The Anisian continental-marine transition in Sardinia (Italy): state of the art, new palynological data and regional chronostratigraphic correlation

The scarce evidence of paleontological records between the upper Permian and the Anisian (Middle Triassic) of Western Europe could reflect (1) large stratigraphic gaps in the continental successions and/or (2) the persistence of disturbed conditions after the Permian–Triassic Boundary extinction event and the succession of ecological crises that occurred during the Early Triassic. In this context, the study of palynological associations, integrated with the stratigraphical and sedimentological data, plays a key role in dating and correlating the successions of the Western European domain and improves our understanding of environmental and paleoclimatic conditions. In some cases, pre-Anisian paleontological evidence is lacking, as in Sardinia (Italy), where a long gap encompasses the middle Permian (pars) to late Lower Triassic successions. Although fragmented and disseminated, the continental Lower-Middle Triassic sedimentary successions (Buntsandstein) of Sardinia have proved crucial to our understanding of the evolution of the southern edge of the Paleo-Europe and the different timings of the Tethys transgression (Muschelkalk facies) in some of these areas. Various paleogeographic reconstructions were attempted in previous works, without providing any consensus on the precise position of Sardinia and its surrounding seaways in the Western Tethys domain during this time interval. At present, the configuration and distribution of the subsiding and emerging landmasses and the temporal development of the transgressions of the Western Tethys during the Middle Triassic remain unclear. This work focuses on the stratigraphical, sedimentological and palynological aspects of three Middle Triassic continental-marine sedimentary successions in Sardinia, with particular attention to the analysis of the palynological associations sampled there, and it also provides a detailed review of all previous palynological publications on the Sardinian Anisian. The studied successions are: Su Passu Malu section (Campumari, SW Sardinia), Arcu is Fronestas section and Escalaplano section (Escalaplano, Central Sardinia). These sections were also correlated to other significant sections in the SW (Scivu Is Arenas) and NW (Nurra) parts of the island.


Introduction
The late Palaeozoic era witnessed the formation of the Pangaea supercontinent, derived from the collision of Gondwana, Laurussia and several other microplates between the Carboniferous and the early Permian (e.g. Stampfli & Borel, 2002;Stampfli et al., 2013;Pastor Galán et al., 2015). The Panthalassa ocean surrounded Pangaea while, to the east, the Tethys Sea occupied a triangular gulf that included the tropics during most of the Mesozoic. Immediately after the formation of Pangaea, during the early-middle Permian, the compressive geodynamic regime which led to the Variscan orogenic cycle progressively gave way to a large-scale transtensional-extensional regime which favoured the beginning of the break-up of the supercontinent, the subduction of the Palaeotethys oceanic ridge beneath Eurasia (Stampfli & Borel, 2002;Stampfli et al., 2013), and the opening and westward expansion of the Neotethyan Ocean (e.g., Angiolini et al., 2013). The dismantling of the Variscan orogenic system and the widespread evolution of the late Palaeozoic rifting phases produced several small basins filled with terrestrial sediments, and the deposition of these sequences occurred in seasonal to semi-arid conditions: the continentalization of the climate after Pangaea led to pronounced warming and aridification throughout the late Permian (Wignall, 2007), which persisted into the Early Triassic. This latter period was characterized by severe hothouse conditions (e.g., Bourquin et al., 2011) and high temperatures (Payne et al., 2007;Sun et al., 2012), which underwent great fluctuations, also influenced by strong and enduring volcanic activity in the Siberian Traps Large Igneous Province (Svensen et al., 2009), thereby contributing to a perturbation of the carbon cycle through the massive injection of CO 2 into the atmosphere, causing ocean acidification and anoxia. The persistence of volatile environmental conditions and the significant instability of the ecosystem throughout the Early Triassic delayed the biotic recovery after the major P-T boundary extinction event (Wignall, 2007).
Our study focuses on Sardinia, Italy, a key location for understanding the Lower-Middle Triassic sedimentary record of the Western Tethys domain. The Triassic of Sardinia is characterized by the classic Germanic threefold lithologic sub-division: Buntsandstein, Muschelkalk and Keuper, which were sometimes incorrectly used as chronostratigraphic units. Sparse clastic terrestrial to transitional Early-Middle Triassic (Buntsandstein) deposits crop out in various parts of Sardinia (Costamagna & Barca, 2002).
The first post-Palaeozoic marine transgression took place during the Middle Triassic and flooded the folded, metamorphosed and smoothed Variscan basement (Sinisi et al., 2014) in the southwestern, northwestern and central-eastern part of the island. This first transgressive pulse was represented by the transitional deposits of the upper Buntsandstein facies, or Röt facies, developed on alluvial fine-to-coarse grained deposits, eroded from the Variscan basement (i.e., Cala Viola, NW Sardinia, Fig. 1A; Monte Maiore and Escalaplano in Central-SE Sardinia, Fig. 1B; Scivu Is Arenas (Arburese) and Campumari in SW Sardinia, Fig. 1C, D). After these transitional deposits, a succession of dolomites, limestones and marls, or Muschelkalk facies, represents the first complete incursion of the Tethys Sea into Sardinia (e.g., Bourquin et al., 2011).
More particularly, central Sardinia is considered a structural highpoint during the Anisian that was progressively submerged until the maximum eustatic flooding during the Ladinian (Costamagna & Barca, 2002, 2016. The palynological characterization of these Middle Triassic deposits in this area and other parts of Sardinia has permitted some authors to propose correlations with the Alpine standard stages and substages (Pittau Demelia & Del Rio, 1980;Frechengues et al., 1983;Pittau Demelia & Flaviani, 1982a, 1982bBarca et al., 1985;Pittau & Del Rio, 2002). However, despite the good characterization and correlation with the adjacent domains, a more detailed study of these successions could enhance our understanding of processes that still remain unclear, such as the detailed configuration and distribution of the subsiding and emerging landmasses and the temporal development of the transgressions of the Western Tethys during the Middle Triassic. New stratigraphic analyses and palynological samplings were made in the Escalaplano area (Central Sardinia,Fig. 1B) and Campumari (SW Sardinia, Fig. 1D) to accurately define the age of such sediments and, thus, the beginning of Triassic deposition and the first marine incursion into this sector of the Western Peri-Tethys realm. Together with these new samplings in the cited areas, the known Anisian palynological associations of Escalaplano, Campumari, Scivu Is Arenas (Arburese) and Nurra were considered to obtain a regional correlation for the Middle Triassic continental-to-transitional deposits in the island.
This work also provides up-to-date stratigraphical and sedimentological analyses, which particularly further our knowledge of the palynostratigraphical zonation of these successions and permit a more accurate determination of the age of such sediments and the beginning of both continental and marine Triassic deposition in this sector of the Western Peri-Tethys domain.

