Cultural Dynamics in the Levantine Upper Paleolithic, ca. 40–33 ky BP: Insights Based on Recent Advances in the Study of the Levantine Aurignacian, the Arkov-Divshon, and the Atlitian

The chrono-cultural sequence of the Levantine Upper Paleolithic went through several major revisions during approximately a century of focused research, each revision contributing to shedding light on the mosaic of cultural entities and the complex social and cultural dynamics composing the Levantine Upper Paleolithic. The current state of research suggests the co-inhabitance of two cultural groups: the Early Ahmarian and the Levantine Aurignacian. Two other cultural entities, the Arkov-Divshon and the Atlitian, are regarded as younger manifestations and were tentatively suggested to relate to the Levantine Aurignacian. This paper presents a research synthesis of two case studies: Manot Cave, located in western Galilee, Israel, and Nahal Rahaf 2 Rockshelter in the Judean Desert. The application of high-resolution excavation methods, alongside detailed documentation of the stratigraphy and site-formation processes and wide-scale radiocarbon-based absolute dating, marked these sites as ideal for chrono-cultural study through the analyses of flint industries. The results indicate a clear distinction between the Levantine Aurignacian and the Arkov-Divshon/Atlitian industries and a chronological overlap between the Arkov-Divshon, Levantine Aurignacian, and possibly with the Early Ahmarian. Subsequently, we suggest another revision of the currently accepted chrono-cultural model: not two, but at least three cultural entities co-inhabited the Levant at ca. 40–30 ky cal BP. This study further suggests an evolvement of the Atlitian flint industries from the Arkov-Divshon and stresses the foreign cultural features of the Levantine Aurignacian. These results were used to construct an updated model of migration and possible interaction patterns.

The six-phase model formed the basis for reconstructing a continuous, linear cultural evolution, encompassing all of the lithic industries from the Middle Paleolithic to the Epipaleolithic of the southern Levant.The presence of Aurignacian elements in Phases III-IV was perceived as a European influence on the local industries (Garrod, 1957).The paucity of osseous artifacts, and particularly of antler-made, split-based points that were hallmarks of European Aurignacian assemblages in the early twentieth century, alongside the presence of el-Wad points, led to the distinction of the Levantine Phases III-IV from the Aurignacian of Europe, ascribing to them a local nomenclature: Lower and Upper Antelian (for Phases III and IV, respectively; Copeland & Hours, 1971;Garrod, 1957).Only later was the possibility that these assemblages reflect actual European populations established, providing a first indication of a "back migration" from Eurasia to the Levant and coining the term "Levantine Aurignacian" (e.g., Belfer-Cohen & Bar-Yosef, 1981;Besançon et al., 1975;Copeland, 1975;Hours, 1974; and see a similar suggestion that was later withdrawn in Garrod, 1953).
Further modifications to the Levantine Upper Paleolithic chrono-cultural model were introduced in the second half of the twentieth century.Accumulation of data from new survey and excavation projects (e.g., Bar-Yosef & Phillips, 1977;Gilead, 1993;Gilead & Bar-Yosef, 1993;Marks, 1976aMarks, , 1977aMarks, , 1983)), alongside comprehensive studies of the lithic industries from key sites in the Mediterranean and arid regions of the southern Levant (e.g., Belfer-Cohen, 1980;Gilead, 1981;Goring-Morris, 1987), showed marked differences between Phases III and IV.These led to a reconsideration of the cultural affiliation of Phase III and to the realization that this phase consists of two distinct components: one containing Levantine Aurignacian hallmarks and another encompassing a very blade-oriented flint industry that was named "Ahmarian" (e.g., Gilead, 1981Gilead, , 1991;;Marks, 1981Marks, , 2003)).In contrast to the Levantine Aurignacian, the Ahmarian was, and at large still is, considered to represent indigenous populations, encompassing local lithic traditions that evolved from earlier Paleolithic industries (e.g., Belfer-Cohen & Goring-Morris, 2018;Gilead, 1981Gilead, , 1991;;Kuhn, 2004;Marks, 1981Marks, , 2003;;Williams, 2003).Most significantly, however, the archaeological data accumulated by the end of the twentieth century indicated the co-existence of the Ahmarian and the Levantine Aurignacian in the Levant ca.40-30 ky BP, leading to a revision of the linear cultural evolution approach.Instead, a new model was established: The "Two Traditions" or "Parallel Phyla" model (e.g., Bar-Yosef & Belfer-Cohen, 1996;Belfer-Cohen & Bar-Yosef, 1981;Gilead, 1981Gilead, , 1991;;Goring-Morris, 1980;Marks, 1981Marks, , 2003)).The assumption that some degree of contact between two populations sharing a geographic region was inevitable led to further investigation into the manifestations of such contacts within the archaeological record, trying to extrapolate on aspects of social behaviors (e.g., Gilead, 1991;Goring-Morris & Belfer-Cohen, 2003, 2018;Lengyel, 2007;Marks, 2003;Williams, 2003;Williams & Bergman, 2010).
