Drowning, extinction, and subsequent facies development of the Devonian Hönne Valley Reef (northern Rhenish Massif, Germany)

The Hagen-Balve Reef is one of the largest Devonian carbonate complexes in the Rhenish Massif exposed in many former or active, economically significant quarries, especially in the Hönne Valley region at its eastern end. The timing and patterns of reef drowning, final extinction, and the middle Frasnian to middle Famennian post-reefal facies history, including details of the global Kellwasser Crisis, were studied based on two boreholes (HON_1101 and B102) and one outcrop at the Beul near Eisborn. More than 100 conodont samples provided a fine biostratigraphic framework and included new forms left in open nomenclature. The ca. upper 80 m of the new Asbeck Member of the Hagen-Balve Formation consists of relatively monotonous lagoonal successions assigned to four microfacies types. The local diversity of reef builders, mostly stromatoporoids, is low. Fenestral microbialites indicate very shallow and rather hostile back-reef settings. Near the Middle/Upper Devonian boundary, the eustatic pulses of the global Frasnes Events led to a significant backstepping of the reef margin, with reef core/outer slope facies overlying lagoonal facies. This flooding drastically reduced the carbonate accumulation rate and enabled the invasion of drowned back-reef areas by open-water organisms, such as polygnathid conodonts. Within this Eisborn Member, five microfacies types and numerous subtypes are distinguished including low-diversity “coral gardens” and a final, top lower Frasnian parabiostrome dominated by tabulate and colonial rugose corals. There was no cap stage (“Iberg Facies”). Two phases of the Basal Frasnes Event are marked by dark, organic rich limestones with subordinate reef builders. Based on conodont fauna from overlying nodular limestones of the new, (hemi-)pelagic Beul Formation, the final Hönne Valley reef extinction was caused by the eustatic Middlesex Event at the lower/middle Frasnian boundary. Within the Beul Formation, eight subphotic submarine rise microfacies types are distinguished. After a lower middle Frasnian phase of extreme condensation, rich conodont faunas enable the recognition of most upper Frasnian to middle Famennian zones. The global semichatovae Event led to a regionally unique intercalation by four phases of organic-rich, laminated black shales and intervening thin limestones in core HON_1101. The Lower Kellwasser Event is represented in HON_1101 by atypical, moderately Corg-rich, recrystallized, peloidal ostracod-mollusk pack-grainstones. The Upper Kellwasser level begins with an ostracod bloom, followed either by recrystallized mollusk wacke-packstones (HON_1101) or laminated, argillaceous mudstones (B102). The first indicates a rarely documented shallow subphotic, better oxygenated setting than typical Upper Kellwasser facies. As elsewhere, the top-Frasnian conodont extinction was severe. The lower/middle Famennian carbonate microfacies of the Beul Formation is relatively monotonous and typical for an oxic, pelagic submarine rise. The youngest recorded nodular limestones fall in the Palmatolepis marginifera utahensis Zone. Regionally uniform lydites of the Hardt Formation show that the local palaeotopography was levelled before the base of the Viséan. The Hönne Valley case study and comparisons with western parts of the Hagen-Balve Reef and other Rhenish reefs underline the significance of Givetian to middle Frasnian eustatic and anoxic events as causes for reef extinctions.


Introduction
On a global scale, the Givetian and Frasnian were the time of maximum Phanerozoic reef growth (e.g. Kiessling et al. 1999;Flügel and Kiessling 2002;Copper 2002a;Copper and Scotese 2003). In this context, the Givetian-Frasnian Hönne Valley Reef, as the northeastern part of the elongated Hagen-Balve Reef Complex, is one of the best-known examples in the northern Rhenish Massif. The Hagen-Balve Reef Belt is exposed in many outcrops along the northern, eastern, and southeastern margin of the 1 st order Remscheid-Altena Anticline between Hagen in the West and Balve in the East (compare Becker et al. 2016c; Fig. 1).