Geological setting
In Sardinia, the post-Variscan deposits are arranged in three main sedimentary sequences: the first (lower Permian) is related to the development of the aforementioned intra-Pangaea lateral mega shear system; the second (lower Permian to early middle Permian) is related to a more mature post-orogenic rifting phase; and the third (Early-Middle Triassic) is associated with fluvial sedimentation in medium-sized basins and the first eastward marine incursions through narrow corridors (Cassinis et al., 2003). This third sedimentary sequence led to the deposition of red continental terrigenous sequences in Buntsandstein facies, represented by the Punta S'Arridelli Formation and the Riu Is Corras Formation (SW Sardinia Fig. 2: 1,2), the Escalaplano Formation in the Sarcidano area (Central Sardinia Fig. 2: 3, 4) and the Conglomerato del Porticciolo and Arenarie di Cala Viola Formation in the Nurra region (NW Sardinia Fig. 2: 5, 6). According to Costamagna and Barca (2002), these units were mainly formed in a small ephemeral distal fan environment, as the development of a significant fan-delta complex could have been hindered by the low landforms resulting from the several post-Variscan erosion cycles, which severely reduced most of the reliefs through intense chemical and mechanical weathering processes (Scotese & Schettino, 2017). The passage from the Buntsandstein to the shallow marine, carbonate platform environment in Muschelkalk facies is marked by the presence of transitional deposits in Germanic-like "Röt" facies, which present different lithologies from, for instance, those defined in Iberia. The first accurate and illustrated documentation of the Sardinian "Röt" facies can be found in Costamagna, (2011; it is characterized by marls and evaporitic lithologies which can be seen, for example, in the Punta S'Arridelli Formation and at the uppermost levels of the Riu Is Corras Fm (Fig. 2). These fms. also show a prominent carbonate influence, the genesis is still subject to debate; it is difficult to interpret due to the occurrence of pedogenetic processes, as testified by the presence of caliches (Barca et al., 1995b;Cocozza & Gandin, 1976). As regards the NE and SE successions, the transition to the marine environment is quite abrupt, with only a few meters of marls (1) Su Passu Malu (Campumari, Iglesiente) (2) Punta su Nuraxi (Scivu is Arenas, Arburese) (3) Arcu is Fronestas (Escalaplano) (4) Escalaplano SW (Escalaplano) (5) Cala Viola (Nurra) (6) Cugiareddu well (Nurra). (1) Marls (2) Sandstones (3) Limestones (4) Crossbedded conglomerates, (5) Sandy limestones (6) Gypsum (7) Breccias (8) Bedded sandstones (9) Dolomitic limestones (10) Interbedded sandstones and siltstones (11) Schist (12) Dolostones (13) Conglomerates (14) Crossbedded sandstones (15) Calcretes (16) Interbedded shales and silty limestones (17) Clays (18) Silty limestones. See Fig. 1 for locations passing into the dolostones of the Muschelkalk, as in the Escalaplano Formation, where the influence of the marine environment is testified by the presence of the aforementioned "Röt" facies, consisting of calcareous-evaporitic intercalation containing wrinkled algal mats (Costamagna & Barca, 2002). The studied sections are included in four regions of the island: Escalaplano-Sarcidano/Gerrei region (Central-SE Sardinia), Campumari-Iglesiente region (SW Sardinia), Scivu Is Arenas-Arburese region (SW Sardinia) and Nurra region (NW Sardinia).

Central-SE Sardinia (Sarcidano-Gerrei region)
Close to the village of Escalaplano, to the SW (Sarcidano-Gerrei region, Fig. 1B), the post-Variscan deposition is mainly represented by a lower Permian volcano-sedimentary sequence that fills a small (about 15 km 2 ) intramontane basin (Fig. 1B). On the whole, the extruded calc-alkaline magmatic products in this succession have an extrabasinal origin and largely exceed the strictly sedimentary units.
According to Pecorini (1974), Ronchi (1997) and Ronchi et al. (2008), the entire Permian succession can be subdivided into two main parts, namely "Lower" and "Upper" volcanic and sedimentary successions: both of these begin with a coarse sedimentary unit, continue with reworked volcaniclastites, tuffites, lacustrine-to-palustrine deposits and end with, respectively, an acidic ignimbrite deposit and an andesitic lava flow. Nine lithostratigraphic units have been distinguished within the two cited volcano-sedimentary portions by Ronchi et al. (2008), which seem to be separated by a paraconformity(?). The estimated average thickness is approximately 150 m, possibly reaching a maximum of almost 200 m in the basin's northwestern sector.

SW Sardinia: Campumari (Iglesiente region)
The Campumari-Coremò area is located along the southwestern coast of Sardinia, in the Iglesiente area, and it consists of a series of small tabular reliefs with slightly different heights, forming a plateau with a maximum altitude of 219 m (Fig. 1D).
The post-Variscan "Permo-Triassic" auctt. deposits vary from a minimum of 10 m in the "Case Miniera Paneccani" locality up to a maximum of 45 m in the Su Grifoneddu de S'Acqua area, and they lie in angular unconformity over the low metamorphic basement (Fig. 2B 1).
The Campumari plateau has been studied since the end of the nineteenth century (Lovisato, 1881;Tornquist, 1902;Novarese, 1914;Taricco, 1928) but there has never been any agreement about the chronology of its sedimentary formations, which have been attributed to very different ages (Permian, Triassic, Cretaceous and Eocene). It was not until the work of Cocozza and Gandin (1976) that the sequence was studied in detail; these authors recognized a stratigraphic gap between the true Eocene deposits and the underlying carbonate lithologies, and these latter deposits were finally attributed to the Triassic, based on facies analogies with the Triassic sediments of the Scivu-Is Arenas and Monte Maiore areas. Later, Pittau Demelia & Del Rio (1980) described a palynological association with a discontinuous gray-blackish clay level located in the upper part of the "B interval" (Cocozza & Gandin, 1976), now formalized as "Su Passu Malu Member" (Barca & Costamagna, 2003). They therefore attributed at least the upper part of the clastic unit of the Campumari succession to the Middle Triassic or "Lower Muschelkalk''. Subsequently, Barca and Costamagna (2003) performed another detailed sedimentological study of the Campumari sequence, identifying and formalizing two formations and their respective members, the Riu Is Corras Formation and Campumari Formation (Costamagna & Barca, 1997b) (Fig. 2:1). The former consists mainly of conglomerates and calichified, brecciated horizons liying unconformably over the basement; it has been interpreted as the product of a coastal plain, with a distal fan environment in a semi-arid hot climate (Cocozza & Gandin, 1976;Costamagna & Barca, 2002). The overlying Campumari Formation is sub-divided into two members: the Su Passu Malu Member begins with dark marly-clay deposits (locally with thin carbonate intercalations) and mainly consists of thinly stratified, laminated, yellowish marly dolostones bearing sulphate pseudomorphs and chert nodules, and the Grifoneddu de S'acqua Member, which is composed of dolostones and starts with a breccia horizon marking the passage to a shallow marine, restricted lagoonal environment (Costamagna & Barca, 2002).