Marked differences between lithic industries associated with the Atlitian and Arkov-Divshon on the one hand and the Ahmarian on the other led to the distinction of the former two from the latter, primarily based on the absence of systematic blade reduction and the lack of blade predominance.Instead, the possible association of the Arkov-Divshon and the Atlitian with the Levantine Aurignacian was tentatively suggested (e.g., Gilead, 1981Gilead, , 1991;;Marks, 1981Marks, , 2003)), somewhat deferring to Garrod's initial suggestion of specialized evolvement (Garrod, 1953(Garrod, , 1957)).However, a secure connection was never fully established due to wide variability in some of the key attributes associated with each cultural entity.Table 1 presents a summary of characteristic features of the flint assemblages from primary occupation layers associated with the three cultural entities, demonstrating this issue.Some of the variability was addressed, ascribed to differences in excavation and processing methodology (i.e., in el-Wad and el-Khiam), to the incompleteness of the examined assemblages (i.e., in Yabrud and Ksâr 'Akil), to the effect of post-depositional erosion (i.e., in Arkov and possibly Ramat Materd I), and to possible local adaptation and differences in site function (i.e., in Nahal Ein Gev I and Fazael IX, e.g., Belfer-Cohen et al., 2004;Ghazi, 2013;  Shemer et al., 2023;Williams, 2003).As an alternative, the Arkov-Divshon and the Atlitian were grouped under the general description of "flake-based industries" of the Levantine Upper Paleolithic (e.g., Bar-Yosef & Belfer-Cohen, 1996;Belfer-Cohen & Goring-Morris, 2003, 2018;Gilead, 1981Gilead, , 1991;;Marks, 1981Marks, , 2003;;Williams, 2003).
This paper encompasses the research synthesis of a study conducted in the recent years that examined the archaeological evidence for a possible connection between the Levantine Aurignacian and the "flake-based" industries of the Levantine Upper Paleolithic.It was based on two case studies: Manot Cave in the Levantine Mediterranean region and Nahal Rahaf 2 Rockshelter in the Judea Desert (Fig. 1).The two sites contain well-preserved stratified sequences ascribed to the Levantine Aurignacian and the Atlitian at Manot Cave, and Arkov-Divshon at Nahal Rahaf 2 Rockshelter (Shemer et al., 2023(Shemer et al., , 2024)).The study focused on the characterization of lithic industries, combining radiocarbon chronology and considering high-resolution stratigraphy and site formation processes.These were used as the basis for chrono-cultural affiliations.The complete analyses of each of the case studies were published separately (Shemer et al., 2023(Shemer et al., , 2024) ) and are summarized here.These provided the database for the suggestions and ideas presented in this paper.The data collected from each of the case studies were tested here from a regional perspective.As a result, the current study suggests another revision of the Levantine model of cultural dynamics in the Upper Paleolithic period.Notably, however, issues regarding the Levantine Ahmarian industries are beyond the scope of this study and, therefore, presented here only in broad lines.

Nahal Rahaf 2 Rockshelter and the Arkov-Divshon Industries of the Arid Regions
The first case study, Nahal Rahaf 2 Rockshelter (henceforth NR2), is located in the Judean Desert, ca. 5 km west of the Dead Sea and ca.90 km east of the Mediterranean shoreline.The rockshelter is composed of a large, partly roofed open space (ca.35 m 2 ) and two additional inner chambers with unknown extents that are currently completely filled with sediments.Excavations on site revealed ca.1.5 m of deposits, including four occupation layers and three in situ combustion features that were associated with the Arkov-Divshon layers, together forming ca.80 cm of archaeological accumulations (Barzilai et al., 2020;Marom et al., 2022;Shemer et al., 2023).The hyper-arid conditions in the Judean Desert were favorable for the preservation of organic remains, and accordingly, in addition to rich flint assemblages, the site yielded numerous animal bones and charcoal fragments.These attributes made NR2 a high-quality case study, providing a unique insight into a cultural entity whose definition and chronology were not well established, having relied primarily on surface collections with poor organic preservation (e.g., Baruch & Bar-Yosef, 1986;Belfer-Cohen & Goring-Morris, 1986;Ferring, 1976;Gilead, 1993;Marks & Ferring, 1976;Marks, 1976b;Schyle & Richter, 2015;Schyle & Uerpmann, 1988;Williams, 2003).
The study of the lithic assemblages from NR2 indicated an industry that was equally focused on the production and use of thick flakes and blades on the one hand and twisted bladelets on the other (Shemer et al., 2023).Several refitted sequences implied the prevalence of a two-stage reduction sequence, where stage 1 was used to produce thick artifacts, and in stage 2, twisted bladelets were produced using lateral carination, applied on items reduced in stage 1 (Fig. 2).Among the tools, the microlithic component is dominant, averaging ca.42% of the retouched tools category.Bladelets most commonly presented partial, light retouch on one or both lateral edges (Fig. 3a-e).Bladelets with alternate retouch (i.e., Dufour bladelets) were present but not common, as were el-Wad points (Fig. 3f; Shemer et al., 2023).Macrolithic tools were most commonly shaped from flakes, with a notable preference for thick items, (i.e., thicker than 1 cm), commonly produced in stage 1.Endscrapers are the most common tool-type, specifically flat endscrapers (Fig. 3m-n), followed by burins, mostly of the dihedral type (Fig. 3i).Burins on truncation are rare (Shemer et al., 2023).