The complete disappearance of metazoan reefs at the end of the Frasnian, during the global Kellwasser Crisis, was dramatic (e.g. Copper 2002b) but is not well understood (Racki 2005). It is intriguing that the reef majority did not reach the end-Frasnian with its anoxic to hypoxic Upper Kellwasser level. This is especially true for the Rhenish Massif (e.g. Eder and Franke 1982;Stritzke 1989;Aboussalam and Becker 2016;Brinkmann and Stoppel, in press) or westwards, for the Ardennes Shelf (e.g. Denayer and Poty 2010;Mottequin et al. 2015; Mottequin and Poty 2016). The terminal Frasnian transgression and spread of hypoxia killed in Central Europe obviously only small biostromes and the last remaining carbonate platforms. The latter differed from the main Givetian/Frasnian reefs since they were predominantly formed by calcimicrobes and harbored only subordinate last stromatoporoids, rugose and tabulate corals (e.g. Elbingerode Reef, Fuchs 1990;Wülfrath Reef, Becker et al. 2016b;; upper Frasnian biostrome south of Heiligenhaus, Becker et al. 2016b;Hartenfels et al. 2018). In this context, there is a surprising lack of modern research on Rhenish Devonian reef extinctions, with very few highresolution data concerning the precise timing of extinctions and of possible stepwise drowning episodes, the palaeoecology of terminal reef stages, and facies-based reconstructions of triggering factors. An exception is the rarely quoted Ph.D. study for reefal limestones of the Lahn Syncline by Oetken (1997). Eder and Franke (1982) claimed that local factors caused variable pre-Kellwasser extinctions. Is this generally true or did the now wellestablished sequence of 2 nd to 4 th order global events (summarised by Becker et al. 2016a, 2020) play a major role? The latter interpretation has been preferred for reefal episodes of the Aachen region  and of the Velbert Anticline (Wülfrath, e.g. Becker et al. 2016b;Hofermühle;Becker et al. 2016b). These questions are significant for a deeper understanding of global reef megacycles (Hladil 1986) and a refined view on global fluctuations of reef abundance in time and space (e.g. Kiessling 2008).
In the frame of a cooperation project, the Lhoist Germany Rheinkalk GmbH provided drill cores and access to their quarries in the Hönne Valley region. This enabled us to analyse the regional microfacies developments and biostratigraphy of the upper parts of the Devonian carbonate succession, from the final reef stage (the up to 200 m thick "Upper Micrite Formation", compare Eiserlo 1987) through the open platform Eisborn Member of Becker et al. (2016b) to post-reefal nodular limestones. Applying the wide definition of the term reef by Flügel and Kiessling (2002), we focused on the following approaches: 1. Detailed, bed-by-bed logging of macroscopic lithology and macrofauna in drill cores HON_1101 and B102, with a focus on the reef builders of the last reef stage. 2. Microfacies documentation and analysis of all carbonate types, with emphasis on the, so far, poorly known final reef drowning stages. 3. Continuous conodont sampling and biostratigraphic dating, applying the most recent conodont scales, recognising new and rare forms, and with a focus on the hardly known faunas from final reef stages. 4. Lithological and biostratigraphical correlation between the two cores and the re-sampled Beul outcrop of Nowak (2010) and Becker et al. (2016c). 5. Reconstruction of palaeoecological trends and the precise timing of the stepwise regional reef extinction and postreefal history in the context of known global events.
Our study yielded the most detailed data set for the extinction of any German Devonian reef. On a global scale, there are few other examples, where microfacies successions of a drowning Givetian carbonate platform have been documented with a similarly precise biostratigraphic scale. This will enable future comparisons and provide a base for the recognition of potential patterns that cannot be concluded from the study of a single reef. A refined knowledge of Middle Palaeozoic reef extinctions holds clues for a better general understanding how reefs reacted and developed in times of biotic crises. Coral and stromatoporoid genera: Alv. = Alveolites, Am. = Amphipora, Hex. = Hexagonaria, K. = Keega, St. = Stachyodes, Th. = Thamnopora.

Repository
All specimens are stored in the Geomuseum of the Westfälische Wilhelms University Münster (GMM), conodont originals under B9A.14, brachiopods under B5B.17, ammonoids under B6C.56. Slides with the majority of other conodonts and thin-sections with reef faunas are stored according to their localities, bed numbers, or core depth.

