Early multicellular organisms had two distinct types of photoreceptor cells, apparently with different functions. How these cells combined to form modern eyes turns out to be a complicated story.
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Lacalli, T. Light on ancient photoreceptors. Nature 432, 454–455 (2004). https://doi.org/10.1038/432454a
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DOI: https://doi.org/10.1038/432454a
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