SW Sardinia: Scivu Is Arenas (Arburese region)
The Triassic of the Scivu-Is Arenas area overlies the Variscan metamorphic basement and is in turn covered by Quaternary deposits (calcarenites and aeolian sands), forming small reliefs (Figs. 1C, 4A). The area was first studied by Bornemann (1881), who attributed the outcrops to the Triassic sedimentary record on the basis of the fossil content (e.g., presence of Myophoria goldfussi Von Alberti, Gervilleia subglobosa Credner, Rhyzocorallium jeneense Zenker, among others) and correlated the succession to the Grès Bigarré of Provence, due to similarities in facies. It was only after more recent studies (Barca et al., 1995b;Costamagna & Barca, 2002;Damiani & Gandin, 1973b) that these deposits received more detailed chrono-and litho-stratigraphic subdivisions, as reported below. The older Triassic unit, namely Punta S'Arridelli Formation (Costamagna & Barca, 2002), lies nonconformably on top of the Palaeozoic basement ( Fig. 2: 2, 4A) and consists of two sub-units, the lower Su Ripostiggiu Member and the upper Bruncu Pilloni Member, representing the evolution from a continental braided stream environment, to a more transitional, sabkha-to-shallow marine setting (Costamagna & Barca, 2002). The Su Ripostiggiu Member starts with thin reddish breccias continuing to coarse sandstones with angular-to-subrounded clasts coming from the eroded metamorphic basement and volcanic detritus, and it then evolves upwards to purple-reddish paleosols and calicified carbonate layers containing fragments of basement and reddish sandstone (Fig. 4C). From the base to the top of this member, there is a decrease in the clast size and a variation in clast composition, which changes from almost totally volcanic to exclusively metamorphic basement fragments, marking an increase in erosive sub-aerial conditions (Barca et al., 1995a(Barca et al., , 1995b. The Su Ripostiggiu Member (about 20 m) passes upwards (or laterally to the south) to the Bruncu Pilloni Member, formed by dedolomitized, locally cavernous limestone, bearing siliceous pseudomorphs, levels of dissolution-collapse breccias and traces of calicitization processes in its upper part.
The Punta Su Nuraxi Formation (Costamagna & Barca, 2002) overlies Punta S'Arridelli Formation with a slight angular unconformity (Fig. 4A) and is divided into two different sub-units: a lower Case Pisano Member and an upper Bruncu Zippiri Member (Costamagna & Barca, 2002). The Case Pisano Member starts locally with a dark, marly mudstone layer, yielding a microflora assemblage suggesting an upper Anisian age, according to Barca et al. (1995b).
There follow metric alternations of massive, cavernous, calcareous mudstones bearing siliceous pseudomorphs and evaporitic minerals, and whitish layered and cross-laminated calcarenites-calcilutites. Salt tectonic structures (folds several metres in size) occur in this sub-unit, along with collapse-dissolution breccias and localized calcrete structures. The Case Pisano Member evolves upwards in the Bruncu Zippiri Member through a thick layer of dissolution-collapse breccias. The Bruncu Zippiri Member consists of two facies, the first being represented by blue-gray limestone with reddish intercalation of bioclastic grainstones-packstones, locally presenting cross, wavy and planar lamination, fossil content (bivalves, foraminifera and algae) and rare evidence of storm beds eroding the substratum (Costamagna & Barca, 2002). The second facies is represented by massive calcarenite-calcilutite (packstones to wackestones) evolving upwards into gray-reddish (Fig. 4E), marly limestones (wackestones to mudstones), showing bioturbation structures classified as Planolites and Paleophycus tubularis (Costamagna & Barca, 2002;Moore, 1962).

NW Sardinia (Nurra region)
The post-Variscan succession of the Nurra region (Northwestern Sardinia, Fig. 1A) was deposited on the Variscan crystalline basement during the early Permian to Middle Triassic time-span; it is represented by a thick pile of continental sediments (about 700 m thick), mainly Permian, evolving into continental-to-shallow marine Triassic deposits with the classic Germanic facies (Buntsandstein, Muschelkalk and Keuper Auct.) (Fig. 4B). The first and second sequence belong to the Permian and are represented by the stacking of different fluvial to lacustrine formations (e.g., Cassinis et al., 2003;Costamagna, 2019).
The third sequence (approx. 50 m thick) consists of two formations, namely the Conglomerato del Porticciolo and Arenarie di Cala Viola ( Fig. 2: 5), separated by a minor erosional surface. The former (up to 12 m thick) was accumulated in a gravelly braided river setting under conditions of persistent aridity, as suggested by the common occurrence of wind-worn clasts (Cassinis et al., 2003;Durand, 2008). Its base consists of the alternation between massive to cross-bedded ortho-and para-conglomerates (mainly polycrystalline quartz), presenting imbrication features and cross-laminated siltstones to sandstones. The top of the formation consists of a 4-5 m alternation of trough cross-bedded, medium-fine sandstones of fluvial and aeolian origin and coarser pebbly sandstones.
The Arenarie di Cala Viola Formation is about 40 m thick and unconformably overlies the Conglomerato del Porticciolo Formation. Its base is mainly represented by dark continental red sandstones and siltstones arranged in thin beds that present small, pedogenic-origin, intraformational breccias, sets of climbing ripples evolving into horizontal to low-angle laminated or trough cross-bedding, with subordinate mudstones in the lower part. It is overlain by a few meters of gray-green, pink and reddish, medium-to coarse-grained sandstones, with the presence of lensoid geometries and trough cross-bedding (Fig. 4D). Further up, in the middle-upper part of the unit, there are thinly bedded to well-bedded, medium-to fine-grained, dark orange/red to purple sandstones and siltstones, while, finally, the top of the unit consists mainly of whitish-green to gray siltstones and claystones. The Arenarie di Cala Viola Formation marks a dramatic change in sedimentary and climatic conditions. In fact, while the Conglomerato di Porticciolo was probably deposited during the late Early Triassic (late Olenekian) sedimentary cycle, under arid to hyper-arid conditions (Durand, 2006(Durand, , 2008Bourquin et al., 2007Bourquin et al., , 2011Borruel-Abadía et al., 2019), similarly to the Poudingue de Port-issol in Provence, the Arenarie di Cala Viola Formation with its finer grained facies, was deposited in a terminal fan setting under semi-arid conditions (Cassinis et al., 2003;Citton et al., 2020;Durand, 2006Durand, , 2008, as testified by the presence of pedogenetic carbonate concretions, mud cracks, tree-related bio-sedimentary structures and the presence of inundite sequences ( Fig. 4F) (sensu Seilacher, 1982). Some authors (i.e. Fontana et al., 2001) have postulated a tidal influence in the Arenarie di Cala Viola Formation, based on the sedimentology and stratigraphic position of this unit, which narrowly predates the widespread Middle Triassic marine transgression documented by the 'Muschelkalk'-type carbonates.

