In horses it is due to homozygosity for the d allele; actually this color is expressed when the genetic constitution of the animals is AAbbCCdd.
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(2008). Chestnut Color. In: Encyclopedia of Genetics, Genomics, Proteomics and Informatics. Springer, Dordrecht. https://doi.org/10.1007/978-1-4020-6754-9_2810
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DOI: https://doi.org/10.1007/978-1-4020-6754-9_2810
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