Animals may have beaten upright plants to land.
References
MacNaughton, R. B et al. First steps on land: Arthropod trackways in Cambrian-Ordovician eolian sandstone, southeastern Ontario, Canada. Geology 30, 391 - 394 (2002).
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Clarke, T. Oldest fossil footprints on land. Nature (2002). https://doi.org/10.1038/news020429-2
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DOI: https://doi.org/10.1038/news020429-2
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