Stratigraphy
For this work, detailed bibliographical research was undertaken, in order to review and update the stratigraphical and sedimentological data related to the areas of interest. The sections mentioned below were re-examined and studied, with particular attention to sedimentary facies recognition and associations. They crop out in the Iglesiente (SW Sardinia) and the Sarcidano-Gerrei (Central-SE Sardinia) regions of the island (Fig. 1).
In the former area, more specifically on the Campumari plateau, the Su Passu Malu section was analyzed. In the latter area, two sections were described: the first cropping out at the southern entrance of the Escalaplano village, and the second a few km to the north, in the Arcu is Fronestas locality (Fig. 1B). The stratigraphy and facies analyses provided by the fieldwork were integrated into the pre-existing information and complemented by new palynological data and revised older data. Based on the new interpretation of the microfloristic associations, correlations were drawn at a regional level. In a broader interregional context, the data have been used to better frame Sardinia in the palaeogeographic evolution of the Western Tethys during the Anisian.

Methodology for palynological revision
A precise review of all the previous palynological publications on the Sardinian Anisian was undertaken for the present work, and it was necessary to establish the validity of these data for our purposes.
The minimum conditions for considering the data valid were: 1) correct figuration: lists of taxa without representation were not considered valid since there was no possibility of confirming or identifying the palynomorph. 2) correct location: each palynomorph association must be referred to an exact position in the stratigraphic section in which it was sampled.
Based on these premises, identifications from previous papers that meet these requirements could be incorporated into the discussion, together with unpublished data (Diez, 2000) and new data obtained in the last field trips undertaken by the authors of the present paper.

Palynological analyses
This paper presents unpublished and revised results from Diez's PhD thesis (2000) collected in 1996 in the Escalaplano area: three samples (PC-2, PC-3 and PC-4) at the southern entrance to Escalaplano village (Escalaplano SW section, Fig. 5) and another sample (PC-7.2) taken from the Arcu is Fronestas section, north of the same village ( Fig. 6). Three new samples (AIF-2, AIF-5, and AIF-9) were later collected in 2015 and 2016 fieldwork in the Arcu Is Fronestas section and another one (GRIFO 3) was collected from the Su Passu Malu section on the Campumari site (Campumari section, Fig. 7).
Palynological samples were processed using HCl-HF-HCl classic attack techniques, as described by WOOD et al. (1996), in the Palynology Laboratory in the Department of Geosciences at the University of Vigo (AIF-2, AIF-5, AIF-9, GRIFO 3) and the Paleobotany Laboratory in the University Pierre et Marie Curie, Paris-6 (PC-2. PC-3, PC-4 and PC-7.2). A dispersing agent was added to facilitate filtering and sieving at 10 µm. The palynological slides were studied under a Leica DM2000 LED, and the photomicrographs were taken with a Leica ICC50 W camera using × 1000 magnification.   The slides are stored in the Palynology Laboratory in the Department of Geosciences at the University of Vigo.

Escalaplano
Reddish clastic sediments with chalky and marly-clayey intercalations, up to about 20 m in thickness, unconformably overlie the Variscan metamorphic basement at the northeastern margin of the Permian Escalaplano basin (Fig. 1). The marked unconformity between the crystalline basement and these Middle Triassic siliciclastics can be observed along the SS 128 Ballao-Escalaplano road, just before the first buildings at the southern entrance to the village of Escalaplano (Fig. 1B). A stratigraphic superimposition of the Triassic deposits on the Permian succession has been described so far (Ronchi, 1997) only in the Arcu 'e Azzinuri locality (i.e., along the Escalaplano-Orroli road, just a few hundred metres west of the village).
Some representative stratigraphic sections have been described in the Triassic sedimentary deposits in the Escalaplano area (Fig. 1). Two of these are described herein, with indications of the palynomorph horizons. At Escalaplano's southern entrance (SW Escalaplano section, Fig. 5), the depositional environment evolves from continental alluvial environment (Buntsandstein) to transitional salted lagoons, forecasting the marine (Muschelkalk) transgression (Costamagna et al., 2000). The following lithostratigraphic units can be recognized, from their base lying on violet schists from the basement to the Eocene siliciclastics: -reddish basal polygenic conglomerates and breccias (Fig. 5A), with angular elements of the underlying metamorphic basement, quartz pebbles and Permian volcanic rocks, embedded into a sandy matrix alternating with reddish sandstones (0-2 m). This unit is very reduced northwestwards, in the area where the unconformable contact with the basement is underlined by fine-grained deposits; -dark red to violet to gray-green, evenly bedded in cm-dm thick layers, mica-rich siltstones, sandstones, marly claystones and marls. Irregular white-to-pink satin spar veins occur frequently (0-10 m) (Fig. 5B). This lithofacies can reach up to 15 m in the area; in the upper part, there are alternating beds of dark claystones, thin dolomitic marls, siltstones and cm-irregular layers of pink and white satin spar; -yellowish light brown to gray irregularly bedded limestones and dolomitic limestones (Mushelkalk Auctt.) (3 m) (Fig. 5C). These carbonates, which can reach about 20 m in the area, are unconformably overlain by alluvial coarse Eocene deposits in this locality. To the north of the village of Escalaplano (i.e., Is Forreddu and Arcu is Fronestas sections), the same Middle Triassic siliciclastics are in turn directly overlain by Middle Jurassic quartz conglomerates (Genna Selole Formation, Costamagna, 2016;Costamagna et al., 2007;Dieni et al., 1983) (Fig. 3).
In the Arcu Is Fronestas locality (Arcu Is Fronestas section, Fig. 6), located about 3 km to the NE of Escalaplano, on the road to Perdasdefogu (Fig. 1B), the Middle Triassic succession is less than 20 m thick. It lies unconformably over the Variscan basement and is unconformably overlain by the Middle Jurassic quartz conglomerates of the Genna Selole Formation. The section can be described as follows: -the base of the succession consists of pinkish-reddish polygenic conglomerates and breccias and it unconformably overlies the schists of the Variscan basement (Fig. 6A). Quartz pebbles, angular clasts of metamorphic basement and Permian volcanites are supported by a sandy matrix alternating with reddish sandstones (80 cm); -it follows an approximately 4 m-thick alternation of dark gray to reddish claystones and greenish-yellow marly claystones, with centimetric intercalations of medium-coarse sandstones bearing halite casts, mudcracks and bioturbation (AIF2, AIF5 pollen samples, Fig. 6A1); -moving upwards, there is an increase in the carbonate content, resulting in the presence of reddish marls and clayey marls over claystones (10-15 m; Fig. 6B); -dark gray-black claystones to yellowish marly claystones (5 m) (AIF 9 pollen samples, Fig. 6C) make up the upper part of the Escalaplano Formation in this area, and they are overlain by the Jurassic quartz conglomerates of the Genna Selole Formation.

