity (palynological settlement phases I-III) were identified. In view of the relatively vast lake areas, the pollen results reflect both local and regional vegetation around the individual sites (Sugita 1993; Sugita et al. 1999) and cover the land occupied by the groups of people who built and used the islands.
Results
General description of the material
The plant material contained over 90 grassland taxa from meadows and pastures, represented by over 12,200 remains (Table 2). The percentages of individual taxa differ from site to site and from phase to phase, although this may be related to the differences in the number of samples (Figs. 2, 3). The overall comparison of the results shows that grassland plants are better represented in Phase I in Chycina than II or III, whereas at Nowy Dworek both remains and taxa are better represented in Phase II than I. At Lubniewice, only Phase I has been preserved. The comparison of Phase I between the islands shows that grassland plants are better represented at Lubniewice than at Chycina or Nowy Dworek. DCA has demonstrated that the botanical composition of Phase I samples is similar from each island (pseudo-F = 1.32, df = 2, p = 0) (Fig. 4). On the right side of the diagram there are single samples containing mostly seeds of Juncus spp., while on the left are samples with mainly Poaceae remains.
Meadow and pasture communities
The phytosociological analysis of the plant communities represented by the macrofossils shows that the identified species relate to five of the currently distinguished classes of grassland communities (phytocoenoses) used as meadows and pastures (Molinio-Arrhenatheretea, Festuco-Brometea, Nardo-Callunetea, Scheuchzerio-Caricetea and Koelerio glaucae-Corynephoretea canescentis). They were represented in the plant remains from all three islands (Table 2), with small differences in the numbers of taxa and abundance of macroremains (Fig. 2). The latter may result from the accumulation of remains of taxa such as Juncus articulatus and J. conglomeratus/effusus.
The botanical composition of the samples indicates that the most characteristic class of grassland growing near the islands was Molinio-Arrhenatheretea, which represents meso-eutrophic, semi-natural and managed meadows and pastures on soils of average richness and moisture content. This type of grassland is represented by almost 60% of all the macroremains from each island and over 50 species. The proportion of Molinio-Arrhenatheretea species from every island is similar and high, over 50%. The Phase I remains from the islands are especially rich in these macroremains, with 1,216 from Chycina, 3,109 from Lubniewice and 776 from Nowy Dworek. However, these species are even better represented from Phase II at Nowy Dworek than Phase I, not only in terms of the number of remains (1,795), but also in terms of taxa (33) representing Molinio-Arrhenatheretea grassland. It is also worth noting that from Chycina in Phase II and III, only isolated remains of grassland plants were found, but almost all represent this class (Fig. 3). From every island, a similar set of taxa characteristic of Molinio-Arrhenatheretea was found. The greatest number of species was recorded from Lubniewice with 42 taxa, while there were 33 from Chycina and 37 from Nowy Dworek. Among 46 species found in the Phase I samples, 20 were found from all three islands. Among these are Scirpus sylvaticus, Rumex crispus, Ranunculus repens, Prunella vulgaris, Lythrum salicaria, Daucus carota, Plantago major, Filipendula ulmaria, Cerastium holosteoides and Ranunculus acris. At the same time, these were the most frequent species, with 15–23 remains in Phase I samples and also in the other phases. The great richness of Molinio-Arrhenatheretea taxa made it possible to sub-divide them into four phytosociological orders, which include Molinietalia caeruleae, Arrhenatheretalia, Plantaginetalia majoris and Trifolio fragiferae-Agrostietalia stoloniferae. Of these, Molinietalia caeruleae species were best represented from every island. The other orders were equally poorly represented from all islands. The order Plantaginetea majoris was hardly visible and from every island it was represented by isolated remains of only three taxa, Plantago major, Poa annua and Polygonum calcatum.
The next grassland class, Festuco-Brometea, a type of grassland growing on relatively dry soils, is the second best represented in terms of the number of taxa in the entire material, with a total of 13 species found, but only three of them, Acinos arvensis, Anthemis tinctoria and Petrorhagia prolifera occurred from all islands. Most of the taxa were represented by relatively few remains and only from Lubniewice. In 14 out of 19 Phase I samples, a fairly large number of Hypericum perforatum seeds were found (302.5). This was also found from Phase I at Chycina, and only from Phase II from Nowy Dworek. Two Festuco-Brometea species, Anthemis tinctoria and Thalictrum simplex, are characteristic of the order Festucetalia valesiacae, which is grassland of continental character growing on warm dry calcareous soils. Anthemis tinctoria was present from all sites, while T. simplex was found only from Nowy Dworek.
Class Nardo-Callunetea is represented by slightly fewer plants, with nine taxa. This represents semi-natural Nardus grasslands and seven of its species were found from Nowy Dworek, which were mostly from Phase II, while Chycina provided five species, only in Phase I, and another five from Lubniewice, where only Phase I was present. Almost all Nardo-Callunetea species were very poorly represented at each of the sites, except Calluna vulgaris, from which over 250 flowers and seeds were found from Lubniewice. and there were isolated finds from Chycina (Phase I) and Nowy Dworek (Phase II). Despite the relatively small number of Nardo Callunetea taxa, among them there were species characteristic of two orders, Nardetalia from Lubniewice and Nowy Dworek, and Calluno-Ulicetalia from all three islands.
Class Scheuchzerio-Caricetea grassland of marshy meadows and fens was distinguished on the basis of eight characteristic species which were mainly found in Phase I of all islands. With the exception of Juncus articulatus, other plants are represented by individual remains. The seeds of J. articulatus were abundant in the samples from Phase I in Chycina and Lubniewice (275 in two samples and 378 in four samples, respectively), but from Nowy Dworek there were far fewer remains from Phase I (26 in one sample) and Phase II (44 in four samples).
The final class, Koelerio glaucae-Corynephoretea canescentis, which is a type of grassland growing on sandy soils and dunes, was represented by four species. It is worth noting that one of these, Rumex acetosella, was plentiful from all phases and islands, with over 350 remains from Chycina and Nowy Dworek and over 200 from Lubniewice.