Most intriguing was the presence of several bone awls (Fig. 3p), previously not considered integral components of Arkov-Divshon assemblages, and of perforated marine shells (Fig. 3o), indicating contact with the Mediterranean region.
Based on a comprehensive characterization of many of the sites associated with the Arkov-Divshon, conducted by J. Williams (2003), the flint industry reflected in NR2 was suggested to bear the closest resemblance to the sites of Arkov (D22; Marks & Ferring, 1976), Ein Aqev (D31; Williams, 2003), Ramat Matred I (Gilead, 1993), and possibly also Har Horsha I (Belfer-Cohen & Goring-Morris, 1986) and Palm View 1: areas 3-4 (Parow-Souchon, 2020; Schyle, 2015;Schyle & Richter, 2015).The association relied on both the reconstruction of a primary, two-stage reduction sequence and on the abundance of lateral carination, while other aspects, such as the capacity of the microlithic component, presented high variability (Shemer et al., 2023).Notably, the presence of perforated marine shells was also reported from the Arkov-Divshon site of Ein-Aqev (Marks, 1976b;Williams, 2003).
Fifteen samples dated independently in two laboratories provided a chronological range of 39.5-34.0ky cal BP.A more confined range of 37.5-34.0ky cal BP was suggested based on the stratigraphy and the contextual quality of the samples (Shemer et al., 2023).Considering the current

Manot Cave, the Levantine Aurignacian, and the Atlitian of the Mediterranean Region
The second case study, Manot Cave, is located in the western Galilee, ca. 9 km east of the Mediterranean shoreline.
The entrance to the cave collapsed ca.30 kya, contributing to the creation of a stable, humid environment within the cave, which resulted in the good preservation of organic remains.
The cave is composed of a main, elongated hall (ca. 100 × 30 m) that branches into two smaller chambers.The primary area of human activity was identified near the entrance to the cave, on a plateau located in the topographically most elevated part of the cave.From there, a large talus steeply slopes to the southwest, ending in a second plateau (e.g., Barzilai et al., 2016Barzilai et al., , 2021;;Marder et al., 2021).Excavations in the cave implied intensive human and carnivore activity at the site and its vicinity roughly ca.55-33 kya (e.g., Abulafia et al., 2021;Alex et al., 2017;Barzilai et al., 2016Barzilai et al., , 2021;;Hershkovitz et al., 2015;Marder et al., 2017Marder et al., , 2018Marder et al., , 2021;;Shemer et al., 2024).The main occupations were ascribed to the Upper Paleolithic period, specifically to the Early Ahmarian, Levantine Aurignacian, and Atlitian (previously referred to as "post-Levantine Aurignacian"; Shemer et al., 2024).
In the entrance hall, ca.2.5 m of archaeological accumulation contained three distinct in situ occupation phases.Characterization of the flint industries alongside a high-resolution dating enabled the establishment of a refined definition and chronology for each of the cultural entities (Shemer et al., 2024).
The oldest, Phase 3, currently including unaffiliated lithic industries, was defined at the base of the sequence and will not be discussed here (Shemer et al., 2024).
Phase 2 was attributed to the Levantine Aurignacian (Shemer et al., 2024; Layers VI-IV and probably also Layers VII-VIII that were excluded from that study) and was characterized by a diversified lithic industry, where distinct trajectories were used for the reduction of different target products: blades, flakes, curved/twisted bladelets, and straight bladelets.The technological breakdown of the analyzed assemblages suggested that blade production was primarily conducted off-site (Shemer et al., 2024).These blades were an integral component of the Levantine Aurignacian industries, used for shaping typological hallmarks such as scrapers and blades with invasive, scalar retouch (i.e., "Aurignacian retouch"; Fig. 4i, j, m-n), but also items with only marginal or limited retouch included in the general "retouched blades" category.Flakes were produced in a separate sequence, often from multiple-platform cores.These were often used for the shaping of frontal carination (i.e., carinated endscrapers; Fig. 4h, l-n).Notably, the curved bladelets produced from the shaping of frontal carination were exclusively used to shape Dufour bladelets (Fig. 4f-g).Finally, straight bladelets were produced separately from designated bladelet cores and constitute the majority of microlithic tools (Fig. 4a-b).The occasional use of Middle Paleolithic artifacts via recycling marks an additional Levantine Aurignacian attribute found in Manot Cave, Phase 2 (Belfer-Cohen & Bar-Yosef, 2015; Marder et al., 2018;Shemer et al., 2024).