Campumari
Sparse and thin "Permian-Triassic" auctt. successions unconformably cover the early Palaeozoic slightly metamorphic rocks in the Iglesiente and Arburese regions (SW Sardinia, Cocozza & Gandin, 1976;Barca & Costamagna, 2003a). The studied section is about 50 m thick and is located in the Su Passu Malu locality, along the SP83, between the villages of Nebida and Gonnesa ( Fig. 7 A and  Fig. 1). From the base to the top, the section shows: -the Riu Is Corras Formation lies unconformably over the folded Variscan basement and shows marked lateral heterogeneity. The first 14 m of the studied section consist of yellowish calcrete, organized in thick banks, often brecciated and with evidence of calichitization processes; -this is followed by about 8 m of well-cemented, heterogeneous conglomerates organized in metric strata (Fig. 7B). Subordinated, dolomitic crusts, nodular and brecciated dolomitic mudstones, rare sandstones with caliches and reddish argillaceous siltites occur; -the passage to the Campumari Formation lower member, i.e., the Su Passu Malu Member, is marked by 1.5 m-thick, thinly bedded light gray mudstones. This level is followed by a conglomerate bank about 1 m thick, which is again followed by 2.5 m of gray mudstones and dolomitic mudstones, locally rich in plant debris, sulphate pseudomorphs and carbonate-rich silt intercalations (pollen sample GRIFO 3) (Fig. 7C); -there follow 4 m of laminated dolostones (Fig. 7D), organized in centimetric to decimetric strata and showing bioturbation and tepee structures; -the passage to the Campumari Formation upper member, i.e., the Grifoneddu de S'acqua Member, is marked by a collapse breccia horizon (1-2 m thick) (Fig. 7E) followed by massive to well-stratified dolostones, showing laminations, evidence of bioturbation (Rhyzocorallium jenense Zenker; Cocozza & Gandin, 1976), and pseudomorphs after evaporitic minerals.
The first reference to the presence of a Middle Triassic pollen association is found in Damiani and Gandin (1973c). These authors mentioned a productive sample collected in reddish marly layers just southwest of the village of Escalaplano (level 2 of the section that corresponds to km 47.4 of the s.s. Nurri-Escalaplano, now SP10) (Fig. 2). At the time of the publication of Damiani & Gandin's work, the palynological sample was being studied by Del Rio and, for this reason, the authors limited themselves to preliminarily pointing out the presence of a continental floristic association from the Middle Triassic, represented mainly by gymnosperms.
The first formal palynostratigraphic study on the Anisian of Sardinia was undertaken by Pittau Demelia & Del Rio (1980) via a sparse and poorly preserved palynological assemblage (see the complete list of taxa in Supplementary Material) in the clay levels of the upper part of member B (sensu Cocozza & Gandin, 1976) of the stratigraphic section of the Campumari outcrop, at km 1.7 between Funtanamare and Nebida (Fig. 2). These authors assigned to this sample a "Lower Muschelkalk" age due to the presence of Stellapollenites muelleri (= Hexasaccites muelleri), Minutosaccus crenulatus and the genus Triadispora. Although the figures do not confirm all the taxonomic attributions, we could suggest an Anisian sensu lato age through the correct identification of Hexasaccites muelleri (Visscher & Brugman, 1981;Brugman, 1983Brugman, , 1986Diez, 2000;Kürschner & Herngreen, 2010).
In Flaviani (1980), the palynostratigraphy from the subsurface of Nurra (Cugiareddu well, Figs. 1A, 2: 6) was presented. This is an unpublished study whose results were used at the time to make lithostratigraphic correlations between Triassic outcrops in Sardinia. In this work, new palynomorphs were described; the authorship of these new species was attributed to P. Pittau in later papers (Pittau & Del Rio, 2002;Pittau Demelia & Flaviani, 1982a, 1982b, and they were used only for Sardinian Triassic studies. Normally this type of publication should not be treated in the context of a review as it is unpublished research, but it has been necessary to include it due to its importance in the development of our review. The samples used for palynostratigraphic studies in Flaviani's thesis were provided from a previous work by Pomesano Cherchi (1968) on the Cugiareddu well (Fig. 1). These samples correspond to level 545 (sample 970 in Flaviani, 1980), ascribed to the Permian; levels 477 and 443 (samples 756 and 755 respectively, in Flaviani, 1980), which Pomesano Cherchi (1968) described as "Buntsandstein'' but Flaviani attributed to the Lower and Middle Muschelkalk respectively; finally, a level 429 (sample 754 in Flaviani, 1980) was described as Lower Muschelkalk by Pomesano Cherchi (1968), and later attributed to the Upper Muschelkalk by Flaviani (1980).
As for sample 970, Flaviani (1980) showed a poor assemblage (see the complete list of taxa in Supplementary Material) and ascribed it to an inconclusive Permian-Triassic age through comparison with the Germanic Zechstein associations. However, the joint appearance of the guide taxa Lueckisporites virkkiae, typical of the middle and upper Permian, and Hexasaccites muelleri that appears during the Anisian (Middle Triassic) and other Triassic taxa, such as Triadispora crassa, Enzonalasporites leschikii and Illinites sp., suggests an Anisian sensu lato age. The presence of a single specimen of Lueckisporites virkkiae would not be sufficient for any Permian affinity.
The next two associations, of samples 756 and 755 (see the complete list of taxa in Supplementary Material), were included in "Zone A": this Zone would correspond to the Lower and Middle Muschelkalk interval (Anisian). It would be characterized by the significant presence of Hexasaccites muelleri and Triadispora crassa, and the Alisporites-Cuneatisporites-Sulcatisporites bisaccate pollen morphogroup. Flaviani also differentiated two subzones: A1 (sample 756) and A2 (sample 755). Subzone A1 (Lower Muschelkalk, Anisian) would be characterized by the presence of Aratrisporites bulloides (n. sp. in Flaviani, 1980), Lunatisporites rhaeticus and Lunatisporites acutus. Subzone A2 (Middle Muschelkalk, Anisian) would be characterized by Myriamsporites triassicus (n. sp. in Flaviani, 1980) and Aratrisporites distalirugulatus (n. sp. in Flaviani, 1980), as well as Krauselisporites cuspidus, Aratrisporites paraspinosus, Ellipsovelatisporites plicatus, Striatiabieites aytugii and Angustisulcites klausii. As we explain in the next section, the presence of Hexasaccites muelleri would indicate an Anisian age for samples 756 and 755. Unfortunately, we believe that the sub-divisions A1 and A2 could not be justified when using new species from levels that have not been geochronologically dated or correlated with other sources previously. Flaviani (1980) assigned the fourth association (see the complete list of taxa in Supplementary Material), referred to as sample 754 (429 m.), to the Upper Muschelkalk ("Zone B", Anisian-Ladinian transition), based on the absence of Hexasaccites muelleri and the presence of Ovalopollis pseudoalatus. However, following our methodology, we cannot endorse this datum, since we do not accept the criterion of absence.
Pittau Demelia and Flaviani (1982b) included a partial publication of the study by Flaviani (1980) of the Cugiareddu well. The featured palynomorphs are not correctly referenced, however, and neither do these authors show photos of all the listed taxa, nor any references to the studied samples. It can be deduced, solely from the wording of the text, that the specimens are the same, and the new species described in Flaviani (1980) already appear as Pittau's in Flaviani, 1980. Therefore, the same considerations applied to Flaviani (1980) can be used for Pittau Demelia and Flaviani (1982b). Some photos of taxa are included, without mentioning what level they correspond to. In this case, we can confirm the existence of Hexasaccites muelleri and the possible classification of Triadispora crassa and Prodiploxypinus gracilis, which reaffirms the notion that nothing can be said about those levels containing H. muelleri that does not correspond to the Anisian sensu lato.