Discussion
General picture of the early medieval landscape around the islands
Changes in the environment were the inevitable result of the growing amount of settlement in central Europe in the early medieval period. Woodland clearance as well as land management connected with animal husbandry gradually shaped not only ruderal plant communities, but also semi-natural meadows and pastures too, which became an integral part of the human landscape (Dutoit et al. 2009; Overland and Hjelle 2013; Kuneš et al. 2015), and this was no different in Lubusz land. Pollen results from the three lakes show a distinct wooded phase (FL) covering the Migration Period and the beginning of the early Middle Ages, followed by a settlement phase (FO) with varying degrees of human pressure (Fig. 5). Before the development of local settlements and island enclosures there, the area was dominated by mixed deciduous woodland with a significant but variable proportion of Carpinus, Fagus, Quercus and some Fraxinus, Tilia, Ulmus and Corylus. Less fertile areas like glacial outwash plains must have been covered mainly by Pinus and Quercus woods and wet habitats by Alnus and riverside woods (Noryśkiewicz 2020).
There is evidence from the Jezioro Długie lake pollen results of early settlement at Chycina in the 7th century indicated by crops and grasslands. Cereal cultivation around Lubniewice and Nowy Dworek began in the mid 7th century, and meadows and pastures appeared there too. The pollen diagrams show increasing values of Poaceae, Plantago lanceolata and Ranunculus undiff., while Calluna vulgaris, Juniperus and Jasione montana appear sporadically. Importantly, the beginning of early medieval use of this area according to the pollen evidence started earlier than the construction of the timberwork of the islands. The mid 8th century marks an increase in the numbers and pollen values of herbaceous taxa connected with human activities (Fig. 5), which suggests that there was more intensive settlement when the islands were in use (Phase I). The pollen indicates growing numbers and pollen values of taxa characteristic of dry grasslands and pastures. Poaceae and Calluna vulgaris records increase, while Jasione montana forms a clear curve with a maximum above 1%. Juniperus is also sporadically present. This may suggest the development of dry grassland communities around the lakes and these are also in evidence from the macrofossil results. These grasslands may have spread as a result of heavy grazing by livestock on the light sandy soils around the sites. The results show not only the intensive development of agriculture with the cultivation of cereals, mainly Triticum aestivum and Panicum miliaceum as shown by the analysis of plant macroremains, but also of meadows and pastures. At the start of the 10th century, there was probably a change in the way the land was used, so that fields were cultivated further away from the islands, but meadows and pastures remained nearby. The widespread reduction in Alnus recorded by pollen diagrams, not only in northern Poland, but also in large areas of northern central Europe, occurred at the same time. The area of alder woods decreased around the 10th/11th centuries as a result of climate change (Stivrins et al. 2017; Latałowa et al. 2019) and this could coincide with Phase II of the islands. From the mid 11th century less settlement activity near the islands is shown by the pollen results, which is demonstrated by lower representation of meadow and pasture indicators and more alder (Fig. 5). A continuous rise of lake water level stables in the three lakes studied here was also observed (Chudziak et al. 2020).
Development and botanical composition of the meadows and pastures around Chycina, Nowy Dworek and Lubniewice
The archaeobotanical material from the islands is highly diversified, which represents a great variety of types of grassland which grew nearby. We should always remember that the recognition of past plant communities based on modern phytosociological classification is prone to inadequacies. These are mainly because of uncertainty about when the plant communities took their present form. It should also be remembered that there is no absolute phytosociological connection of a species to a particular plant community, which may be subject to modifications even today in different areas where the plants occur. A species may be recorded as growing within a vegetation community (phytocoenosis) which we might assign to a given syntaxonomic unit, such as an association, alliance, order or class, even though that community may not be its typical or associated unit (Zarzycki et al. 2002). Phytosociological systems also vary, and the species in the syntaxonomic units vary according to how the units are defined and what are the range limits of the individual syntaxa. If modern phytosociological classification is to be applied to ancient plant assemblages, it should be considered that it is not certain what past plant communities were like (Wasylikowa 1991). However, the use of phytosociology for archaeobotanical material can provide important clues for its interpretation. Assuming that the various plant communities in the past were at least similar to those of today, it is possible to arrange a long list of species into their probable communities and so describe the features of the past vegetation and the conditions of its occurrence (Rybníček and Rybníčková 1995; Kalis et al. 2006; Karg 2008; Gao et al. 2018; Pini et al. 2021).
The development of settlement activity in the microregions within our study area was connected with the individual islands. The selection of places for settlement was not a matter of chance, but was related to local catchments (Chudziak and Kaźmierczak 2020b). There was a more favourable climate in the whole of Europe between the 7th and 9th century and this would have encouraged economic development in the region (Büntgen et al. 2011). The water levels in the lakes at the time of island construction seem to have been considerably lower than today. The quite similar archaeobotanical results for every island indicate that well developed Molinio-Arrhenatheretea grasslands grew on the dry land around all the lakes at the time. It is interesting that similar results were obtained from Żółte, another early medieval semi-artificial island in Pomorze Zachodnie (west Pomerania), Poland (Pińska and Latałowa 2014). Grasslands of this type are particularly strongly related to human activities such as using meadows to mow for hay or as pastures for grazing livestock (Klarzyńska and Kryszak 2016; Kulik et al. 2016). Modern research shows that cattle, for example, prefer moist and productive habitats such as these grasslands provide (Putfarken et al. 2008). This is important, for archaeozoological results from this area indicate that cattle were important in the region in the early Middle Ages (Makowiecki 2020). They also show the presence of Cervus elaphus (red deer) and Capreolus capreolus (roe deer) there, which play a major part in the shaping of grassland communities (Bradshaw and Mitchell 1999; Schütz et al. 2003; Kuiters et al. 2005).