Manot Phase 1, defined as Atlitian (Shemer et al., 2024; Layers III-I-previously referred to as "post-Levantine Aurignacian," e.g., Barzilai et al., 2016;Marder et al., 2021), is characterized by reduced technological variability and increased dominance of the microlithic component.Two primary reduction methods are evident: The first is a focused production of thick flakes and blades (i.e., thicker than 1 cm) that were subsequently used to produce twisted blades and bladelets via lateral carination (i.e., carinated burins).These twisted bladelets constitute the majority of the microlithic component.A second reduction sequence was used to produce bladelets from single-platform cores.These cores are distinguished from the bladelet cores of Phase 2 by their extremely narrow reduction surface (ca. 1 cm wide; Shemer et al., 2024).Among the tools, the microlithic component (Fig. 5a-d) constitutes an average of ca.40%.Burins are the most abundant among macrolithic tools, comprising ca.13-21% in the Atlitian layers.Thirty to fifty percent of them were shaped on truncation, notch, or lateral preparation varieties (Fig. 5f, k); Shemer et al., 2024).Notably, an increase is observed in the dominance of the microlithic component among the tools and of burins on truncation among the burins between the oldest Atlitian Layer III and the youngest Layer I (Shemer et al., 2024).Based on the technological reconstruction of a prominent Page 10 of 23 two-stage reduction sequence, the primary contribution of lateral carination to bladelet production, and the abundance of burins on truncation among the burins (≥ 30%), Manot Phase 1 was suggested to correspond to the Atlitian assemblages of Ksâr 'Akil Rockshelter, Phase 6 (Williams & Bergman, 2010;Phase IV in Bergman et al., 2017; M.S. personal observations) and Yabrud II Rockshelter, Layer 1 (Ghazi, 2013;Ziffer, 1981).Techno-typological similarities are also suggested with el-Wad Cave, Layer C (Garrod & Bate, 1937), and el-Khiam Terrace, Layer E (Neuville, 1951), albeit the available data for these assemblages is severely meager.(Alex et al., 2017;Shemer et al., 2024).These are minimum ages, as all dated samples were collected from the youngest Layer I, while absolute dating was not achieved for the two underlying Atlitian layers (Layers II-III).However, when considering the absolute chronology of the Atlitian Phase 6 in Ksâr 'Akil Rockshelter (Bosch et al., 2015;Douka et al., 2013) alongside the minimum ages of the underlying Phase 2 in Manot Cave, a general age range of ca.35.0-33.0ky cal BP can be suggested for the Atlitian (Shemer et al., 2024).

Revisiting the Issue: The Levantine Upper Paleolithic "Flake-Based" industries and Their Relation to the Levantine Aurignacian
Comprehensive analyses of the flint assemblages from Manot Cave and Nahal Rahaf 2 Rockshelter (Shemer et al., 2023(Shemer et al., , 2024) ) provided the techno-typological basis for testing possible connections-developmental or adaptivebetween the Levantine Aurignacian and the "flake-based" industries, the Arkov-Divshon and the Atlitian.In search of shared traditions, approaches, or preferences that may be used to extrapolate on the possible assimilation between these cultural entities, an emphasis was placed on the technological attributes of the lithic industries, as they are slower to change, less affected by site and considered to reflect deeper, more inherent connections, compared to typological attributes (e.g., Bar-Yosef & Belfer-Cohen, 2013;Kadowaki, 2013;Pelegrin, 1990Pelegrin, , 1991Pelegrin, , 2000)).Nevertheless, typological aspects of the assemblages, as well as other features such as the presence and attributes of osseous industries and personal ornaments, were also examined, being integral elements of cultural characterization.
The first indicator that a revision of the accepted chronocultural model is required was the dating results of the Arkov-Divshon presence in NR2.Taken at face value, the suggested age range of ca.37.5-34.0ky cal BP places the Arkov-Divshon within the EUP rather than the LUP, implying its co-inhabitance of the southern Levant alongside the Levantine Aurignacian and possibly with the Early Ahmarian.This chronological overlap hinders a scenario encompassing its local evolvement from the Levantine Aurignacian.
Intriguingly, comparative analyses of the flint industries indicate a close resemblance between the Arkov-Divshon and the Atlitian, whereas the Levantine Aurignacian seems to encompass distinctly different technological and typological attributes.The observed resemblance between the former two is reflected primarily in the technological aspects: the Arkov-Divshon and the Atlitian share a similar reduction approach in their reliance on a primary two-stage sequence and in the systematic use of lateral carination for bladelet production (e.g., Shemer et al., 2023Shemer et al., , 2024;;Williams, 2003).Accordingly, the flint industry in both cultural entities equally emphasized the production of thick flakes and blades on the one hand and twisted bladelets on the other (e.g., Shemer et al., 2023Shemer et al., , 2024;;Williams, 2003).
The similarities between the Arkov-Divshon and the Atlitian flint industries are inherent.Hallmark attributes, such as the abundance of lateral carination commonly associated with the former and burins on truncation associated with the latter, were found in this study to be integral components in the flint assemblages of both cultural entities.In addition, the assemblages from the two case studies presented here indicate an increasing dominance of the microlithic component, possibly reflecting a general process of microlithization (Shemer et al., 2023(Shemer et al., , 2024)).The technological and typological affinities obscure the distinction between the Arkov-Divshon and the Atlitian and indicate shared lithic traditions, supporting an observation made by J. Williams two decades ago (Williams, 2003).Based on the shared reduction approach mentioned above, Williams grouped assemblages from the two cultural entities, suggesting that the geographical distribution of the sites might reflect a migration route from the Mediterranean to the arid regions of the southern Levant (Williams, 2003).Considering the chronological frames now available, it is plausible that the Atlitian represents a younger manifestation of the Arkov-Divshon, possibly indicating population movements from arid environments to the Mediterranean woodland region.