Subsequently, new palynological data (see the complete list of taxa in Supplementary Material) were described from the Punta del Lavatoio section (Fig. 1A) by Pittau Demelia and Flaviani (1982a). The authors sampled different dark clay layers located in the "Lithofacies C" (Costamagna, 2002). Due to the presence of the morphological group Alisporites-Sulcatisporites-Cuneatisporites, Triadispora crassa, Aratrisporites sp., Aratrisporites paraspinosus, Camarozonosporites cf. rudis, Polypodiaceoisporites sp. and Verrucosisporites sp, in sample 1275, coming from one of these levels, it was possible to correlate it with subzone A2 of the Cugiareddu well (Flaviani, 1980;Pittau Demelia & Flaviani, 1982a, 1982b. With this correlation, the authors pointed out that, as in the A2 subzone defined in the Cugiareddu well (not in Punta del Lavatoio), Stellapollenites muelleri (= Hexasaccites muelleri) occurs, and so this level can be ascribed to the late Anisian, and the upper part to the early Ladinian due to the presence of Diplopora. This reasoning is not viable under our methodology because photos of only seven taxa are presented (Aratrisporites sp., Chordasporites singulicorda, Concavisporites sp., Polypodiaceoisporites sp., Striatoabieites sp., Triadispora plicata and T. suspecta). Furthermore, the determination is based on a taxon that would not even have been present in the sample.
Subsequently, Frechengues et al. (1993) described palynological results for a sample (see the complete list of taxa in Supplementary Material) in a dark clay level, included in the "Escalaplano section", which they also compare to those mentioned above by Damiani and Gandin (1973a) (Fig. 2: 4). The authors suggested a Ladinian age to this sample due to the presence of a large amount of Triadispora spp., numerous Alisporites, Pityosporites neomundanus and Cycadopites, the absence of Ovalipollis and Circumpolles and the low quantity of spores, except for Aratrisporites saturni and A. fisheri. In the same work, the authors compared this assemblage with similar ones in the Pyrenean ranges and with the association described in Pittau Demelia and Flaviani (1982a) for the section of Punta del Lavatoio. Of all the taxa referenced, only the photo of Heliosaccus dimorphus is published. This does not allow us to conclude that the classification is correct and, because of the lack of characterization of the forms presented in that work, we prefer not to attribute these levels to the Ladinian.
In Barca et al. (1995b), a well-preserved association (see the complete list of taxa in Supplementary Material) is yielded by the basal marly levels of the carbonate Case Pisano Mb (Punta Su Nuraxi Formation, Costamagna & Barca, 2002) in the Scivu-Is Arenas section (Figs. 1C, 2: 2). The authors reported a very diverse palynomorph content with fifty-three different taxa, but photos of only twenty-six of these are shown. Due to the presence of Triadispora crassa, together with Illinites chitonoides, Angustisulcites klausi, Microcachrydites fastidioides, Stellapollenites thiergartii (= Hexasaccites muelleri), Perotriletes minor and Cristianisporites triangulatus, they consider that the association would be Pelsonian-Illyrian in age and would be correlated with the "phase Mu-1" of Dockter et al. (1980). Barca et al. (1995b) Flaviani, 1980 for Triadispora sp., Varirugosisporites roeticus (Schulz) Pittau Demelia in Flaviani, 1980 for Verrucosisporites sp., Verrucatosporites cugiareddu Pittau Demelia in Flaviani, 1980 for Verrucosisporites thuringiacus Mädler 1964. This new interpretation of the data has allowed us to specify the previous dating in the Discussion section.
In Ronchi (1997), two palynological associations (see the complete list of taxa in Supplementary Material) from samples collected in levels known as the Buntsandstein of the Triassic basin of Escalaplano were studied by Pittau. The ARPE-02 sample was obtained at the northeast of the basin (Arcu Is Fronestas) along the Escalaplano-Perdasdefogu road and the ARPE-04 sample at the southern entrance to Escalaplano (Fig. 1B). Both levels were attributed to the Anisian by Pittau (A. Ronchi Pers. Com.), as the presence of Stellapollenites thiergartii is referable to Anisian. Unfortunately, we cannot make any assessment as the work does not present any photos of the taxa mentioned.
In order to formalize the Escalaplano Formation, Costamagna et al. (2000) described three levels with palynomorphs (see the complete list of taxa in Supplementary Material) of two different sections of the "lithofacies B": two levels in a section (nowadays covered by a road wall) not far from the southern entrance to the village of Escalaplano-along the Escalaplano-Orroli road, 2 and 4 m from the underlying contact on the Palaeozoic basementand the third level along a cut on the Escalaplano-Perdasdefogu road in the locality of Arcu Is Fronestas. Based on the presence of Stellapollenites muelleri, Cristianisporites triangulatus and Dyupetalum vicentinense, these levels were considered upper Anisian (Pelsonian), correlating with the "vicentinense-muelleri" phase, according to Brugman (1986), Visscher and Brugman (1981) and Van Der Eem (1983), and the Escalaplano Fm is thus upper Anisian-basal Ladinian.
Observation of the photographs allows us to verify the referenced taxa and propose a new attribution for some of them, as appropriate. We consider that the form attributed to Dyupetallum vicentinense could also be Cristianisporites triangulatus, among others (see supplementary material). According to the authors, the coexistence of these taxa would justify a late Anisian (Pelsonian-Illyrian interval) to early Ladinian attribution. Unfortunately, it is impossible to know the stratigraphic relationship between the studied samples and their relative position since these authors do not provide separate lists and do not attribute their published photos to specific sampled levels.
The last published reference would be Pittau and Del Rio (2002), in which the authors publish a compilation of all the previous palynostratigraphic works on the Permo-Triassic sequences of Sardinia. For the Scythian to early Anisian interval, the authors refer to a listed association (with only three photos, however) consisting of Enzonalasporites leschikii Mädler 1964, Microcachryidites fastidiosus Jansonius 1962, Sulcatisporites splendens Leschik 1955, Triadispora crassa Klaus 1964, cf. Stellapollenites muelleri (Reinardt & Schmitz) Pittau Demelia 1993 andVoltziaceosporites heteromorpha Klaus 1964. This is the same association first described by Flaviani (1980) and related to the Permian in Pittau Demelia and Flaviani (1982b) for level 545 (sample 970). In this work, the authors reevaluate this pollen association and consider a possible Olenekian (late Scythian) or, more probably, early Anisian due to the impossibility of affirming the identification of Stellapollentes muelleri (= Hexasaccites muelleri). However, Voltziaceaesporites heteromorpha made its first appearance in the late Olenekian, according to Brugman (1986). This deduction based on the criterion of absence is not acceptable, and therefore the attribution of level to the "Scythian" is similarly unacceptable. The following section of this compilation is dedicated to the Anisian and it repeats the arguments presented in Pittau Demelia & Flaviani, (1982b), Barca et al. (1995b), Pittau & Del Rio, (2002) and Costamagna et al. (2000).