Another way in which these grassy areas near the islands may have been formed is by the clearance of deciduous woodlands from both dry and damp land (Pawłowski and Zarzycki 1977). The diversity of grassland species and the ample occurrence of Molinio-Arrhenatheretea plant remains suggest that this class developed rapidly and spread into various habitats. Moisture was undoubtedly the main diversifying factor, but the soil type must have been significant too (Table 2). A closer analysis of the species within the Molinio-Arrhenatheretea allows the taxa to be sub-divided into finer vegetational divisions, resulting in the identification of four orders and five associations. Since these plant communities must have developed at various distances from the wet ground near the lakes or watercourses, differences in the amount of soil moisture would have affected them too. The largest number of plants belong in the Molinietalia caeruleae order. It represents meadow and stream bank communities on moist or wet meso- and eutrophic soil (Maslovsky 2009; Grzegorczyk 2010; Zelnik and Čarni 2013). The presence of plants in this order may indicate the intensive use of this type of grassland near the islands. Within Molinietalia caeruleae are three related alliances, Calthion palustris, Filipendulion ulmariae and Molinion caeruleae. Calthion palustris today comprises nutrient rich, moist or wet meadows, on mineral soils which remain wet in these places even during droughts. These meadows are traditionally mown several times a year, and are of human origin. Species related to this alliance include Epilobium palustre, Caltha palustris, Cirsium oleraceum and Myosotis palustris. Another alliance, Filipendulion ulmariae, includes species such as Stachys palustris, Valeriana officinalis, Filipendula ulmaria and Lythrum salicaria, which may have grown in wet ground along the watercourses around the islands. Molinion caeruleae an alliance of nutrient poor grassland, mown annually, was also present near every lake and island as indicated by species including Linum catharticum and Molinia caerulea. Meadows and pastures of the order Arrhenatheretalia elatioris are also the outcome of intensive human activity. One of the conditions for their existence is traditional livestock grazing and these grasslands are now under protection (Chabuz et al. 2019). The presence of such types of grassland around the sites is shown by macrofossils of species such as Daucus carota, Achillea millefolium and Bromus hordeaceus. They are found on slightly moist mineral soils, but can also grow by watercourses (Grzegorczyk 2010), which were plentiful around the islands. This set of species includes plants characteristic of lower syntaxonomic units. Four species (Campanula patula, Galium mollugo, Knautia arvensis, Pastinaca sativa) are now assigned to the alliance Arrhenatherion elatioris, which are meadows mown repeatedly. Another alliance, Cynosurion, which forms fertile pastures, was represented by three species, Leontodon autumnalis, Ranunculus sardous and Trifolium repens. Archaeobotanical studies indicate that this type of pasture appeared in Europe during the Roman period (Knörzer 1975). The other two orders within the Molinio-Arrhenatheretea may have only occurred locally or were of little use for the people there. The order Trifolio fragiferae-Agrostietalia stoloniferae develops as grasslands in places which are periodically flooded, but it is tolerant of trampling and it is indicated by species like Potentilla reptans, Ranunculus repens, Agrostis stolonifera and Juncus glaucus. There were also nitrophilous communities composed of species resistant to mechanical damage, such as modern blanket grasslands of the Plantaginetalia majoris order, of which only two species possibly connected with such communities have been found, Plantago major and Poa annua. What these two orders have in common is their resistance to intensive use, including trampling. The considerable presence of Ranunculus repens, Rumex crispus and Plantago major remains is noteworthy, and these plants can also spread into ruderal communities and even crops.
Unlike the Molinio-Arrhenatheretea, the other classes of grasslands from the dry land near every island are much less well represented. This may be because they were less useful or there were fewer suitable habitats for them, thus limiting their range.
Several species within class Festuco-Brometea represent thermophilous grasslands developing on rather dry soils, as shown by macroremains including Acinos arvensis, Arabis hirsuta, Campanula glomerata, Centaurea scabiosa, Scabiosa columbaria and Stachys recta. Although the origins of Festuco-Brometea communities are mainly due to environmental conditions, human activities, including keeping grazing livestock, are also important factors (Bąba 2004; Dostálek and Frantík 2008; Pokorný et al. 2015; Büchler et al. 2020). Plant remains from the islands indicate that there could have been such conditions on the dry land around every island, but these grasslands were only present during part of the period when the sites were in use. At Nowy Dworek, Festuco-Brometea is recorded in Phase I, while the number of both taxa and seed remains rises in the following phases. In the case of Chycina, the evidence of this type of community is only in Phase I.
Another grassland class which also seems to have been present in our study area is Koelerio glaucae-Corynephoretea canescentis, which grows on sandy soil. The community is represented by only a few species. It is worth mentioning that in the pollen diagrams, the presence of occasional grains of Jasione montana, which is connected with this class, were recorded from horizons dating from before the formation of the islands. Numerous fruits of Rumex acetosella were found at each of the sites and it may have spread from this grassland to cultivated or ruderal habitats and arrived at the islands with the sediment which was brought there to strengthen the wooden structures (Badura et al. 2018).
Another important type of vegetation that must have developed near the islands includes communities such as the class Nardo-Callunetea grasslands and heath. They may have been formed from degraded Molinio-Arrhenatheretea grasslands, on disused and impoverished land or where woodlands had been cleared (Roo-Zielińska 2014). Macrofossils representing this class include Luzula campestris, L. multiflora, Viola canina and Calluna vulgaris. The latter, heather, grows on particularly poor soils, for instance sand, although sometimes also on nutrient-poor and acidic mires (Britton et al. 2003; Savukynienė et al. 2003). Calluna inflorescences and seeds were plentiful, especially at Lubniewice, which suggest that heather had been brought to the island on purpose (Badura and Noryśkiewicz 2020).
The list of taxa includes species of the class Scheuchzerio-Caricetea which are characteristic of fens and transitional mires, but in small amounts, which may indicate that communities of this kind did not play a major part in the early medieval landscape around the lakes. Only in the case of Lubniewice were more remains found, suggesting that the people had more use for this plant material and brought it to the island. It should be remembered, on the other hand, that such wetlands are easily transformed into Molinio-Arrhenatheretea meadows on a peaty substrate (Osadowski et al. 2019).
In some papers on meadows (such as Grynia and Kryszak 2003; Maslovsky 2009; Wyłupek et al. 2019), the class Phragmitetea, formed by reeds, bulrushes and tall sedges, is also included with meadow and pasture communities. Although we have not listed taxa from this group, a considerable number of such remains have been identified in the material discussed here (Table 2). Some of them must have grown along the lake shores or local watercourses. When the water table was particularly low, such communities may have been colonised by plants of wet grassland. A reed belt, on the other hand, certainly formed around the islands and stabilised them.
The usefulness of meadows and pastures for the local human communities
In the past, meadows and pastures were the main source of grazing and fodder for livestock. Therefore, the emergence of such grassland communities may be linked to animal husbandry (van Geel et al. 2003; Putfarken et al. 2008; Poschold et al. 2009; Florenzano 2019), although we have to remember that it would have been a gradual process. Archaeozoological data for the early Middle Ages in this area show that cattle, pigs, goats, sheep and horses were kept. At the same time, on the islands themselves, numerous fragments of animal bones were found, which were the post-consumption remains from food (Makowiecki 2020). Undoubtedly, livestock needed suitable sources of fodder from various plant communities, including grasslands. Naturally, the surrounding meadows and pastures may also have been influenced by wildlife such as deer (Gebert and Verheyden-Tixier 2001; DeVoe et al. 2019).