Regional geological setting
The lower Givetian of the Remscheid-Altena Anticline, located east of the Rhine river, belonged to the southeastern shelf of Laurussia (Old Red Continent), which was characterised by the deposition of prodeltaic siliciclastics (Honsel Formations, compare Hartkopf-Fröder and Weber 2016, fig. 2). Only small biostromes formed in protected positions or on small topographic highs (e.g. May 1983May , 1992May , 1994bMay , 2003. These lasted only for brief episodes and were buried by shifting silt-and sandstone wedges. Higher in the lower Givetian succession, the detrital discharge decreased strongly, probably because of increasing aridity in the Old Red source area (Becker et al. 2016c) or by transgressive pulses. Consequently, reef growth was established widely in the Rhenish Massif, ranging from small, isolated biostromes to laterally extensive and hundreds of metres thick bioherms that are of economic significance (e.g. Jux 1960aJux , 1960bKrebs 1971Krebs , 1974Burchette 1981). East of the Rhine river, well-known examples are the reefs of Hofermühle, Wülfrath, Neandertal, Wuppertal, Hagen-Balve, Attendorn, Warstein, and Brilon (Fig. 1). Traditionally, very different reefal carbonates, including marginal slope debris, were summarised under the term "Massenkalk", which referred to the hardness and often poor stratification of the rocks. The rock name was adopted as a common formation name on many geological maps, even if there was clearly no continuity between the individual reefs and irrespective of different facies developments and age. Although the term "Massenkalk" is still used, e.g. by workers of limestone quarries and industrial plants, it is more correct to call such reefal deposits carbonate complexes (Krebs 1974) or, after Burchette (1981), reef complexes, as most build-ups are partly stratified. Regionally, different formation names (e.g. Hartkopf-Fröder and Weber 2016, Bergisch Gladbach-Paffrath Syncline: Büchel Formation; Becker et al. 2016b, Wülfrath: Wülfrath Formation;Becker et al. 2016c, Hagen to Hönne Valley: Hagen-Balve Formation) have been introduced but this process of lithostratigraphical revisions has not yet been completed.
The studied reefal limestones of the Hönne Valley region belong to the Hagen-Balve Reef Complex (Fig. 1) and Hagen-Balve Formation. Following the Variscan strike direction, it stretches for more than 20 km along the northern limb and NE-diving axial ramp of the Remscheid-Altena Anticline (Fig. 1). The reef belt has been exploited economically in numerous quarries along its entire range but active quarrying concentrates at the western (Donnerkuhle Quarry at Hagen-Eppenhausen, Steltenberg Quarry at Hagen-Hohenlimburg; Fig. 1), and eastern (Hemer-Becke and Hönne Valley) ends. The elongated structure probably represents the eroded remnants of a fringing reef north of a structural high in the core of the anticline (Paproth 1986). A maximum thickness of nearly 1000 m of Givetian shallow water carbonates accumulated in the Hönne Valley region. This enormous carbonate succession proves a period of stable conditions, in which reef growth and subsidence were in balance. As mentioned by Becker et al. (2016c), the old geological map by Paeckelmann (1938) displays a complex pattern of numerous faults. However, recent mapping by Münster (e.g. Kruse 2013;Richter 2013) and Cologne (Bahr 2021) students clearly suggest a smaller fault number.

Research history
Detailed geological research started in the Hönne Valley in the early 20th century with the pioneer work of Denckmann and Lotz (1900) and Denckmann (1901aDenckmann ( , 1903Denckmann ( , 1905 in the frame of the initial geological mapping by the Prussian Geological Survey. Paeckelmann (1924Paeckelmann ( , 1938 continued the mapping and added details concerning faunas, lithostratigraphy, and local facies developments. For example, he separated the true reef limestones from marginal slope deposits and lower slope to off-reef, dark, organic-rich Flinz facies. He also described the condensed, post-reefal carbonates and adjacent basinal shales. The lithostratigraphic scheme developed by the time does not agree with modern standard terminology but has hardly been revised since. Therefore, we introduce two new formations and two new members. Jux (1960aJux ( , 1960b studied Hönne Valley quarries in his general, descriptive review of Rhenish Devonian reefs. As a supposed typical example, he logged the now abandoned Emil Quarry west of the valley (Fig. 1) and documented the different, thin-bedded "Massenkalk" facies further to the south, near Garbeck and at the famous Balve Cave (Fig. 1). Subsequently, Eder (1970Eder ( , 1971) analysed this Givetian Garbeck Limestone and recognised it as the marginal slope deposits of the southern Hönne Valley Reef. In the huge Asbeck Quarry (Fig. 2), Schudack (1993) logged the middle Givetian inner platform succession and recognised cyclic reef growth, which was explained by relative sea-level change due to regional tectonic movements since the lack of biostratigraphical data prevented a correlation with eustatic changes. Strutz (2004) analysed a drill core (HON 01/02) of the Beul area with special emphasis on the microfacies of the upper lagoonal part of the Asbeck succession. He recognised the potential of thin bentonites for regional correlation. Polenz (2008) published a popular scientific booklet with geological, historical, and recent ecological  Nowak (2010) dealt with the youngest reef deposits at the Beul, whilst the B.Sc. Thesis of Stichling (2011) was devoted to the microfacies and stratigraphy of the adjacent drill core B102. Some of the results are presented herein. Becker et al. (2016c) provided an overview of the reef development in the Hönne Valley, with a more complete compilation of former research, first data on the initial phase at Binolen, the main reef phase of Emil Quarry (Fig. 1), and the final phase on the Beul outcrop, including its carbon isotope stratigraphy. Preliminary data of our joint project with the Lhoist Germany Rheinkalk GmbH were presented during the IGCP 596-SDS Symposium in Bruxelles and the post-meeting excursion (Stichling et al. 2015;Becker et al. 2016c), Closing Meeting of IGCP 591 at Ghent (Stichling et al. 2016), and at the ICOS IV in Valencia (Stichling et al. 2017).