Compared to the Arkov-Divshon and the Atlitian, the Levantine Aurignacian flint industries display a wider technological diversity.Distinct reduction sequences were used for the production of different target artifacts: large blades, possibly produced off-site; flakes from single-platform and multiple-platform cores; straight bladelets from designated cores; and small, curved bladelets-products of frontal carination.Blades were highly favored for retouch in Levantine Aurignacian industries (Table 1), and the bladelet component was present but not dominant, constituting ca.20% or less of both the debitage and tools categories (Table 1, e.g., Belfer-Cohen & Bar-Yosef, 1981;Lengyel, 2007;Shemer et al., 2024;Williams, 2003;Williams & Bergman, 2010).
Frontal carination, a hallmark of Levantine Aurignacian industries, was secondary in the Atlitian and the Arkov-Divshon, where lateral carination was more common.The comparison between the industries further suggests that these two types of carination served different purposes: frontal carination of the Levantine Aurignacian (Phase 2) in Manot Cave was often accompanied by other forms of retouch, implying its use for multiple functions (Shemer et al., 2024).Reduction surface attributes and the scarcity of typical maintenance spalls indicated few rounds of reduction.Thus, there is no clear evidence for systematic bladelet production from frontal carination despite the targeted use of frontal-carination-bladelets for the shaping of Dufour bladelets.These too few to define a designated, systematic production rather than the use of reduction by-products (Shemer et al., 2024).In contrast, lateral carination in the Atlitian and the Arkov-Divshon flint industries was an integral component of the two-stage reduction sequence and a primary contributor to the production of bladelets.Intriguingly, in the Atlitian assemblages of the Jordan Valley, where lateral carination is scarce, the abundance of the microlithic component is also notably low (Belfer-Cohen et al., 2004;Goring-Morris, 1980).
The techno-typological differences between the Arkov-Divshon and the Atlitian on the one hand and the Levantine Aurignacian on the other might have been suggested to represent evolvement into a more time-efficient, microlithic-oriented approach, giving a time-lapse between these industries.However, considering the chronological overlap now suggested between the Levantine Aurignacian and the Arkov-Divshon, the observed differences seem too vast to be considered as two manifestations of a single tradition.Extrapolating from the archaeological evidence, two distinct cultural groups seem more likely to be reflected here: one corresponding with the Levantine Aurignacian and the other with the Arkov-Divshon and Atlitian.

The Origins of the Levantine Aurignacian
Numerous studies have discussed the possible origins of the Aurignacian culture, trying to trace its emergence and dispersal throughout Eurasia and often equating the patterns with the arrival and establishment of modern human populations (e.g., Anderson et al., 2018;Barshay-Szmidt et al., 2018a, 2018b;Belfer-Cohen & Goring-Morris, 2014;Bon, 2002;Bordes, 2006;Conard & Bolus, 2008;Falcucci et al., 2020;Garrod, 1953;Higham et al., 2011;Mellars, 2006;Michel, 2010Michel, , 2012;;Nigst et al., 2014;Tejero et al., 2021;Teyssandier & Zilhão, 2018;Teyssandier et al., 2010;Zilhão & d'Errico, 2006).From a local perspective, the study presented here stresses the already established view of the Levantine Aurignacian as representing traditions and ideas that were foreign to the cultural entities of the southern Levant.The techno-typological characteristics presented in this study suggest a low probability of evolutionary connections with the Arkov-Divshon and/or the Atlitian, further highlighting the detachment of the Levantine Aurignacian assemblages from the regional record.With the short chronology suggested for the Levantine Aurignacian (Phase 2) in Manot Cave, the archaeological record seems to show increasing support for a scenario in which a foreign band of hunter-gatherers arrived in the southern Levant ca.38 ky cal BP, bringing with them traditions and ideas that disappeared a few millennia later.
Searching for the roots of the Levantine Aurignacian within the Aurignacian technocomplex of Eurasia is intriguing, as Levantine assemblages contain a distinct set of attributes that is comparable to both the Early and Late phases of the Aurignacian of western Europe (Archaic/Aurignacian I and Evolved/Aurignacian II, respectively, e.g., Bon, 2002;Bordes, 2006;Davies, 2001;Dinnis et al., 2019;Douka et al., 2011;Tejero et al., 2016Tejero et al., , 2021)).
Chronologically, the ages received for the Levantine Aurignacian are young compared to the west European Early Aurignacian occupations.The current state of research suggests an age range of ca.38-34/33 ky cal BP for the Levantine Aurignacian (Alex et al., 2017;Bosch et al., 2015;Douka et al., 2013;Lengyel, 2007;Lengyel et al., 2006), correlating perfectly with the ages of the Late Aurignacian in Europe.The shorter chronology of ca.37.5-36.0ky cal BP, recently suggested based on the Manot Cave sequence, places the Levantine Aurignacian closer to the latest noted appearance of the west European Early Aurignacian, but nonetheless within the European Late Aurignacian chronology.