The plant remains from the islands include a number of important forage plants, which may indicate that herbaceous fodder such as hay was brought to the sites. These plants are mainly connected with the class Molinio-Arrhenatheretea and the order Molinietalia caeruleae. The legumes Trifolium repens and T. pratense were of particular nutritional value, as well as grasses such as Poa pratensis and P. annua (Miller and Thompson 2007; Komainda et al. 2019). Valuable grazing herbs like Taraxacum officinale, Plantago lanceolata and Achillea millefolium were also abundant. They made the fodder more nutritious (Putfarken et al. 2008; Pirhofer‐Walzl et al. 2011; Supek et al. 2016). The accumulation of goat or sheep dung at Nowy Dworek (Phases I/II) is the only suggestion that live animals were kept on the islands. The pollen analysis of this dung showed that the goats or sheep had eaten tree leaves and inflorescences as well as herbaceous plants (Badura et al. 2018; Badura and Noryśkiewicz 2020). Some dung contained mostly pollen of herbaceous plants such as Knautia arvensis, Plantago lanceolata, Rumex, Ranunculus and Poaceae, which may represent associations within Molinio-Arrhenatheretea. Pollen of Apiaceae, Brassicacaeae and Cichorioideae, possibly also from grassland plants, was also frequent in the goat or sheep dung, but in variable amounts.
The results include some plants with no special nutritional value, like Bromus hordeaceus, Galium mollugo, Leontodon autumnalis and Angelica sylvestris, which can be eaten by animals in small amounts or at certain stages of plant development. Some plants are unpleasant or poisonous (when fresh), such as Ranunculus acris, R. repens and Caltha palustris (Siminska et al. 2009; Grzegorczyk 2010; Nawara 2015).
The remains of Campanula patula, C. glomerata, Centaurea jacea, C. scabiosa, Calluna vulgaris, Taraxacum officinale, Senecio jacobaea, Caltha palustris and Anthemis tinctoria were also found, which are insect pollinated flowers, one of the features of traditional meadows and pastures (Steffan-Dewenter 2003; Meyer et al. 2017; Tonietto and Larkin 2018). Therefore, beekeeping may have developed as a side effect of the formation of such communities around the islands. One of the ways in which plants adapt to entomophily is through bright, colourful flowers (McCall and Primack 1992; Gumbert et al. 1999; Hoyle et al. 2018). Showy purple or pink flowers are characteristic of species like Campanula patula, C. glomerata, Centaurea jacea, C. scabiosa, Scabiosa columbaria or Calluna vulgaris (Van Treuren et al. 1994; Steffan-Dewenter and Tscharntke 2000; Denisow and Wrzesień 2015). Other species such as Taraxacum officinale, Senecio jacobaea and Caltha palustis have yellow-coloured flowers (Andersson 1996; Arnold et al. 2009; Lázaro and Totland 2010). Archaeologists suggest that the islands may have played a sacred role, so the plants mentioned above could have been used as decorations during the festivities held there. It may also be assumed that the nearby meadows and pastures were a source of medicinal plants for the local people. Hypericum perforatum, Plantago major, Filipendula ulmaria, Centaurea scabiosa, Angelica sylvestris or Achillea millefolium may have been gathered for this purpose (Grieve 1971).
Conclusions
The continuation of research on the samples obtained from the semi-artificial islands in the three lakes has provided new evidence of the presence and role of local grasslands in Ziemia Lubuska (Lubusz land) during the early Middle Ages. It may be assumed that in the 9th and 10th centuries these unusual sites were constructed near already well developed meadows and pastures. These particular types of grassland communities appeared around the 7th century with the clearance of woodland and establishment of local settlements. It seems that the grasslands were used by the people who built and used the islands at Nowy Dworek, Chycina and Lubniewice. The main types of grassland, according to the remains studied here, include meadows or pastures on slightly moist to wet soil. These, together with a significant number of species from other communities resilient to trampling and water level fluctuations, provide an image of strong economic uses of the areas around the lakes and related watercourses. Grasslands on drier, sandier, soil were also indicated by the plant remains, although these must have been located some distance away from the lakes themselves, but still within reach of the island users. The pastures would have been extensively grazed during spring and summer and haymaking cannot be excluded, either. Winter fodder was probably obtained from trees. The grasslands may have provided not only a source of fodder for farm animals, but also a source of medicinal and ornamental plants, as well as a grazing place for deer and other herbivores.
This study of the early medieval meadows and pastures represents only one part of the work on the natural history of this area. Another aspect is the rich collection of remains of segetal and ruderal weeds that will allow us to study the changes that occurred in the landscape which was created by the people who built and used the islands.
