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Review of the Family Tococladidae Carpenter (Insecta: Eoblattida) from the Permian of North America and Europe

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Abstract

The family Tococladidae is transferred to the order Eoblattida. The family consists of three Permian genera: Tococladus Carpenter, 1966 from USA, France and Russia, Tshekardocladus Rasnitsyn et Aristov, 2015 from Russia and Elmocladus gen. nov. (type species Opistocladus strictus Carpenter, 1976) from the Lower Permian of USA. Opistocladus kargalensis Rasnitsyn et Aristov, 2013 from the middle Permian of Russia is redescribed and transferred to the genus Tococladus. A key to the genera of Tococladidae is provided.

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REFERENCES

  1. Aristov, D.S., New Grylloblattida (Insecta) from the Upper Permian and Lower Triassic of European Russia and Kazakhstan, Paleontol. J., 2013, vol. 47, no. 1, pp. 66–76.

    Article  Google Scholar 

  2. Aristov, D.S., Classification of the order Cnemidolestida (Insecta; Perlidea) with description of new taxa, Far East. Entomol., 2014, no. 277, pp. 1–46.

  3. Aristov, D.S., Classification of order Eoblattida (Insecta; Blattidea) with description of new taxa, Far East. Entomol., 2015, no. 301, pp. 1–56.

  4. Aristov, D.S., Revision of the family Necrophasmatidae (Insecta: Cnemidolestida), Far East. Entomol., 2018, no. 359, pp. 7–11.

  5. Aristov, D.S., A revision of the genera Heteroptilon, Nugonioneura, and Opistocladus from Lower Permian of USA (Insecta: Cnemidolestida: Tillyardembiidae, Nugonioneuridae and Parmapteridae), Far East. Entomol., 2020a, no. 401, pp. 1–9.

  6. Aristov, D.S., New Gryllones insects (Insecta: Gryllones) from the Babii Kamen’ locality (Upper Permian of Russia). 2. Order Reculida and Gryllones ordinis incertis, Paleontol. J., 2020b, vol. 54, no. 2, pp. 132–142.

    Article  Google Scholar 

  7. Aristov, D.S. and Rasnitsyn, A.P., New insects from the Kungurian of Tshekarda fossil site in Permian territory of Russia, Russ. Entomol. J., 2015, vol. 24, pp. 17–35.

    Article  Google Scholar 

  8. Bethoux, O., Ordinal assignment of the genus Tococladus Carpenter 1996 (Insecta: Archaeorthoptera), Alavesia, 2007, no. 1, p. 3.

  9. Béthoux, O. and Nel, A., Venation pattern and revision of Orthoptera sensu nov. and sister groups. Phylogeny of Palaeozoic and Mesozoic Orthoptera sensu nov, Zootaxa, 2002, vol. 96, pp. 1–88.

    Article  Google Scholar 

  10. Béthoux, O., Nel, A., Galtier, J., et al., A new species of Tococladidae Carpenter, 1966 from the Permian of France (Insecta: Archaeorthoptera), Geobios, 2003, vol. 36, pp. 275–283.

    Article  Google Scholar 

  11. Carpenter, F.M., The Lower Permian insects of Kansas: Part 11. The orders Protorthoptera and Orthoptera, Psyche, 1966, vol. 73, no. 1, pp. 46–88.

    Article  Google Scholar 

  12. Carpenter, F.M., The Lower Permian insects of Kansas: Part 12. Protorthoptera (continued), Neuroptera, additional Palaeodictyoptera & families of uncertain position, Psyche, 1976, vol. 83, pp. 336–376.

    Article  Google Scholar 

  13. Carpenter, F.M., Treatise on Invertebrate Paleontology, pt. R: Arthropoda 4, vol. 3: Superclass Hexapoda, Lawrence: Boulder, 1992.

  14. Coty, D., Hava, J., Procop, J., et al., New archaeorthopteran insects from the Late Carboniferous of the Nord and Pas-de-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera), Zootaxa, 2014, vol. 3878, no. 5, pp. 462–470.