Intriguingly, a somewhat similar admixture of attributes has been reported in recent years from a few sites in western Europe, such as Layer 8 in Abri Pataud (Michel, 2010(Michel, , 2012)), the open-air site of Régismont-le-Haut (e.g., Anderson et al., 2018;Barshay-Szmidt et al., 2018b), and Unit 2 in Les Cottés (Jacobs et al., 2015;Roussel & Soressi, 2013), all dated roughly ca.38-36 ky cal BP.The prevalence of Early Aurignacian attributes alongside the absence of Late Aurignacian hallmarks in the form of lateral carination and burin dominance on the one hand and the relatively young chronology on the other hand led to the suggestion that these assemblages reflect a late segment within the Early Aurignacian or an intermediate phase between the Early and Late Aurignacian industries (i.e., "Middle Aurignacian"; Anderson et al., 2018;Barshay-Szmidt et al., 2018b;Jacobs et al., 2015;Michel, 2010Michel, , 2012)).
Within the framework of the present research, a similar suggestion might be made for the Levantine Aurignacian.The techno-typological attributes of Levantine assemblages correlate better to the Early rather than the Late phase of the west European Aurignacian technocomplex.Considering their chronology, the Levantine Aurignacian is likely to reflect population movement at the peak of expansion of Early Aurignacian traditions throughout Eurasia, shortly before their replacement by Late Aurignacian industries.

Cultural Dynamics and Migration Patterns: Revising the Chrono-cultural Model of the Levantine Upper Paleolithic
The study presented here provides a new perspective on the Early Upper Paleolithic period in the southern Levant.Technological and typological analyses and radiocarbon chronology established the distinction of the "flake-based industries" from the Levantine Aurignacian on the one hand, stressing the foreign nature of the latter and its association with incoming populations.On the other hand, inherent similarities were identified between the Arkov-Divshon and the Atlitian, suggesting a shared lithic tradition for the two entities, implying that both reflect a single cultural group.Adding these observations to the bank of data accumulated throughout almost a century of research, three main cultural entities can now be securely associated with the Early Upper Paleolithic period in the southern Levant: the Ahmarian, the Levantine Aurignacian, and the Arkov-Divshon/Atlitian.Each contains a distinct set of technological and typological attributes that may imply different sets of social behaviors (Table 2).
A significant contribution to the study of the Levantine Upper Paleolithic is the absolute chronology achieved for the Arkov-Divshon at NR2 Rockshelter, implying the antiquity of a cultural entity commonly regarded as a much younger manifestation (Shemer et al., 2023).Thus, further supporting the observations gained by the lithic industries: not two, but at least three cultural entities co-resided in the southern Levant ca.40-30 ky cal BP.The social complexity implied by the presence of these hunter-gatherer groups in a shared geographic region raises questions regarding inter-group dynamics and the nature of potential interactions.However, it is hard to extrapolate on these subjects based solely on the archaeological record, where the resolution is rarely sufficient for identifying and characterizing daily activities and short-term processes.Nevertheless, important data is provided by the stratigraphic record of the Levantine sites, eliciting insights about geographic distribution and relative chronology.
One of the main in the study of cultural dynamics is the question of the nature of interaction between groups.For example, what happens to indigenous groups when a new population arrives in an already populated region?Such a scenario is suggested, for example, for the Levantine Aurignacian.Regional absolute chronology indicates the presence of Early Ahmarian populations in the Mediterranean region roughly at, or close to, the time of the arrival of Levantine Aurignacian populations (e.g., Alex et al., 2017;Bar-Yosef et al., 1996;Bosch et al., 2015;Douka et al., 2013;Kuhn et al., 2009;Rebollo et al., 2011).The archaeological record suggests some degree of replacement rather than the co-existence of the two, an observation based primarily on the inner stratigraphy of the sites.Thus, when present together in a single sequence, Levantine Aurignacian layers always overlay the ones ascribed to the Early Ahmarian (e.g., Abulafia et al., 2021;Bar-Yosef & Belfer-Cohen, 2019;Barzilai et al., 2016;Garrod & Bate, 1937;Marder et al., 2017;Neuville, 1951;Perrot, 1955;Williams & Bergman, 2010).None of the sites display intermittent occupations that would correspond with the co-existence of the two cultural entities in the vicinity of a specific site.Nevertheless, the possibility that "pockets" of Early Ahmarian groups remained in the Mediterranean region throughout the Levantine Aurignacian occupation cannot be discarded.Similarly, when present in a  Garrod, 1957;Neuville, 1951) The 'Parallel Phyla' model Late 20th century (e.g., Bar-Yosef & Belfer-Cohen, 2010;Belfer-Cohen & Goring-Morris, 2003, 2014;Gilead, 1981Gilead, , 1991;;Marks, 1981Marks, , 2003) ) The single sequence, Atlitian layers always overlay the ones associated with the Levantine Aurignacian (e.g., Garrod & Bate, 1937;González Echegaray, 1964;Marder et al., 2021;Neuville, 1951;Shemer et al., 2024;Williams & Bergman, 2010), likewise implying a replacement of the Levantine Aurignacian by Arkov-Divshon/Atlitian tions ca.35 ky cal BP.
