The Middle Miocene mole Desmanodon crocheti sp. nov. (Talpidae, Mammalia): the last representative of the genus in the North Alpine foreland basin Jérôme Prieto Research Paper 29 August 2009 Pages: 217 - 225
The oldest stylophoran echinoderm: a new Ceratocystis from the Middle Cambrian of Germany Imran Alexander RahmanSamuel ZamoraGerd Geyer Research Paper 22 September 2009 Pages: 227 - 237
First porcupine fossils (Mammalia, Rodentia) from the late Miocene of NW Iran, with notes on late Miocene–Pliocene dispersal of porcupines Sevket SenZeynalabedin Purabrishemi Research Paper 17 September 2009 Pages: 239 - 248
Melanosclerites from the Wilhelmi Formation (Lower Silurian, north-eastern Illinois, USA) Claudia TrampischAnthony Butcher Research Paper 15 October 2009 Pages: 249 - 257
Eptingiacea and Saturnaliacea (Radiolaria) from the middle Carnian of Turkey and some late Ladinian to early Norian samples from Oman and Alaska Paulian DumitricaUgur Kagan TekinYavuz Bedi Research Paper 31 October 2009 Pages: 259 - 292
The position of the claws in Noasauridae (Dinosauria: Abelisauroidea) and its implications for abelisauroid manus evolution Federico L. AgnolinPablo Chiarelli Research Paper 19 November 2009 Pages: 293 - 300
Biostratigraphic distribution of ammonites and calpionellids in the Tithonian of the internal Prerif (Msila area, Morocco) Mohammed BenzaggaghFabrizio CeccaIsabelle Rouget Research Paper 24 November 2009 Pages: 301 - 315