A re-assessment of the taxonomy, palaeobiology and taphonomy of the rangeomorph organism Hapsidophyllas flexibilis from the Ediacaran of Newfoundland, Canada Rod S. TaylorJack J. MatthewsDuncan McIlroy Research Paper 27 March 2021 Pages: 187 - 207
Houcaris gen. nov. from the early Cambrian (Stage 3) Chengjiang Lagerstätte expanded the palaeogeographical distribution of tamisiocaridids (Panarthropoda: Radiodonta) Yu WuDongjing FuXingliang Zhang Research Paper 28 May 2021 Pages: 209 - 221
Timsaloproetus alissae sp. nov. (Trilobita: Proetidae) from the lower Devonian of southern Morocco Allart van ViersenFrederik Lerouge Research Paper 06 March 2021 Pages: 223 - 230
Symbiotic worms in the inner aragonitic layer of Leptodesma (Bivalvia) from the Přídolí (Upper Silurian) of Saaremaa Island, Estonia Olev VinnMark A. WilsonUrsula Toom Research Paper 27 March 2021 Pages: 231 - 236
Upper Frasnian Tornoceratidae (Ammonoidea) from the Sand Formation (Bergisch Gladbach-Paffrath Syncline, Rhenish Massif) Till SöteRalph Thomas BeckerJürgen Bockwinkel Research Paper Open access 28 May 2021 Pages: 237 - 273
Cambrian brachiopods from the area of the former Semipalatinsk nuclear-testing site, Chingiz Ranges, Kazakhstan Leonid E. PopovOlga I. NikitinaGappar Kh. Ergaliev Research Paper 10 February 2021 Pages: 275 - 290
The oldest centrosaurine: a new ceratopsid dinosaur (Dinosauria: Ceratopsidae) from the Allison Member of the Menefee Formation (Upper Cretaceous, early Campanian), northwestern New Mexico, USA Sebastian G. DalmanSpencer G. LucasPeter Dodson Research Paper 10 May 2021 Pages: 291 - 335
A partial skeleton of a new species of Tynskya Mayr, 2000 (Aves, Messelasturidae) from the London Clay highlights the osteological distinctness of a poorly known early Eocene “owl/parrot mosaic” Gerald Mayr Research Paper Open access 01 March 2021 Pages: 337 - 357
Paleobiology of Hoplophorus euphractus Lund, 1839, a large cingulate from Brazil Intertropical Region Yumi AsakuraEdison V. Oliveira Research Paper 06 March 2021 Pages: 359 - 372