Reexamined and new palynological data
The original palynological slices analyzed in Diez (2000) have been revised and re-photographed with better image capture systems, offering a slightly more comprehensive taxa list. These slices correspond to samples PC-2, PC-3, PC-4 ("Escalaplano SW", Fig. 5) and PC-7.2 ("Arcu is Fronestas" section, Fig. 6).
Synthetic plates of all the samples studied are presented (Figs. 8 and 9). The complete palynological assemblages are shown in the supplementary material (Table 1 and Figs. S1-S8). Based on the taxonomic similarity of the palynomorph assemblages found and their stratigraphic location, these assemblages have been grouped into several synthetic associations indicating the same age.

Palynostratigraphical discussion
From the set of results obtained from the review of previous works, as well as from unpublished data and new pollen samples, we can infer that there has been no palynological dating corresponding to the Early Triassic in Sardinia, in contrast with neighbouring paleogeographical regions such as the Iberian Peninsula (Diez et al., 2005) and the South of France (Diez, 2000).
Furthermore, sample 1275 collected in the section of Punta del Lavatoio (Pittau Demelia & Flaviani, 1982a) shows photos of only seven taxa and, although the authors initially ascribed it to the Anisian, this age is not consistent. The three species identified have a wide biozone distribution: Chordasporites singulichorda, which corresponds with the base of the Anisian to the Longobardian (Eshet, 1990); Triadispora plicata, from the Anisian to the Carnian; and Triadispora suspecta, from the Pelsonian to the Carnian (Doubinger & Adloff, 1983). Therefore, our dating would be broader, and a Pelsonian-Longobardian interval is proposed for this sample (Fig. 10). A later publication by Posenato et al. (2002) studied the same outcrop and, through biostratigraphic analyses of various groups of marine invertebrates, attributed the entire section to the Ladinian. Likewise, palynological data from the same Muschelkalk facies from the Monte di Santa Giusta section to the north of Alghero (Carrillat et al., 1999) are consistent with this proposed age. Therefore, since our study is focused on the Anisian, the work of Pittau Demelia & Flaviani (1982a) has not been considered.
As regards the rest of the works referred to in our study, it has been possible to verify the presence of Hexasaccites muelleri, which indicates that all levels can be dated as being of the Anisian age (Visscher & Brugman, 1981;Brugman, 1983;Diez, 2000;Diez et al., 2005;Kürschner & Herngreen, 2010).
Regarding the Cugiareddu well (Fig. 10), although most of the Flaviani (1980) work remains unpublished, the few photos published in Pittau Demelia and Flaviani (1982b) and Pittau and Del Rio (2002) allow us to address the following arguments. In their final work of compilation (Pittau & Del Rio, 2002), the authors reevaluated the palynological association, first described by Flaviani (1980) and referred to in Pittau Demelia and Flaviani (1982b), for level 545 (sample 970), and considered it a possible late Scythian or early Anisian, due to the impossibility of confirming the identification of Stellapollentes muelleri (= Hexasaccites muelleri). However, Voltziaceaesporites heteromorpha made its first appearance in the upper Olenekian, according to Brugman (1986). As we said before, the criterion of absence is not acceptable, but since this publication presents photos of three taxa we can infer from the text that they come from level 545. For this reason, we can confirm the identification of Triadispora crassa and Voltziaceaisporites heteromorpha. Therefore, we agree that V. heteromorpha begins in the upper Scythian, but its biozone extends to the Pelsonian (Brugman, 1983(Brugman, , 1986Diez, 2000;Visscher & Brugman, 1981) (Fig. 10). Furthermore, the Triadispora crassa biozone extends from the Anisian to the base of the Carnian. If we add the fact that the association is very poor based on the above, we can only say that it is Anisian without much precision and possibly does not extend to Illyrian.
Regarding level 429 (sample 754), we reject the use of the criterion of absence. Moreover, the presence of Ovalopollis pseudoalatus, with a photo in Pittau and Del Rio (2002), does not allow us to say that the level is later than the base of the Pelsonian (Brugman, 1983;Diez, 2000;Doubinger & Bühmann, 1981;Visscher & Brugman, 1981) or be more specific.
Most of the published works (e.g., Pittau Demelia & Del Rio, 1980;Ronchi, 1997;Costamagna et al., 2000 andDiez, 2000), as well as the new data presented in this work, refer to the outcrops of the Sarcidano-Gerrei region. Nevertheless, as we have already explained in the previous section, the data presented in Pittau Demelia & Del Rio (1980), Ronchi (1997), Costamagna et al. (2000) cannot be used, due to their incomplete or non-existent sheets of photographs or the unknown stratigraphic locations (level and section) of each of the taxa photographed and classified. We could only attribute all the levels of these sections to the Anisian for the reasons mentioned above (Fig. 10).
The second section studied in the Escalaplano basin is called Arcu Is Fronestas (Figs. 1D, 6). Four samples were studied but, as mentioned in the previous section, they have been unified in two synthetic associations (see the complete lists of the samples in the supplementary material,) due to their composition and stratigraphic position. The first corresponds to the lower part of the sedimentary unit in Buntsandstein facies composed of samples AIF-2 and AIF-5, and the second to the upper part composed of samples PC-7.2 AIF-9. The two synthetic associations present few differences between them; both can be attributed to the lower Pelsonian, following the same reasoning used for those of Escalaplano, and based on the same taxa sets. The only appreciable difference is the appearance of Cristianisporites triangulatus in the upper-part association, which has its first record in Central Europe in the Pelsonian (Doubinger & Adloff, 1983). Its presence does not allow us to say that the age is more modern than the base samples, but its relative stratigraphic position could be used as a correlation criterion with the Grifo-3 sample in Campumari Section, which also presents this taxon (Fig. 10).
In the Iglesiente region ( Fig. 1), the first reference is that of Pittau Demelia & Del Rio (1980), in which a poor association is described for the Campumari outcrop. Due to the presence of Hexasaccites muelleri, the sampled unit can be attributed to Anisian sensu lato, as suggested by the authors (Fig. 10). We can confirm this attribution thanks to the photo of the taxon, as discussed in the previous section.
Furthermore, in the same region, we carried out a new sampling with a productive result in the section of Su Passu Malu (Figs. 1D, 7), where, as we reported in the previous section, a rich well-preserved Anisian palynological association with Cristianisporites triangulatus, Enzonalasporites vigens, Illinites chitonoides, Triadispora falcata, T. plicata and T. falcata together with Hexasaccites muelleri would indicates a Pelsonian-Illyrian age (Brugman, 1983;Diez, 2000;Doubinger & Adloff, 1983;Visscher & Brugman, 1981), which would be restricted by the presence of Duplicisporites granulatus, whose genus has its first record in Illyrian.
In addition to the data presented in Barca et al. (1995b), new samplings attempted to provide fresh data from the Scivu Is Arenas section, in the Arburese region (Figs. 1C, 4), but they were all unsuccessful. However, thanks to the partial figuration of the aforementioned paper, a taxonomic reinterpretation of this section was attempted (see also the supplementary data). Therefore, a more precise age for the basal marly levels of the carbonate member of the Punta Su Nuraxi Formation has now been provided. As the authors correctly deduce the presence of Hexasaccites muelleri and Cristianisporites triangulatus, they consider that the association would be from the Pelsonian-Illyrian age (Doubinger & Bühmann, 1981;Doubinger & Adloff, 1980;Visscher & Brugman, 1981;Brugman, 1983;Eshet, 1990;Diez, 2000;Diez et al. 2010), which would be supported by the coexistence with Cyclotriletes oligogranifer and Triadispora suspecta. However, a reinterpretation of the published photos allows us to identify Punctatisporites triassicus and Verrucosisporites thuringiacus, which, on the basis of their last records, would restrict the dating to the Pelsonian (Doubinger & Bühmann, 1981;Doubinger & Adloff, 1980) (Fig. 10).