While the stratigraphic records provide some indication regarding the Mediterranean region, there is much more ambiguity regarding processes that occurred at the same time in the arid regions of the Levant.There, many of the sites are open-air and single-layered, often showing signs of deflation and post-depositional erosion (e.g., Ferring, 1976;Mark & Ferring, 1976;Marks, 1977b;Baruch & Bar-Yosef, 1986;Belfer-Cohen & Goring-Morris, 1986;Gilead, 1993;Gilead & Bar-Yosef, 1993;Schyle & Richter, 2015;Parow-Souchon, 2020).Other sites present a multilayered sequence but were ascribed in their entirety to a single cultural entity-the Arkov-Divshon, further impeding stratigraphy-based relative chronology (Marks, 1976b;Shemer et al., 2023).Therefore, the degree of interaction and possible contemporaneity between the Arkov-Divshon and the Ahmarian industries in these regions are indeterminable.Similar difficulties arise in the attempt to estimate the chronological correspondence of the Arkov-Divshon in the Negev Desert, Sinai Peninsula, and southern Jordan, on the one hand, and the dated Arkov-Divshon assemblages of the Judean Desert and the Atlitian of the Mediterranean region, on the other hand.
Nevertheless, the accumulated data allows us to suggest a partial reconstruction of migration patterns in the Levantine Upper Paleolithic period, encompassing two major shifts.The Levantine Early Upper Paleolithic seems to have seen the extensive presence of Early Ahmarian populations in the Mediterranean region (Fig. 6a).In the arid regions, the chronological data does not allow a secure determination, as most of the dated samples were obtained from open-air sites and analyzed by the end of the twentieth century, prior to the establishment of the rigorous protocols applied nowadays.Therefore, many of them present wide error margins and dated sites often present a broad scatter, falling somewhere in the range between ca.50/45-30 ky cal BP (e.g., Boaretto et al., 2021;Gilead & Bar-Yosef, 1993;Kadowaki et al., 2015;Marks, 1977b;Phillips, 1988).Nonetheless, the stratigraphy of el-Quseir in the Judean Desert, where a sequence of Early Ahmarian and Levantine Aurignacian layers was identified, presents evidence for the presence of Ahmarian populations outside the Mediterranean region prior to the arrival of the Levantine Aurignacian (Gilead, 1981;Perrot, 1955).
The arrival of the Levantine Aurignacian ca.38 ky cal BP marks the disappearance of Early Ahmarian populations from most key sites in the Mediterranean region and the northern Judean Desert (Fig. 6b, e.g., Abulafia et al., 2021;Bar-Yosef & Belfer-Cohen, 2019;Garrod & Bate, 1937;Marder et al., 2021;Neuville, 1951;Perrot, 1955;Williams & Bergman, 2010).It is possible that the Early Ahmarian groups were pushed at that point to the arid regions of the Levant; however, the limited chronological data available implies its disappearance from there as well, shortly after.It is possible that by that time, Early Ahmarian industries had already started placing an increasing focus on microliths, marking the beginning of their evolvement into Late Ahmarian/Masraqan industries (e.g., Kadowaki, 2013).The only indication of a possible long-term Early Ahmarian presence in the Levant after the arrival of the Levantine Aurignacian comes from the site of Üçağızlı, where no occupation layers associated with the latter were identified (Kuhn et al., 2009).The chronology of the site, based on radiocarbon dating of shell and charcoal samples, implies the possibility of Ahmarian presence there until ca.32.5 ky cal BP (Douka et al., 2013;Kuhn et al., 2009).However, this young chronology is viewed with caution by some scholars (e.g., Alex et al., 2017).
In the southern Judean Desert, the presence of Arkov-Divshon populations is indicated in close proximity to the estimated arrival of the Levantine Aurignacian to the Mediterranean region (Shemer et al., 2023).Chronological data on this cultural entity is still severely lacking, and it is unclear whether the Arkov-Divshon occurrences in the Negev Desert, the Sinai Peninsula, and southern Jordan can also be ascribed to this phase.Hypothetically, these sites may mark the filling of a "void" created by the disappearance or the depleting presence of the Early Ahmarian in these regions.The finds from NR2 Rockshelter imply highly desert-adapted social behaviors, including the targeting and field butchering of small-and medium-sized ungulates and the preferred use of juniper as fuel for fire (Marom et al., 2022;Shemer et al., 2023).The degree of social interactions between the Arkov-Divshon and Levantine Aurignacian is still a research subject.The presence of perforated marine shells in Arkov-Divshon sites implies contact with the Mediterranean region (Marks, Shemer et al., 2023), suggesting that perhaps the two cultural entities were aware of each other.The presence of Levantine Aurignacian in the site of el-Quseir, in the northern Judean Desert, further supports this suggestion, indicating geographic proximity and possible utilization of similar regional resources (e.g., water sources, game herds).
The high techno-typological resemblance between the Atlitian and the Arkov-Divshon has been suggested here to indicate a single cultural group.Therefore, the appearance of the Atlitian in the Mediterranean region ca.36/35 ky cal BP may be regarded as an indication for the dispersal and establishment of these cultural groups beyond the arid regions of the southern Levant.While the chronological record is insufficient in determining whether the Negev sites were still occupied at this point, in the case of NR2 Rockshelter, this timeframe marks the end of human occupation at the site.Thus, it possibly reflects a wide trend of migration towards preferable environmental conditions.