Lithostratigraphy
The Hönne Valley sedimentary history and palaeogeography were complex due to the interplay of laterally variable carbonate facies/accumulation, synsedimentary tectonics (fluctuating subsidence), and volcanism. This resulted in a small-scale palaeotopography with either local or more widespread lithological units. The thick reefal succession is underlain by the lower Givetian Upper Honsel Formation, which represents an offshore, fine siliciclastic prodelta depositional system (Çinar 1978). Embedded are small-sized lenses of biostromal limestones with reef corals (e.g. May and Marks 2014). They represent initial reef phases that failed due to renewed high clastic influxes.
The thick succession of reef carbonates started with a sharp boundary after the shedding of silt and fine sand ended completely (Becker et al. 2016c). This Hagen-Balve Formation is subdivided vertically and laterally into different members ( Fig.  3). At the base, the Binolen Member, described by Löw et al. (2022), comprises the initial, dark limestones with corals, stromatoporoids, brachiopods, and abundant crinoid debris. It represents the Schwelm Facies sensu Krebs (1974). Dated by brachiopods and conodonts, biostrome growth began in higher parts of the lower Givetian.
The succession turns into the main reef stage, the biohermal Dorp Facies, which characterises the new Asbeck Member. It is named after its type locality, the Asbeck Quarry (Fig. 2). The base is exposed at Binolen (Löw et al. 2022, this issue) and defined by the first appearance of back reef limestones, which reflect the change from an open platform into a bioherm with a protected inner lagoon. From the Asbeck Quarry, Schudack (1993) described more than 600 m of cyclic lagoonal and reef core facies and distinguished six major facies associations: reef-core, sublagoonal, lagoonal, restricted lagoonal, intertidal, and supratidal environments. His five "formation" names do not agree with the standard lithostratigraphic nomenclature. They are treated by us as subunits (submembers) of the Asbeck Member. The first and oldest stage, the "Lower Micrite Formation", is characterised by micritic limestones and fenestral fabrics; it falls in the lower/ middle Givetian due to the presence of Stringocephalus. The "Lower Stromatoporoid-Limestone Formation" represents the second stage with extensive reef growth by massive stromatoporoids. It is followed by a mixed phase that exhibits micritic lagoonal deposits as well as reefal carbonates dominated by large stromatoporoids, the so-called "Micrite-Stromatoporoid-Formation". The overlying unit is, again, a stromatoporoid reefal stage, the "Upper Stromatoporoid-Limestone Formation". Micritic lagoonal carbonates dominate the last phase, the "Upper Micrite Formation". The top of the reef is not exposed in the Asbeck Quarry.
The term "Eskesberg Facies" was introduced by Krebs (1974) for well-bedded, organic rich, and dark back reef facies. These are well-exposed in the northern wall of the abandoned Emil Quarry west of the Hönne Valley (see Becker et al. 2016c). They deposited during deepening phases in concert with decreasing bottom turbulence while organic productivity was high. In the Asbeck Quarry, a corresponding thick black marl interrupts as a marker interval the backreef limestones. The lagoonal Asbeck Member at Emil is well stratified, differentiated (Jux 1960a), and subdivided by a marker bentonite (Becker et al. 2016c, figs. 8-9). The source of the volcaniclastics lay in the south, in the Balve area, where thick volcanic sequences occur. The Emil succession is dated as lower/middle Givetian by storm layers enriched in mostly fragmented, thick-shelled Stringocephalus (e.g. Becker et al. 2016c, fig. 10).
