Skip to main content
Log in

Isolation and the origin of the khoisan: Late pleistocene and early holocene human evolution at the southern end of Africa

  • Published:
Human Evolution

Abstract

The debate between the proponents of the Recent African Origin and the Multiregional Evolution models for the origin of anatomically modern humans has not been of great importance to the interpretation of late Pleistocene human fossils in South Africa. The main reason is that both models propose that evidence of anatomically moderns should happen in South Africa at an early date. A more important issue to the African context is whether or not those early transitions to modernity were accompanied by the development of the distinctive local populations known today as the Khoisan. Serogenetic evidence suggests a relative antiquity for the origins of living Khoisan peoples, but this may simply reflect the longer time that anatomically modern peoples have lived in the sub-continent. A search for features of distinctive Khoisan morphology in the early remains may be of more value as it might indicate the presence of a local adaptive pattern to the special environment conditions of the region.

This is a preview of subscription content, log in via an institution to check access.

Access this article

Price excludes VAT (USA)
Tax calculation will be finalised during checkout.

Instant access to the full article PDF.

Similar content being viewed by others

References

  • Barnicot, N.A., Mukherjee, D.P., Woodburn, J.C. & Bennett, F.J. (1972) Dermatoglyphics of the Hadza of Tanzania. Human Biology 44(4):621–648.

    Google Scholar 

  • Barnicot, N.A. & Woodburn, J.C. (1975) Colour-blindness and Sensitivity to PTC in Hadza. Annals of Human Biology 2(1):61–68.

    Article  Google Scholar 

  • Bleek, D.F. (1931) Traces of former Bushman occupation in Tanganyika Territory. South African Journal of Science 28:423–429.

    Google Scholar 

  • Brauer, G. (1976) Morphological and multivariate analysis of human skeletons from Iron Age graves northeast of Lake Eyasi (Tanzania). Homo 27(3/4):185–196.

    Google Scholar 

  • Brauer, G. (1978) The morphological differentiation of anatomically modern man in Africa, with special regard to recent finds from East Africa. Zeitschrift fur Morphologie & Anthropologie, 69(3):266–292.

    Google Scholar 

  • Brauer, G. (1980) Human skeletal remains from Mumba rock shelter, northern Tanzania. American Journal of Physical Anthropology. 52:71–84.

    Article  Google Scholar 

  • Brooks, A.S. & Robertshaw, P. (1990) The Glacial Maximum in Tropical Africa: 22000-12000 B.P. In: The World at 18000 BP, Vol.2: Low Latitudes, edited by C. Gamble & O. Soffer, pp. 121–169. London: Unwin Hyman.

    Google Scholar 

  • Cavalli-Sforza, L.L., Menozzi, P. & Piazza, A. (1994) The History and Geography of Human Genes. Princeton: Princeton University Press.

    Google Scholar 

  • Dart, R.A. (1939) Population fluctuation over 7000 years in Egypt. Transactions of the Royal Society of South Africa 27:95–145.

    Article  Google Scholar 

  • Deacon, J. (1990) Changes in the archaeological record in South Africa at 18000 B.P. In: The World at 18000 BP, Vol.2: Low Latitudes, edited by C. Gamble & O. Soffer, pp. 170–188. London: Unwin Hyman.

    Google Scholar 

  • Deacon, J. & Lancaster, N. (1988) Late Quaternary Palaeoenvironments of Southern Africa. Oxford: Oxford University Press.

    Google Scholar 

  • De Villiers, H & Fatti LP (1982) The antiquity of the Negro. South African Journal of Science 78: 321–332.

    Google Scholar 

  • Ehret, C. (1985) Proposals on Khoisan reconstruction. Paper presented at International Symposium on African Hunter-Gatherers. Cologne: Institut fur Afrikanistik.

    Google Scholar 

  • Fagan, B.M. & van Noten, F. (1971) The hunter-gatherers of Gwisho. Tervuren: Musee Royale de l’Afrique Centrale. Belgique Annales, Sciences Humaines no 74.

  • Gabel, C. (1962) Human crania from the Later Stone Age of the Central Kafue Basin, Northern Rhodesia. South African Journal of Science 58:307–314.