Much remains unclear regarding this phase of the Levantine Upper Paleolithic and the transition to full-fledged Epipaleolithic industries.Both techno-typological and chronological studies are required to establish a wide scope encompassing the various manifestations of the Atlitian.One important issue, for example, is represented by the Atlitian sites of the Jordan Valley: Fazael IX and Nahal Ein Gev I (Belfer-Cohen et al., 2004;Goring-Morris, 1980).Both sites are open-air, presenting lithic industries in which lateral carination and the bladelet component are severely under-represented compared to other Atlitian assemblages (e.g., Shemer et al., 2024;Williams & Bergman, 2010), while burins on truncation are highly dominant.Nevertheless, basic technological affinity is kept in the reconstruction of one primary reduction sequence, from which flakes as well as bladelets were produced (Belfer-Cohen et al., 2004).Considering the typological dominance of burins on truncation, the focus put on flake production, and the virtual absence of Aurignacian elements, the assemblages from the Jordan Valley seem to portray a close resemblance to the Atlitian/Arkov-Divshon industries.The underrepresentation of lateral carination and of the microlithic component in these assemblages emphasizes the correlation between these elements and strengthens the notion that lateral carination was a primary contributor to bladelet production in these industries (Shemer et al., 2024;Williams, 2003).Their low frequencies in the Jordan Valley sites may indicate less focus on bladelet-related activities.Therefore, the suggestion that these assemblages represent task-specific or locally adapted occurrences (e.g., Williams, 2003) seems plausible.
Another issue that was not addressed here is the question of Early Ahmarian evolvement into the Late Ahmarian/Masraqan industries, the chronology, geography, and technological characteristics of this transition, and the mechanisms leading to it.Late Ahmarian/Masraqan sites were excluded from the model suggested here due to their young chronology: ca.33/30-23 ky cal BP (e.g., Kadowaki, 2013;Goring Morris & Belfer-Cohen, 2017).However, the scenario of local evolvement from the Early Ahmarian suggests that the roots of these industries, possibly represented as a transitional phase, should appear in the last phase of the suggested model, ca.36/35-33 ky cal BP.

Concluding Remarks
The two key sites analyzed in this research, Manot Cave and Nahal Rahaf 2 Rockshelter, present new insights regarding the cultural dynamics in the Levantine Upper Paleolithic period.These insights suggest revisions to the commonly accepted chrono-cultural model and indicate more social complexity than previously thought.
The study suggests a low probability of an evolutionary connection between the Levantine Aurignacian and the "flake-based" industries of the southern Levant-the Arkov-Divshon and the Atlitian.Instead, close techno-typological similarities were observed between the two, suggesting that they represent a single cultural group.The results of this study stress the foreign nature of Levantine Aurignacian industries compared to the local Levantine record.An attempt is made to correlate between the Levantine Aurignacian and the west European Aurignacian technocomplex, suggesting an affiliation of the former with the end of the Early Aurignacian in Europe.
Techno-typological characterization combined with absolute chronology achieved for the archeological sequence from NR2 indicates the presence of the Arkov-Divshon in the region as early as ca.37 ky cal BP, placing it within the early phase of the Levantine Upper Paleolithic, alongside the Early Ahmarian and the Levantine Aurignacian.Considering the regional data accumulated in the past century, a model of population movements and possible social interaction is suggested and discussed, encompassing two major shifts.The first marked by the arrival of the Levantine Aurignacian to the region ca.38 ky cal BP, replacing, to a degree, the indigenous Early Ahmarian population in the Mediterranean region.The second shift, at ca. 36/35 ky cal BP, is marked by the disappearance of the Levantine Aurignacian and the arrival of the Atlitian to the Mediterranean region, possibly reflecting the dispersal of population from the arid regions.

Fig. 5
Fig. 5 Typical components of the Atlitian lithic industries from Manot Cave (Phase 1): a bladelet with complete retouch; b bladelet with inverse retouch; c-d Dufour bladelets; e multiple dihedral burin; f burin on a concave truncation; g dihedral burin; h, l lateral carination; i-j endscrapers on flakes; k burin on an oblique truncation

Fig. 6
Fig. 6 Suggested model of cultural dynamics in the Levantine Upper Paleolithic period, showing possible migration patterns ca.46-38 ky cal BP (a), ca.38-36/35 ky cal BP (b), and ca.35-33 ky cal BP (c).Curved arrows mark primary areas and direction of migration.Small arrows assumed interaction between cultural entities.Diagonal lines mark areas in which the chronological data (both absolute and relative) is severely lacking.(1) Üçağızlı; (2) Ksâr 'Akil;

Table 1
Summary of characteristic features of the flint assemblages associated with the Levantine Aurignacian, Arkov-Divshon, and Atlitian from primary sites in the southern Levant.*Calculated based on published data.n/a data not available.Italics font marks the case studies at the base of this research.Average values are presented for multi-layered occurrences

Table 2
Primary steps in the evolvement of the chrono-cultural model of the Levantine Upper Paleolithic, including the adjustments suggested in the current study.Arrows mark suggested techno-typologi-cal continuity.Dashed arrows marked tentatively suggested continuity, which have not been fully established