The Eisborn Member, introduced by Becker et al. (2016c), is characterised by a strongly reduced carbonate accumulation and an influx of open shelf organisms, such as conodonts, brachiopods, and trilobites. The type-section is a small natural cliff in the forest at the Beul (Locality 3 in Fig.  2). Denckmann (1903) and Paeckelmann (1924Paeckelmann ( , 1938 described from the top of the reef succession locally fossiliferous crinoidal limestones containing brachiopods. Such proximal slope deposits resemble the well-bedded Schleddenhof Member of the western parts of the Hagen-Balve Reef Complex (Fig. 3), especially from Iserlohn-Letmathe (Beckmann 1961) and from the famous brachiopod locality at Iserlohn-Bilveringsen (e.g. Torley 1908Torley , 1934Fig. 1).
Outcrops to the south, from near the Balve Cave to Garbeck (Fig. 1), exhibit the mentioned Garbeck Limestone that was studied intensively by Eder (1970Eder ( , 1971; it is re-named as Garbeck Member of the Hagen-Balve Formation. It represents an intergradation and transition from reefal to marginal slope sediments including proximal debris flows with corals and stromatoporoids, crinoid limestones, and organic-rich turbidites as well as darkgrey shales/marls. Conodont re-sampling yielded only sparse faunas from 3-4 kg samples. Records of L. weddigei from the upper 8-10 m confirmed a lower/middle Givetian age, in accord with Stringocephalus occurrences (Paeckelmann 1938) and previous conodont data (Eder 1971;Dornsiepen 1973). The Garbeck Member accumulated synchronously with the Asbeck Member on the southern reef slope.
At Balve (Fig. 1), marginal slope limestones grade into contemporaneous volcaniclastics, tuffites, and overlying iron ore bodies. The proposed new Husenberg Formation is named after the Husenberg, the type locality forming the eastern end of the Balve village. The lower and upper boundaries of the formation are defined by the onset and disappearance of volcanic rocks. Locally, there are intercalated middle Givetian debris limestones (Paeckelmann 1938;Dornsiepen 1973), resulting in a total thickness of up to 290 m (Paeckelmann 1938). As mentioned before, the pyroclastic complex is the source for thin bentonites intercalated in the reef platform succession in the north.
A steep reef slope, with dark turbiditic limestones and shales ("Flinz Facies") existed also just west of the Hönne Valley. In the Emil Quarry, Jux (1960a) noted black Flinz beds that followed above a 3 m thick upper slope reef breccia. Unfortunately, this important level has been mined away. A similar succession continues westwards towards Hemer-Apricke (Spiske 2003;Fig. 3).
The Asbeck and Horst quarries ( Fig. 2) sit on the axial ramp of the Remscheid-Altena Anticline and represent the central area of the Hönne Valley Reef, which turned after its drowning into a large, subphotic submarine rise. This resulted in the accumulation of grey to reddish (hemi-)pelagic limestones with goniatites embracing the two black Kellwasser levels (Paeckelmann 1924) that has no valid lithostratigraphical name. The previously used terms Adorf Limestone and "Nehden-Schichten" (e.g. Paeckelmann 1938) refer to units of the eastern Sauerland that differ partly in facies and stratigraphical range. More importantly, there was no depositional continuity between the eastern pelagic limestones and shales and those of the Hönne region. The type Adorf Limestone (= Adorf Formation) was restricted to small volcanic submarine rises (e.g. Saupe and Becker 2022, this issue), the distinctive goniatite shales of the type Nehden Formation accumulated only in isolated drowned lagoons of the Brilon Reef (Becker 1993;Becker et al. 2016d). The term Cheiloceras Limestone, used in the Hönne Valley region by Paeckelmann (1938), does not conform with modern lithostratigraphical standards and could be confused with the Cheiloceras Limestone of the Aachen region, which has a very restricted stratigraphic position (e.g. Schmidt 1951). Therefore, we propose the new term Beul Formation, defined by the onset of condensed, light-grey nodular or flaser-bedded limestones with pelagic fauna, intercalated by subordinate marls, black shales or limestones, such as the famous Kellwasser Beds. The Upper Kellwasser level provides a lithostratigraphic boundary to separate Lower and Upper Members. The type-section is the natural cliff above our Locality 3 at the Beul, with further outcrops to the north (Schäfer 1978;Hacke 1999). The maximum thickness is ca. 35-40 m.