Paleogeographic inferences
The precise configuration of the Western Tethys during the Middle Triassic, and in particular the position of Sardinia in this domain, has always been subject to debate (e.g., Costamagna & Barca, 2002). The data presented in this work shed more light on this topic, as the microfloristic associations studied, along with the regional facies correlations, enhance our understanding of the timings of the Western Tethys transgression in the western Mediterranean Triassic domain. This allows us to propose a new hypothesis regarding the paleogeographical evolution of this domain during the Anisian. It has been possible to identify three important steps/stages of evolution: 1) First step (late Pelsonian, Fig. 11a). The northwestern and southeastern part of Sardinia (data from Cugiareddu well, Nurra region and ourcrops in the Sarcidano region) were the first sectors of the island to be affected by the first transgressive pulse, which occurred through the development of small seaways that previously crossed the southern Catalan basin and parts of the eastern Iberian basin (Fig. 12), determining the deposition of the lower Muschelkalk facies or Landete Fm, represented by a shallow marine carbonate platform environment (Escudero-Mozo et al., 2015). During this period, most of the islands of Sardinia and Minorca still constituted high terrestrial areas (Costamagna & Barca, 2002;Escudero-Mozo et al., 2014) where erosion or continental sedimentation persisted. At this stage, these elevated areas acted as a barrier between the different Tethys realms and still experienced continental sedimentation in the Buntsandstein facies. 2) Second step (early Illyrian, Fig. 11b). As the transgression progressed, during the Pelsonian/Illyrian, there was a gradual evolution to a more transitional marine environment. Following this new sea level rise, related to the northward shift of Cimmeria (Escudero-Mozo et al., 2014, more areas were progressively covered by epicontinental seas, including the southernmost sector of Sardinia (Sulcis-Iglesiente region), while Minorca and the northern Catalonian basin (Escudero-Mozo et al., 2014 and the eastern Castellón area (López-Gómez et al., 1998;Escudero-Mozo et al., 2015;Ortí et al., 2020) in eastern Iberia were still represented by continental elevated areas during the middle Illyrian (Fig. 12). This configuration represented an important paleogeographic barrier in the Tethys Sea and it determined the distribution of fauna in northern and southern Cimmeria (Escudero-Mozo et al., 2015). 3) Third step (Fassanian, Fig. 11c). The new transitional pulse that took place during the middle-late Illyrian represented the complete incursion of the Tethys sea in its western domain, including central-eastern Iberia and the Balearic Islands, as well as most of Central and Southern Europe (Gianolla & Jaquin, 1998;Ziegler, 1999;  (2001), Stampfli and Borel (2002) and Muttoni et al. (2009) Sanz De Galdeano et al., 2001;Martín-Algarra & Vera, 2004;Pérez-López & Pérez-Valera, 2007;Martín-Rojas et al., 2009;Mercedes-Martín et al., 2013;Escudero-Mozo et al., 2015;Ortí et al., 2020). Over the course of this step, the evaporitic conditions (sabkha environment) which characterized most of the upper Anisian in Sardinia eventually gave way to a shallow marine environment in the whole island.

Conclusions
The new stratigraphic analyses and palynological samplings presented in this work, as well as the close review of all the previous palynological publications on the Sardinian Anisian, enhance our framing of the stratigraphical, sedimentological and palynological aspects of the three key Middle Triassic successions of Sardinia: the Su Passu Malu section (Campumari, SW Sardinia), the Arcu is Fronestas section and the Escalaplano SW section (Escalaplano, Central Sardinia). Moreover, the integration of these data with those obtained from other sectors of Sardinia and Western Europe make it possible to evaluate the paleogeographical evolution of the Western Tethys realm during the late Anisian-early Ladinian with greater precision. The main conclusions can be summarized as follows: 1) The palynostratigraphical associations analyzed provided precise data on the age of the studied sections. It was possible to attribute the Escalaplano Formation to the Pelsonian, thanks to the data obtained from the Escalaplano SW and Arcu is Fronestas sections. As regards the Campumari succession, the sample from Su Pasu Malu Member (Campumari Formation) gave an Illyrian age-constraint. No indications of Early Triassic or lower Anisian associations were identified. 2) These data allow us to define with greater accuracy the Tethys Sea incursion towards its westernmost domains during the Middle Triassic, as well as the different stages of the transgression in Sardinia during the late Anisianearly Ladinian. 3) Sardinia, together with Minorca and the eastern area of Castellón (easternmost Iberia), constituted a topographical high area until the middle-late Illyrian (late Anisian), when an important change in the configuration of the Western Tethys realm occured. Up until that point, the Sardinia-Minorca block had played a key role by acting as a barrier between the realms of the Paleotethys and Neotethys seas. 4) From the late Illyrian to the early Fassanian, the new transgression of the Tethys Sea progressively covered Sardinia with shallow marine deposits that also reached the neighbouring easternmost Iberia and Minorca areas. During this transgression, almost all the elevated areas of the Western Tethys domain were finally covered by the shallow seas.