References
Akeret Ö, Kiefer S, Kühn M, Rentzel P, Rösch M, Wick L (2018) The buried medieval pasture of Onoldswil (Niederdorf BL, Switzerland, AD 1295): an example of a well preserved palaeobiocoenosis. Veget Hist Archaeobot 27:137–149. https://doi.org/10.1007/s00334-017-0623-1
Andersson S (1996) Floral display and pollination success in Senecio jacobaea (Asteraceae): interactive effects of head and corymb size. Am J Bot 83:71–75. https://doi.org/10.1002/j.1537-2197.1996.tb13876.x
Arnold SEJ, Savolainen V, Chittka L (2009) Flower colours along an alpine altitude gradient, seen through the eyes of fly and bee pollinators. Arthropod-Plant Inte 3:27–43. https://doi.org/10.1007/s11829-009-9056-9
Badura M, Noryśkiewicz AM (2020) Wyspy jeziorne jako element wczesnośredniowiecznej struktury osadniczej w świetle danych archeobotanicznych. In: Chudziak W, Kaźmierczak R (eds) Człowiek na pograniczu. Na peryferiach Civitas Schinesghe. T. I:2. Ziemia lubuska. Wydawnictwo Naukowe Uniwersytetu Mikołaja Kopernika, Toruń, pp 107–143
Badura M, Noryśkiewicz AM, Chudziak W, Kaźmierczak R (2018) Environmental context and the role of plants at the early medieval artificial island in the lake Paklicko Wielkie, Nowy Dworek, western Poland. Veget Hist Archaeobot 27:99–110. https://doi.org/10.1007/s00334-017-0617-z
Bąba W (2004) The species composition and dynamics in well-preserved and restored calcareous xerothermic grasslands (South Poland). Biologia, Bratislava 59:447–456
Behre K-E, Jacomet S (1991) The ecological interpretation of archaeobotanical data. In: van Zeist W, Wasylikowa K, Behre K-E (eds) Progress in Old World palaeoethnobotany. Balkema, Rotterdam, pp 81–108
Berglund BE, Ralska-Jasiewiczowa M (1986) Pollen analysis and pollen diagrams. In: Berglund BE (ed) Handbook of Holocene palaeoecology and palaeohydrology. Wiley, Chichester, pp 455–484
Beug H-J (2004) Leitfaden der Pollenbestimmung für Mitteleuropa und angrenzende Gebiete. Pfeil, München
Bleile R (2010) Central sites on the periphery? The development of Slavonic ramparts on islands in freshwater lakes in Northern Germany (8th–12th century AD). In: Kiel Graduate School (eds) Landscapes and human development: the contribution of European archaeology. Proceedings of the International Workshop "Socio-Environmental dynamics over the last 12,000 years: the creation of landscapes (1st-4th April 2009). Habelt, Bonn, pp 285–295
Boch S, Becker T, Deák B, Dengler J, Wagner V (2020) Traditional land use, management and biodiversity of European semi-natural grasslands - Editorial to the 15th EDGG Special Feature. Tuexenia 40:401–407. https://doi.org/10.14471/2020.40.026
Bradshaw R, Mitchell FJG (1999) The palaeoecological approach to reconstructing former grazing-vegetation interactions. For Ecol Manage 120:3–12. https://doi.org/10.1016/S0378-1127(98)00538-6
Britton A, Marrs R, Pakeman R, Carey P (2003) The influence of soil-type, drought and nitrogen addition on interactions between Calluna vulgaris and Deschampsia flexuosa: implications for heathland regeneration. Plant Ecol 166:93–105. https://doi.org/10.1023/A:1023248407408
Büchler M-O, Billeter R, Dengler J (2020) Optimal site conditions for dry grasslands of high conservation value in the canton of Zurich, Switzerland. Tuexenia 40:527–546. https://doi.org/10.14471/2020.40.021
Büntgen U, Tegel W, Nicolussi K et al (2011) 2500 years of European climate variability and human susceptibility. Science 331:578–582. https://doi.org/10.1126/science.1197175
Cappers RTJ, Bekker RM, Jans JEA (2006) Digital seed atlas of the Netherlands. Barkhuis Publishing, Eelde
Chabuz W, Kulik M, Sawicka-Zugaj W et al (2019) Impact of the type of use of permanent grasslands areas in mountainous regions on the floristic diversity of habitats and animal welfare. Glob Ecol Conserv 19:e00629. https://doi.org/10.1016/j.gecco.2019.e00629
Chudziak W, Kaźmierczak R (eds) (2020a) Człowiek na pograniczu. Na peryferiach Civitas Schinesghe. T. I:1. Ziemia lubuska. Wydawnictwo Naukowe Uniwersytetu Mikołaja Kopernika, Toruń
Chudziak W, Kaźmierczak R (2020b) Wybrane mikroregiony - analiza struktur osadniczych pod kątem problemu kształtowania się pogranicza i przebiegu szlaków komunikacyjnych. In: Chudziak W, Kaźmierczak R (eds) Człowiek na pograniczu. Na peryferiach Civitas Schinesghe. T. I:2. Ziemia lubuska. Wydawnictwo Naukowe Uniwersytetu Mikołaja Kopernika, Toruń, pp 61–91
Chudziak W, Kaźmierczak R, Kramkowski M (2020) Problem wahań poziomu wody w akwenach lubuskich w okresie wczesnego średniowiecza. In: Chudziak W, Kaźmierczak E (eds) Człowiek na pograniczu. Na peryferiach Civitas Schinesghe. T.I:2, Ziemia lubuska. Wydawnictwo Naukowe Uniwersytetu Mikołaja Kopernika, Toruń, pp 31–37
Denisow B, Wrzesień M (2015) The habitat effect on the diversity of pollen resources in several Campanula spp. – an implication for pollinator conservation. J Apicult Res 54:62–71. https://doi.org/10.1080/00218839.2015.1030243
DeVoe JD, Proffitt KM, Mitchell MS, Jourdonnais CS, Barker KJ (2019) Elk forage and risk tradeoffs during the fall archery season. J Wildl Manage 83:801–816. https://doi.org/10.1002/jwmg.21638
Dostálek J, Frantík T (2008) Dry grassland plant diversity conservation using low-intensity sheep and goat grazing management: case study in Prague (Czech Republic). Biodivers Conserv 17:1,439–1,454. https://doi.org/10.1007/s10531-008-9352-1
Dutoit T, Thinon M, Talon B, Buisson E, Alard D (2009) Sampling soil wood charcoals at a high spatial resolution: a new methodology to investigate the origin of grassland plant communities. J Veg Sci 20:349–358. https://doi.org/10.1111/j.1654-1103.2009.05403.x
Ellenberg H, Weber HE, Düll R, Wirth W, Werner W, Paulißen D (1992) Zeigerwerte von Pflanzen in Mitteleuropa. (Scripta Geobotanica 18) Golze, Göttingen