Laterally, on both limbs of the Remscheid-Altena Anticline, the Beul Formation grades into basinal Frasnian banded or grey-green shales, followed by lower Famennian black shales and sandstones ("Plattensandstein"; Fig. 3), which all lack valid formation names. Paeckelmann (1924Paeckelmann ( , 1938 described the intermediate Frasnian facies, with a mixture of basinal shales, finegrained grey limestones, and dark, turbiditic Flinz limestones. Above the Beul Formation, the change to reddish nodular beds ("Fossley" or "Kramenzelkalk") marks the base of the middle Famennian Hemberg Formation, which has a wide distribution along the northern flank of the Remscheid-Altena Anticline (e.g. Becker 1992; Hartenfels and Becker 2016a) and beyond (e.g. Becker et al. 2016c). Upper Famennian to lower Tournaisian nodular limestones and the middle Tournaisian Lower Alum Shale deposited only on parts of the submarine high (Paeckelmann 1938;Kruse 2013;discussion in Söte et al. 2017;Hartenfels et al. 2022, this issue.). The final drowning of the pelagic platform occurred around the Tournaisian/Viséan boundary, when the uniform lydite facies of the Hardt Formation covered the whole region. At least since the Upper Carboniferous, the area of the Remscheid-Altena Anticline was continuously a terrestrial high. Therefore, younger pre-Quarternary sediments are restricted in the Hönne Valley to Permian redbeds (Menden Conglomerate), Cretaceous karst fillings, and Tertiary weathering products (Polenz 2008).

Studied localities and drill cores
Drill core HON_1101 (coordinates: x = 34 21.227, y = 56 95.936m, map sheet 4613 Balve) was sunk by the Lhoist Germany Rheinkalk GmbH and reached a depth of nearly 160 metres. The borehole is located northeast of the Asbeck Quarry (Fig. 2). As Holocene sediments are either thin or absent, most core material consists of thick reefal limestone (upper ca. 70 m of Asbeck Member, interrupted in the upper part by a 6.7 m thick fault zone), overlain by coarse debris correlating with the Eisborn Member (ca. 12 m), (hemi-)pelagic Frasnian to Famennian limestones (Beul Formation,ca. 19.5 m middle/upper Frasnian, ca. 15 m lower/middle Famennian), followed by ca. 5.5 m of the locally argillaceous Hemberg Formation (Fig. 4). The core was sunk in the frame of an exploration campaign for the local mining and was archived for possible future analysis.
Drill core B102 (coordinates: x = 34 21.192, y = 56 95.769, map sheet 4613 Balve) was sunk for exploration of the limestone resources in the northeastern Beul area (Fig. 2). It also penetrated the upper part of the Asbeck Member and its overlying Upper Devonian (hemi-)pelagic sediments containing the thin Upper Kellwasser level. The final depth was at 106 m (Bed -97, Stichling 2011). The coarse debris interval correlated with the lateral Eisborn Member is only ca. 3 m thick (beds -25 to -27) and overlain by 17 m limestones of the Beul Formation (ca. 12.6 m Frasnian, ca. 4.5 m lower Famennian), and, after a long time of non-deposition, by Viséan cherts (Fig. 5).
Within a forest at the northeastern end of the Asbeck Quarry, the Beul outcrop forms a small cliff and the highest topographic elevation in the adjacent area (Fig. 2). It is located at the eastern end of the Beul hill (x = 34 21.470, y = 56 95.590, map sheet 4613 Balve), approximately 800 m to the northwest of the centre of the village Eisborn. Hacke (1999) recorded it as his outcrop AB6. Access is possible by following in Eisborn the road "Zum Blechen" to the west, leading after some curves eventually to a track branching, where the main downslope track should be taken. The latter is barred at the entrance of the quarry area. Since this is an area of active mining, a permission by the Lhoist Germany Rheinkalk GmbH is mandatory. Nowadays, it is the only location in the Hönne Valley region, where the drowning stage (Eisborn Member) and following (hemi-)pelagic nodular limestone succession (Beul Formation) can be investigated. An adjacent second cliff noted by Hacke (1999) as Outcrop AB5 exposes higher parts of the Beul Formation. Based on the B.Sc. Thesis of Nowak (2010), a summary of the succession and remarks on the younger Famennian strata were published by Becker et al. (2016c).

Methods
Between 2014 and summer 2022, repeated fieldwork provided more than 30 conodont and 53 microfacies samples from the Beul outcrop. During the logging of cores HON_1101 and B102, macrofauna and lithofacies were recorded continuously, with a focus on vertical facies changes. Our study concentrated on the final phase of local reef development, from the upper part of the Asbeck Member, through the Eisborn Member and overlying Beul Formation. The main part of the Asbeck Member has been sufficiently studied by Schudack (1993) and Strutz (2004).