    Article  Google Scholar 

  15. Gorokhov, A.V., Primitive Titanoptera and the early evolution of Polyneoptera, Chteniya pamyati N.A. Kholodkovskogo (N.A. Kholodkovskii Memorial Lectures), St. Petersburg, 2004, vol. 54, no. 1, pp. 1–54.

  16. Gu, J.-J., Béthoux, O., and Ren, D., Longzhua loculata n. gen. n. sp., one of the most completely documented Pennsylvanian Archaeorthoptera (Insecta; Ningxia, China), J. Paleontol., 2011, vol. 85, no. 2, pp. 303–314.

    Article  Google Scholar 

  17. Gu, J.-J., Béthoux, O., and Ren, D., A new cnemidolestodean stem-orthopteran insect from the late Carboniferous of China, Acta Palaeontol. Pol., 2014, vol. 59, no. 3, pp. 689–696.

    Google Scholar 

  18. Novokshonov, V.G., New Fossil Insects (Insecta: Hypoperlida, Panorpida, Ordinis Incertis) from the Chekarda Locality, Paleontol. J., 1999, vol. no., 1. S. 52–56.

  19. Prokop, J., Szwedo, J., Lapeyrie, J., et al., New Middle Permian insects from Salagou Formation of the Lodève Basin in southern France (Insecta: Pterygota), Ann. Soc. Entomol. France, 2015, vol. 51, pp. 14–51.

    Article  Google Scholar 

  20. Rasnitsyn, A.P., Order Hypoperlida, in Istoricheskoe razvitie klassa nasekomykh (Historical Development of Class Insecta), Trudy Paleontol. Inst. Akad. Nauk SSSR, 1980, vol. 178, pp. 41–43.

  21. Rasnitsyn, A.P., Superorder Hypoperlidea Martynov, 1928, in History of Insects, Rasnitsyn, A.P. and Quicke, D.L.Q., Eds., Dordrecht: Kluwer, 2002, pp. 111–115.

    Book  Google Scholar 

  22. Rasnitsyn, A.P., On the discussion of the wing venation of (Archae)Orthoptera (Insecta), Paleontol. J., 2007, vol. 41, no. 3, pp. 341–344.

    Article  Google Scholar 

  23. Rasnitsyn, A.P. and Aristov, D.S., New fossil insects (Insecta: Caloneurida, Hypoperlida, Palaeomanteida, Jurinida) from the Middle and Upper Permian of European Russia, Paleontol. J., 2013, vol. 47, no. 7, pp. 678–704.

    Google Scholar 

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ACKNOWLEDGMENTS

We are grateful to A.G. Ponomarenko (Paleontological Institute, Russian Academy of Sciences) and S.Yu. Storozhenko (Federal Research Center of Biodiversity, Far East Branch, Russian Academy of Sciences) for their comments on the manuscript.

Funding

This work was supported by Program 15 of the Presidium of the Russian Academy of Sciences “Problems of the Origin of Life and the Formation of the Biosphere.”

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Authors and Affiliations

  1. Borissiak Paleontological Institute, Russian Academy of Sciences, 117647, Moscow, Russia

    A. P. Rasnitsyn & D. S. Aristov

  2. Natural History Museum, SW7 5BD, London, UK

    A. P. Rasnitsyn

  3. Cherepovets State University, 162600, Cherepovets, Russia

    D. S. Aristov

Authors
  1. A. P. Rasnitsyn
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  2. D. S. Aristov
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Correspondence to A. P. Rasnitsyn or D. S. Aristov.

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Translated by M. Batrukova

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Rasnitsyn, A.P., Aristov, D.S. Review of the Family Tococladidae Carpenter (Insecta: Eoblattida) from the Permian of North America and Europe. Paleontol. J. 55, 151–156 (2021). https://doi.org/10.1134/S003103012102012X

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  • DOI: https://doi.org/10.1134/S003103012102012X

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