Florenzano A (2019) The history of pastoral activities in S Italy inferred from palynology: a long-term perspective to support biodiversity awareness. Sustainability 11:404. https://doi.org/10.3390/su11020404
Gao G, Jie D, Wang Y et al (2018) Do soil phytoliths accurately represent plant communities in a temperate region? A case study of Northeast China. Veget Hist Archaeobot 27:753–765. https://doi.org/10.1007/s00334-018-0670-2
Gebert C, Verheyden-Tixier H (2001) Variations of diet composition of Red Deer (Cervus elaphus L.) in Europe. Mammal Rev 31:189–201. https://doi.org/10.1111/j.1365-2907.2001.00090.x
Greig J (1988) Some evidence of the development of grassland plant communities. In: Jones M (ed) Archaeology and the flora of the British Isles. Oxford University Committee for Archaeology, Oxford, pp 39–54
Grieve M (1971) A modern herbal. Dover Publications Inc, New York
Grynia M, Kryszak A (2003) Zbiorowiska roślinne łąk i pastwisk w fitosocjologicznym systemie klasyfikacyjnym (Plant communities of meadows and pastures in phytosociological classification system). Biuletyn Instytutu Hodowli i Aklimatyzacji Roślin 225:211–220
Grzegorczyk S (2010) Rośliny zbiorowisk trawiastych. Wydawnictwo Uniwersytet Warmińsko-Mazurski, Olsztyn
Gumbert A, Kunze J, Chittka L (1999) Floral colour diversity in plant communities, bee colour space and a null model. Proc R Soc Lond B 266:1711–1716. https://doi.org/10.1098/rspb.1999.0836
Hájková P, Jamrichová E, Petr L et al (2018) Persistence of a vegetation mosaic in a peripheral region: could turbulent medieval history disrupt Holocene continuity of extremely species-rich grasslands? Veget Hist Archaeobot 27:591–610. https://doi.org/10.1007/s00334-017-0660-9
Hejcman M, Hejcmanová P, Pavlů V, Beneš J (2013) Origin and history of grasslands in Central Europe – a review. Grass Forage Sci 68:345–363. https://doi.org/10.1111/gfs.12066
Hjelle KL (1998) Herb pollen representation in surface moss samples from mown meadows and pastures in western Norway. Veget Hist Archaeobot 7:79–96. https://doi.org/10.1007/BF01373926
Hodgson JG, Halstead P, Wilson PJ, Davis S (1999) Functional interpretation of archaeobotanical data: Making hay in the archaeological record. Veget Hist Archaeobot 8:261–271. https://doi.org/10.1007/BF01291778
Hoyle H, Norton B, Dunnett N, Richards JP, Russell JM, Warren P (2018) Plant species or flower colour diversity? Identifying the drivers of public and invertebrate response to designed annual meadows. Landsc Urban Plan 180:103–113. https://doi.org/10.1016/j.landurbplan.2018.08.017
Kaczmarek M (2019) Urnfields in the middle Oder basin – a perspective of a Lubusz-Greater Polish territorial community. Praehist Z 94:379–413. https://doi.org/10.1515/pz-2019-0017
Kalis AJ, van der Knaap WO, Schweizer A, Urz R (2006) A three thousand year succession of plant communities on a valley bottom in the Vosges Mountains, NE France, reconstructed from fossil pollen, plant macrofossils, and modern phytosociological communities. Veget Hist Archaeobot 15:377–390. https://doi.org/10.1007/s00334-006-0065-7
Karg S (2008) Direct evidence of heathland management in the early Bronze Age (14th century B.C.) from the grave-mound Skelhøj in western Denmark. Veget Hist Archaeobot 17:41–49. https://doi.org/10.1007/s00334-007-0109-7
Klarzyńska A, Kryszak A (2016) Useful plants in grassland communities of river valleys in Wielkopolska. J Res Appl Agric Eng 61:224–228
Knörzer K-H (1975) Entstehung und Entwicklung der Grünlandvegetation im Rheinland. Decheniana 127:195–214
Kobyliński Z (2014) Starosiedle w Ziemi Lubuskiej: osadnictwo starożytne i wczesnośredniowieczne (Starosiedle in the Lubusz Land: Prehistoric and Early Medieval Settlement). Fundacja Res Publica Multiethnica, Warszawa
Komainda M, Küchenmeister K, Küchenmeister F, Breitsameter L, Wrage-Mönnig N, Kayser M, Isselstein J (2019) Forage legumes for future dry climates: lower relative biomass losses of minor forage legumes compared to Trifolium repens under conditions of periodic drought stress. J Agron Crop Sci 205:460–469. https://doi.org/10.1111/jac.12337
Koszałka J (2014) Źródła archeobotaniczne do rekonstrukcji uwarunkowań przyrodniczych oraz gospodarczych grodu w Łęczycy. In: Grygiel R, Jurek T (eds) Początki Łęczycy. T. I. Archeologia środowiskowa średniowiecznej Łęczycy. Muzeum Archeologiczne i Etnograficzne, Łódź, pp 191–241
Körber-Grohne U (1991) Identification key for subfossil Gramineae fruits. (Probleme der Kustenforschung im sudliche Nordeegebiet 18) Lax, Hildesheim
Kreuz A, Schäfer E (2011) Weed finds as indicators for the cultivation regime of the early Neolithic Bandkeramik culture? Veget Hist Archaeobot 20:333–348. https://doi.org/10.1007/s00334-011-0294-2
Kuiters AT, Groot Bruinderink GWTA, Lammertsma DR (2005) Facilitative and competitive interactions between sympatric cattle, red deer and wild boar in Dutch woodland pastures. Acta Theriol 50:241–252. https://doi.org/10.1007/BF03194487
Kulik M, Baryła R, Warda M, Stamirowska-Krzaczek E (2016) Vegetation changes of the Molinio Arrhenatheretea class in the Bystra valley, eastern Poland. Acta Agrobot 69:1689. https://doi.org/10.5586/aa.1689
Kuneš P, Svobodová-Svitavská H, Kolář J et al (2015) The origin of grasslands in the temperate forest zone of east-central Europe: long-term legacy of climate and human impact. Quat Sci Rev 116:15–27. https://doi.org/10.1016/j.quascirev.2015.03.014
Kurnatowska Z, Łosińska A (1996) Perspektywy badań nad wczesnym średniowieczem Ziemi Lubuskiej. In: Leciejewicz L, Gringmuth-Dallmer E (eds) Człowiek a środowisko w środkowym i dolnym Nadodrzu. Spotkania Bytomskie 2. Polska Akademia Nauk, Instytut Archeologii i Etnologii, Wrocław, pp 161–176
Küster H, Keenleyside C (2009) The origin and use of agricultural grasslands in Europe. In: Veen P, Jefferson R, de Smidt J, van der Straaten J (eds) Grasslands in Europe of high nature value. KNNV Publishing, Zeist, pp 9–14