All lithological types were sampled for thin-sections, using sizes of 75 x 100 mm or 100 x 150 mm. Overall 77 conodont samples were taken from both cores and 124 microfacies thinsections were produced from core halves. The microfacies analysis was conducted by using transmitted light microscopes (Leica MZ 6, Leica MZ 12.5, fully motorized Keyence VHX). Facies interpretations follow the Dunham classification for carbonate rocks (Dunham 1962;Embry and Klovan 1971), the characterisation of Facies Zones (FZ) after Wilson (1975), the Standard Microfacies Types (SM) sensu Flügel (1978Flügel ( , 2004, and updates by Hartenfels (2011). These formed the frame for comparisons with Devonian microfacies analyses of other authors.
Conodont samples taken from the core material weighted 300 to 400 g, whereas Beul samples weighted at least 3 kg, since their overall yield was low. The limestones were dissolved by using a 10 % solution of formic acid, with washed residues separated into 0.100-0.315, 0.315-0.630, and > 0.630 mm fractions. The residues of the finest fraction were subject to heavy liquid separation using diluted sodium polytungstate (3Na 2 WO 4 × 9WO 3 × H 2 O, 2.76-2.78 g ml -1 ). The samples yielded between 0 and more than 1.000 platform elements per kg, with peak abundances in Famennian nodular limestones. Because many multi-element reconstructions are still doubtful, mostly the Pa element taxonomy has been utilised, with the exception of Pa. winchelli (= subrecta) and Pa. bogartensis (= rotunda) in the Frasnian. Ramiform elements were picked but not identified; only supposed Pb elements with platforms (Nothognathella) were recorded. Biostratigraphic dating applied the revised zonations of Aboussalam (2003), Aboussalam and Becker (2007), and Narkiewicz and Bultynck (2010) for the Givetian, the Montagne Noire (MN) zonation sensu Klapper (1989), with updates in Girard et al. (2005), Becker et al. (2016aBecker et al. ( , 2020, and Pisarzowska et al. (2020) for the Frasnian. Spalletta et al. (2017) revised the terminology of Famennian zones, but we refer also to the traditional zonation of Sandberg (1984, 1990) and subdivisions proposed by Schülke (1995). Representative conodonts were photographed with a SEM at Münster.

Microfacies analysis
Three facies sets are distinguished that represent the Asbeck Member (MF-A), Eisborn Member (MF-B), and the Beul Formation (MF-C). A coarse detrital facies aligned with Set A is intermediate to Set B. It represents the reefal backstepping correlating in time with the initial lagoon drowning. The distribution of MF-types in the Rhenish Massif and, partly, beyond is given in order to place the Hönne Valley microfacies into a wider context, especially since there is no published review on Givetian/Frasnian MF-type distributions that could be quoted.
A variant, MF-A1*, is characterised in parts of core B102 (e.g. 101.10-101.38 m) by a very low content of branching stromatoporoids; it is intermediate towards wackestones with reef builders. Interpretation: MF-A1 is wide-spread in the upper Asbeck Member both in HON_1101 and B102. Following Krebs (1974), Stachyodes prefers niches between more massive, bulbous reef builders. In back reef environments, it is characteristic for moderately agitated settings (Eichholt and Becker 2016). In contrast to the more delicate Amphipora branches, Stachyodes was stronger and, therefore, more tolerant to currents and wave activity, which is noticeable by occasional micrite winnowing. The co-occurrence of Amphipora and calcispheres/parathuramminid foraminifers speaks for deposition in a relatively calm lagoonal environment (Flügel 2004). The branching stromatoporoids may have formed small patch reefs within the back-reef. Units with poor macrofauna (MF-A1*) accumulated far from these and received mostly finer detritus. According to the modell sensu Wilson (1975), MF-A1 can be integrated in FZ 7 and SMF 7 sensu Flügel (2004). Distribution: Comparable stromatoporoid float-rudstones have been widely described from Rhenish reefs. Krebs (1974), May (1987), Fuchs (1990), Malmsheimer et al. (1991), Schudack (1993), andBecker et al. (2016c) reported this facies type from the Dornap, Hagen-Balve, and Brilon reef complexes. Weller (1991) described a similar facies type from the Elbingerode Reef Complex of the Harz Mountains. A Stachyodes dominated variety is mentioned by Faber (1980) from the Eifel Mountains. Further occurrences are given by Flick and Schmidt (1987) from small atolls in the southern part of the Rhenish Massif. Krebs (1974) described a dark, bulbous stromatoporoid-Amphipora facies, which, in contrast to MF-A1, contains echinoderms, thamnoporids, and alveolitids. Outside the Rhenish Massif, there are similarities with the "stromatoporoid float-/rud-/boundstone" found in patch reefs of the Oued Cherrat Zone of the Moroccan Meseta (Eichholt and Becker 2016, their MF A5).