    Google Scholar 

  • Gabel, C. (1965a) Further human remains from the Central African Later Stone Age. Man 63:38–43.

    Article  Google Scholar 

  • Gabel, C. (1965b) Stone Age Hunters of the Kafue. Boston: University Press.

    Google Scholar 

  • Galloway, A. (1933) The Nebarara Skull. South African Journal of Science 30:585–596.

    Google Scholar 

  • Godber, M., Kopec, A.C., Mourant, A.E., Teesdale, P., Tills, D., Weiner, J.S., El-Niel, H., Wood, C.H., & Barley, S. (1976) The blood groups, serum groups, red cell isoenzymes and haemoglobins of the Sandawe and Nyaturu of Tanzania. Annals of Human Biology 3(5):463–473.

    Article  Google Scholar 

  • Greenberg, J.H. (1963) The Languages of Africa. The Hague: Mouton.

    Google Scholar 

  • Grun, R & Stringer, CB (1991) Electron spin resonance dating and the evolution of modern humans. Archaeometry 33:153–199.

    Google Scholar 

  • Hiernaux, J. (1975) The People of Africa. London: Weidenfeld & Nicolson.

    Google Scholar 

  • Hiernaux, J. & Hartono, D.B. (1980) Physical measurements of the adult Hadza of Tanzania. Annals of Human Biology 7(4):339–346.

    Article  Google Scholar 

  • Irish, J.D. & Turner, C.G. (1990) West African dental affinity of Late Pleistocene Nubians: Peopling of the Eurafrican-South Asian triangle II. Homo 41(1):42–53.

    Google Scholar 

  • Keith, A. (1933) A descriptive account of the human skulls from Matjes River Cave, Cape Province. Transactions of the Royal Society of South Africa, 21(2):151–185.

    Article  Google Scholar 

  • Kingdon, J. (1993) Self-Made Man. London: Simon & Schuster.

    Google Scholar 

  • Lahr, M.M. (1996) The Evolution of Modern Human Diversity: a study of cranial variation. Cambridge: Cambridge University Press.

    Google Scholar 

  • Lahr, M.M. & Foley, RA. (1998) Towards a theory of modern human origins: geography, demography, and diversity in recent human evolution. Yearbook of Physical Anthropology. 41:137–176.

    Article  Google Scholar 

  • Leakey, L.S.B. (1970) The Stone Age Races of Kenya, 2nd edition. Oxford: Oxford University Press.

    Google Scholar 

  • McDermott, F., Stringer, C., Grun, R., Williams, C.T., Din, V.K. & Hawkesworth, B. (1996) New Late-Pleistocene uranium-thorium and ESR dates for the Singa hominid (Sudan). Journal of Human Evolution 31:507–516.

    Article  Google Scholar 

  • Mitchell, P. (1990) A palaeoecological model for archaeological site distribution in southern Africa during the Upper Pleniglacial and Late Glacial. In: The World at 18000 BP, Vol. 2: Low Latitudes, edited by C. Gamble & O. Soffer, pp. 189–205. London: Unwin Hyman.

    Google Scholar 

  • Morris, A.G. (1987) The Reflection of the Collector: San and Khoi skeletons in museum collections. South African archaeological Bulletin 42(145):12–22.

    Google Scholar 

  • Morris, A.G. (1994) The Myth of the East African Bushmen. Khoisan Studies: Multidisciplinary Perspectives. Conference Paper, Munich, 11 July 1994.

  • Newman, J.L. (1995) The Peopling of Africa. New Haven. Yale University Press.

    Google Scholar 

  • Nurse, GT (1983) Population movement around the northern Kalahari. African Studies 42(2):153–163.

    Google Scholar 

  • Jurse, G.T., Weiner, J.S. & Jenkins, T. (1985) The Peoples of Southern Africa and their affinities. Oxford: Oxford University Press.

    Google Scholar 

  • Parkington, J. (1990) A view from the south: southern Africa before, during, and after the Last GlacialMaximum. In: The World at 18000 BP, Vol.2: Low Latitudes, edited by C. Gamble & O. Soffer, pp. 214–228. London: Unwin Hyman.