Latałowa M (1999) Palaeoecological reconstruction of environmental condition and economy in early medieval Wolin. Acta Palaeobot 39:183–271
Latałowa M, Święta-Musznicka J, Słowiński M et al (2019) Abrupt Alnus population decline at the end of the first millennium CE in Europe – the event ecology, possible causes and implications. Holocene 29:1335–1349. https://doi.org/10.1177/0959683619846978
Lázaro A, Totland Ø (2010) Population dependence in the interactions with neighbors for pollination: a field experiment with Taraxacum officinale. Am J Bot 97:760–769. https://doi.org/10.3732/ajb.0900263
Makowiecki D (2020) Fauna pogranicza wielkopolsko-lubuskiego we wczesnym średniowieczu w świetle analizy archeozoologicznej. In: Chudziak W, Kaźmierczak R (eds) Człowiek na pograniczu. Na peryferiach Civitas Schinesghe. T. I:2. Ziemia lubuska. Wydawnictwo Naukowe Uniwersytetu Mikołaja Kopernika, Toruń, pp 145–170
Maslovsky O (2009) Grasslands of the Narochansky National Park, Belarus. In: Veen P, Jefferson R, de Smidt J, van der Straaten J (eds) Grasslands in Europe of high nature value. KNNV Publishing, Zeist, pp 210–217
Matuszkiewicz W (2008) Przewodnik do oznaczania zbiorowisk roślinnych Polski. Wydawnictwo Naukowe PWN, Warszawa
McCall C, Primack RB (1992) Influence of flower characteristics, weather, time of day, and season on insect visitation rates in three plant communities. Am J Bot 79:434–442. https://doi.org/10.1002/j.1537-2197.1992.tb14571.x
Menotti F (2012) Wetland archaeology and beyond: theory and practice. Oxford University Press, Oxford
Meyer S, Unternährer D, Arlettaz R, Humbert J-Y, Menz MHM (2017) Promoting diverse communities of wild bees and hoverflies requires a landscape approach to managing meadows. Agric Ecosyst Environ 239:376–384. https://doi.org/10.1016/j.agee.2017.01.037
Miller SM, Thompson RP (2007) Seasonal patterns of diet composition, herbage intake and digestibility limit the performance of cattle grazing native pasture in the Falkland Islands. Grass Forage Sci 62:135–144. https://doi.org/10.1111/j.1365-2494.2007.00573.x
Mirek Z, Piękoś-Mirkowa H, Zając A, Zając M (2002) Flowering plants and pteridophytes of Poland: a checklist. W. Szafer Institute of Botany, Polish Academy of Science, Kraków
Morrison I (1985) Landscape with lake dwellings: the crannogs of Scotland. Edinburgh University Press, Edinburgh
Nawara Z (2015) Rośliny łąkowe. Multico Oficyna Wydawnicza, Warszawa
Noryśkiewicz AM (2014) Vegetation and settlement history as reflected by pollen analysis. In: Chudziak W, Kaźmierczak R (eds) The Island in Żółte on Lake Zarańskie. Nicolaus Copernicus University, Stanisław Staszic Pomeranian Library, Toruń, Szczecin, Early Medieval Gateway into West Pomerania, pp 49–60
Noryśkiewicz AM (2020) Palinologiczny obraz wczesnośredniowiecznego osadnictwa pogranicza wielkopolsko-lubuskiego (jeziora Paklicko Wielkie, Długie i Lubiąż). In: Chudziak W, Kaźmierczak R (eds) Człowiek na pograniczu. Na peryferiach Civitas Schinesghe. T. I:2. Ziemia lubuska. Wydawnictwo Naukowe Uniwersytetu Mikołaja Kopernika, Toruń, pp 93–105
Osadowski Z, Drzymulska D, Dobrowolski R, Mazurek M (2019) Current state and vegetation history of spring-fed fens in Western Pomerania (Northern Poland): a case study of the Chociel River valley. Wetl Ecol Manage 27:23–38. https://doi.org/10.1007/s11273-018-9640-9
Overland A, Hjelle KL (2013) Pollen analysis in the context of clearance cairns from boreal forests – a reflection of past cultivation and pastoral farming. J Archaeol Sci 40:1029–1041. https://doi.org/10.1016/j.jas.2012.08.031
Pawłowski B, Zarzycki K (1977) Zespoły łąkowe i wrzosowiskowe. In: Szafer W, Zarzycki K (eds) Szata roślinna Polski, vol 1. Państwowe Wydawnictwo Naukowe, Warszawa, pp 338–351
Pärtel M, Bruun HH, Sammul M (2005) Biodiversity in temperate European grasslands: origin and conservation. Grassl Sci Europe 10:1–14
Pini R, Ravazzi C, Comolli R et al (2021) Life on a hilltop: vegetation history, plant husbandry and pastoralism at the dawn of Bergamo-Bergomum (northern Italy, 15th to 7th century BC). Veget Hist Archaeobot 30:525–553. https://doi.org/10.1007/s00334-020-00802-1
Pińska K, Latałowa M (2014) Plant remains. In: Chudziak W, Kaźmierczak R (eds) The island in Żółte on Lake Zarańskie. Nicolaus Copernicus University, Stanisław Staszic Pomeranian Library, Toruń, Szczecin, Early medieval gateway into West Pomerania, pp 367–560
Pirhofer-Walzl K, Søegaard K, Høgh-Jensen H, Eriksen J, Sanderson MA, Rasmussen J, Rasmussen J (2011) Forage herbs improve mineral composition of grassland herbage. Grass Forage Sci 66:415–423. https://doi.org/10.1111/j.1365-2494.2011.00799.x
Pokorný P, Boenke N, Chytráček M et al (2006) Insight into the environment of a pre-Roman Iron Age hillfort at Vladař, Czech Republic, using a multi-proxy approach. Veget Hist Archaeobot 15:419–433. https://doi.org/10.1007/s00334-006-0064-8
Pokorný P, Chytrý M, Juřičková L, Sádlo J, Novák J, Ložek V (2015) Mid-Holocene bottleneck for central European dry grasslands: did steppe survive the forest optimum in northern Bohemia, Czech Republic? Holocene 25:716–726. https://doi.org/10.1177/0959683614566218
Poschold P, Baumann A, Karlik P (2009) Origin and development of grasslands in Central Europe. In: Veen P, Jefferson R, de Smidt J, van der Straaten J (eds) Grasslands in Europe of high nature value. KNNV Publishing, Zeist, pp 15–24
Punt W, Blackmore S, Clarke GCS, Hoen PP, Stafford PJ (eds) (1976–2003) The Northwest European pollen flora, vol 1–8. Elsevier, Amsterdam
Putfarken D, Dengler J, Lehmann S, Härdtle W (2008) Site use of grazing cattle and sheep in a large-scale pasture landscape: a GPS/GIS assessment. Appl Anim Behav Sci 111:54–67. https://doi.org/10.1016/j.applanim.2007.05.012
Qin F, Bunting MJ, Zhao Y, Li Q, Cui Q, Ren W (2020) Relative pollen productivity estimates for alpine meadow vegetation, northeastern Tibetan Plateau. Veget Hist Archaeobot 29:447–462. https://doi.org/10.1007/s00334-019-00751-4
Ralska-Jasiewiczowa M, Madeyska E, Mierzeńska M (2006) Vegetational changes in the montane grassland zone of the High Bieszczady mountains (southeast Poland) during the last millennium-pollen records from deposits in hanging peat bogs. Veget Hist Archaeobot 15:391–401. https://doi.org/10.1007/s00334-006-0057-7
Roo-Zielińska E (2014) Wskaźniki ekologiczne zespołów roślinnych Polski. Wydawnictwo Akademickie SEDNO, Warszawa
Rybníček K, Rybníčková E (1995) Palaeoecological and phytosociological reconstruction of precultural vegetation in the Bílý Kříž area, the Moravskoslezské Beskydy Mts, Slovak Republic. Veget Hist Archaeobot 4:161–170. https://doi.org/10.1007/BF00203934
Savukynienė N, Moe D, Ūsaitytė D (2003) The occurrence of former heathland vegetation in the coastal areas of the south-east Baltic sea, in particular Lithuania: a review. Veget Hist Archaeobot 12:165–175. https://doi.org/10.1007/s00334-003-0008-5
Schütz M, Risch AC, Leuzinger E, Krüsi BO, Achermann G (2003) Impact of herbivory by red deer (Cervus elaphus L.) on patterns and processes in subalpine grasslands in the Swiss National Park. For Ecol Manag 181:177–188. https://doi.org/10.1016/S0378-1127(03)00131-2
Siminska E, Bernacka H, Grabowicz M (2009) Zioła w żywieniu zwierząt, z uwzględnieniem owiec. (Zeszyty naukowe nr 252) Zootechnika 37:89–97
Steffan-Dewenter I (2003) Importance of habitat area and landscape context for species richness of bees and wasps in fragmented orchard meadows. Conserv Biol 17:1036–1044. https://doi.org/10.1046/j.1523-1739.2003.01575.x
Steffan-Dewenter I, Tscharntke T (2000) Resource overlap and possible competition between honey bees and wild bees in central Europe. Oecologia 122:288–296. https://doi.org/10.1007/s004420050034
Stivrins N, Buchan MS, Disbrey HR et al (2017) Widespread, episodic decline of alder (Alnus) during the medieval period in the boreal forest of Europe. J Quat Sci 32:903–907. https://doi.org/10.1002/jqs.2984
Sugita S (1993) A model of pollen sources area for an entire lake surface. Quat Res 39:239–244. https://doi.org/10.1006/QRES.1993.1027
Sugita S, Gaillard M-J, Broström A (1999) Landscape openness and pollen records: a simulation approach. Holocene 9:409–421. https://doi.org/10.1191/095968399666429937
Supek Š, Pavlů V, Pavlů L, Gaisler J, Hejcman M, Ludvíková V, Mikulka J (2016) Effects of long-term grazing management on dandelion (Taraxacum officinale) in Agrostis capillaris grassland. Grass Forage Sci 72:516–523. https://doi.org/10.1111/gfs.12260
Święta-Musznicka J, Latałowa M (2016) From wetland to commercial centre: the natural history of Wyspa Spichrzów (“Granary Island”) in medieval Gdańsk, northern Poland. Veget Hist Archaeobot 25:583–599. https://doi.org/10.1007/s00334-016-0578-7
Tonietto RK, Larkin DJ (2018) Habitat restoration benefits wild bees: a meta-analysis. J Appl Ecol 55:582–590. https://doi.org/10.1111/1365-2664.13012
Van Geel B, Buurman J, Brinkkemper O, Schelvis J, Aptroot A, van Reenen G, Hakbijl T (2003) Environmental reconstruction of a Roman Period settlement site in Uitgeest (The Netherlands), with special reference to coprophilous fungi. J Archaeol Sci 30:873–883. https://doi.org/10.1016/S0305-4403(02)00265-0
Van Treuren R, Bijlsma R, Ouborg NJ, Kwak MM (1994) Relationships between plant density, outcrossing rates and seed set in natural and experimental populations of Scabiosa columbaria. J Evol Biol 7:287–302. https://doi.org/10.1046/j.1420-9101.1994.7030287.x
Walanus A, Nalepka D (1999) POLPAL. Program for counting pollen grains, diagrams plotting and numerical analysis. Acta Palaeobot (suppl) 2:659–661
Wasylikowa K (1978) Plant remains from early and late Medieval time found on the Wawel Hill in Cracow. Acta Palaeobot 19:115–220
Wasylikowa K (1991) Roślinność Wzgórza Wawelskiego we wczesnym i późnym średniowieczu na podstawie badań paleobotanicznych. Studia Do Dziejów Wawelu 5:93–131
Wyłupek T, Klikocka H, Kulik M, Lipińska H, Gawryluk A (2019) Floristic diversity and use value of phytocoenoses in a section of the Wieprz river valley in Roztocze National Park for feeding the konik polski horse. Agron Sci 4:124–134. https://doi.org/10.24326/as.2019.4.9
Zarzycki K, Trzcińska-Tacik H, Różański W, Szeląg Z, Wołek J, Korzeniak U (2002) Ecological indicator values of vascular plants of Poland (Biodiversity of Poland 2) W. Szafer Institute of Botany, Polish Academy of Sciences, Kraków
Zelnik I, Čarni A (2013) Plant species diversity and composition of wet grasslands in relation to environmental factors. Biodivers Conserv 22(2):179–2192. https://doi.org/10.1007/s10531-013-0448-x
Acknowledgements
The paper has been written as part of the statutory research at the Department of Plant Ecology, University of Gdańsk (531-D040-D581-21) and financed by the Research Project of Young Scientists (Projekt Badawczy Młodych Naukowców), Faculty of Biology, University of Gdańsk (539-D040-B855-21). The field work has been carried out as part of the archaeological project of the Polish Ministry of Science and Higher Education 11H 12 0526 81. The authors wish to thank Wojciech Chudziak (archaeology) and Jacek Herbich (botany) for their helpful comments and suggestions. We would like to thank Julian Wiethold and one anonymous reviewer for their fruitful advice and encouraging comments on the manuscript.
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Maciejewska, K., Badura, M. & Noryśkiewicz, A.M. Botanical composition of meadows and pastures and their role in the functioning of early medieval semi-artificial lake islands in Ziemia Lubuska (Lubusz land), western Poland. Veget Hist Archaeobot 31, 579–594 (2022). https://doi.org/10.1007/s00334-022-00877-y
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DOI: https://doi.org/10.1007/s00334-022-00877-y