    Google Scholar 

  • Rightmire, G.P. (1984) Human skeletal remains from eastern Africa. In: From Hunters to Farmers: the causes and consequences of food production in Africa. edited by J.D. Clark & S.A. Brandt, pp. 191–199. Berkeley: University of California Press.

    Google Scholar 

  • Sandelowsky, B.H. (1972) Later Stone Age Lithic Assemblages from Malawi and their technologies. Ph.D. Thesis, University of California at Berkeley.

  • Schepartz, L.A. (1988) Who were the later Pleistocene eastern Africans? African Archaeological Review. 6:57–72.

    Article  Google Scholar 

  • Stokes, S., Thomas, D.S.G. & Washington, R. (1997) Multiple episodes of aridity in southern Africa since the last interglacial period. Nature 388(10 July):154–158.

    Article  Google Scholar 

  • Stringer, C.B. (1979) A re-evaluation of the fossil human calvaria from Singa, Sudan. Bulletin of the British Museum of Natural History (Geology) 32(1):77–83.

    Google Scholar 

  • Stringer, C.B., Cornish, L. & Stuart-Macadam, P. (1985) Preparation and further study of the Singa skull from Sudan. Bulletin of the British Museum of Natural History (Geology) 38(5):347–358.

    Google Scholar 

  • Tills, D., Kopec, A.C., Warlow, A., Barnicot, N.A., Mourant, A.E., Marin, A., Bennett, F.J., & Woodburn, J.C. (1982) Blood Group, Protein, and Red Cell Enzyme Polymorphisms of the Hadza of Tanzania. Human Genetics 61:52–59.

    Article  Google Scholar 

  • Tobias, P.V. (1964) Bushman hunter-gatherers: a study in human ecology. In: Ecological Studies in Southern Africa. D.H.S. Davis (editor) pp. 67–86. Den Haag: W.Junk.

    Google Scholar 

  • Tobias, P.V. (1972) Recent human biological studies in southern Africa, with special reference to Negroes and Khoisans. Transactions of the Royal Society of South Africa 40(3):109–133.

    Article  Google Scholar 

  • Tobias, P.V. (1978) The Bushmen Cape Town: Human & Rousseau.

    Google Scholar 

  • Trevor, J.C. (1947) The physical characters of the Sandawe. Journal of the Royal Anthropological Institute 77(1):61–78.

    Google Scholar 

  • van Andel, T.H. (1989) Late Pleistocene Sea Levels and the Human exploitation of the shore and shelf of southern South Africa. Journal of Field Archaeology 16(2):133–155.

    Article  Google Scholar 

  • Vigilant, L., Stoneking, M., Harpending, H., Hawkes, K. & Wilson, A.C. (1991) African Populations and the Evolution of Human Mitochondrial DNA. Science 253 (27 Sept): 1503–1507.

    Google Scholar 

  • Wadley, L. (1993) The Pleistocene Later Stone Age South of the Limpopo River. Journal of World Prehistory 7(3):243–296.

    Article  Google Scholar 

  • Walker, N. (1994). Later Stone Age funerary practice in the Matopos, Zimbabwe: A contribution to understanding prehistoric death rites in southern Africa. Southern African Field Archaeology 3:94–102.

    Google Scholar 

  • Wells, L.H. (1951) The fossil human skull from Singa. Fossil Mammals of Africa 2:29–42. London: British Museum (NH).

    Google Scholar 

  • Willcox, A.R. (1987) San artistic ability: how far innate? South African Journal of Science 83:200–203

    Google Scholar 

  • Winkler E-M, (1984) The systematic rank of the populations of West-Kenya. Homo 35(2):73–109.

    Google Scholar 

  • Woodward, A.S. (1938) A fossil skull of an ancestral Bushman from the Anglo-Egyptian Sudan. Antiquity 14:190–195.

    Google Scholar 

Download references

Author information

Authors and Affiliations

Authors

Rights and permissions

Reprints and permissions

About this article

Cite this article

Morris, A.G. Isolation and the origin of the khoisan: Late pleistocene and early holocene human evolution at the southern end of Africa. Hum. Evol. 17, 231–240 (2002). https://doi.org/10.1007/BF02436374

Download citation

  • Received:

  • Accepted:

  • Issue Date:

  • DOI: https://doi.org/10.1007/BF02436374

Key words

Navigation