MF-A2: Detrital stromatoporoid grain-rudstone (Fig. 6f) Examples: HON_1101, . Description: The medium grey limestone contains a high number of broken stromatoporoids, mostly dendroid Stachyodes and Amphipora; laminar and bulbous forms occur, too. The assemblage is completed by disarticulated, thickshelled brachiopods (Fig. 6f), minor echinoderm debris (probably crinoids), calcispheres/parathuramminid foraminifers, and ostracods. In accordance with MF-A1, tabulate and rugose corals are absent. Bioclasts are subrounded and their size, especially in the case of laminar stromatoporoids, can reach more than 10 cm in exceptional cases. The poorly preserved and unsorted skeletal remains and associated, small-sized micritic lithoclasts are embedded within a peloidal grainstone matrix with microsparitic cement. Some bioclasts show marginally an initial micritisation. Microstylolites are partly strongly developed. Interpretation: MF A-2 was deposited in a back reef setting adjacent to patch reefs and represents a high energy environment laterally to MF-A1. A prominent feature is the mixture of organisms, which originally belonged to different reefal zones. Transport may have occurred by waves, storm events, or in channels (compare the turbulent back-reef Zone IVb of Machel and Hunter 1994). Considering the close relationship to MF-A1, MF-A2 falls in the FZ7 sensu Wilson (1975). Distribution: Similar facies types are known from the Rhenish Massif in the "detrital-stromatoporoid-coral-algae facies" sensu Krebs (1974). MF-A2 is also documented from the Elbingerode Reef Complex, Harz Mountains (Weller 1991). In the western continuation of the Rhenish Shelf, in SW Devon, Garland et al. (1996) described a corresponding MF5 (Stachyodes rud-and grainstones) as "high-energy accumulation of back reef talus". Eichholt and Becker (2016) distinguished a similar "detrital, bioclastic grainstone/rudstone (back reef breccia)" in the Moroccan Meseta and discussed its affiliation with SMF 5 sensu Flügel (2004).

MF-A4: Fenestral pack-grain-bindstone
Description: MF-A4 occurs macroscopically as medium grey limestone with a low fossil content. It is distinguished by dominant fenestral fabrics, ranging from subrounded to elongate, irregular and laminar spar-filled primary cavities ("birdseyes", Ham 1952). The matrix consists mostly of well rounded, equalsized peloids (Fig. 7c). In a few cases, the intergranular spaces are filled with microspar. Rare faunal elements are Amphipora, Stachyodes, other stromatoporoids (bulbous and laminar forms), dendroid corals, parathuramminids, gastropods, ostracods, and indeterminate fragmented shells. Pressure solution led to the common formation of stylolites; calcite veins reflect diagenetic and tectonic stress. Two MF-A4 subtypes are distinguished. Subtype MF-A4a is characterised by numerous, isolated, up to 2.5 cm long, sparitic birdseye structures. Some voids are filled with dark sediment at their bases. Occasionally (Fig. 7c), there are thin, laminated microbial micrite layers marking minor discontinuity surfaces.
Mixed peloidal-fenestral fabrics indicate intertidal conditions, a shallow lagoon with a permanent influence of waves and currents. The habitat was hostile to the majority of reefal organisms. The alternation of mudstones and peloidal packgrainstones is typical for peritidal to even supratidal environments. The poor preservation of the subordinate bioclasts indicates re-deposition. This facies is comparable with FZ 8 of Wilson (1975) and SMF 21 sensu Flügel (2004), Facies 2 sensu Schudack (1993, laminite facies), and Zone 1b, fenestral laminites, of Machel and Hunter (1994). Distribution: Related facies haven been described from Hagen-Hohenlimburg by Koch-Früchtl and Früchtl (1993) and from the Brilon Reef Complex (May 1987;Mestermann 1995). Krebs (1974) described a fenestral facies from the Langenaubach Reef Complex of the southern Rhenish Massif and suggested an intertidal palaeoenvironment with slightly increased salinity. Salinity fluctuations would exclude stenohaline organisms. In SW Devon, MF-A4a was described by Garland et al. (1996) as MF9 (restricted, shallow subtidal to intertidal), MF-A4b as MF12 (restricted intertidal to supratidal ponds).
Description: Macroscopically, there are medium grey, fossiliferous limestones. In thin-section, poorly preserved, broken, angular stromatoporoid fragments, partly with micritic seams, are the most abundant organisms. Broken branches of Stachyodes, rare Amphipora and thamnoporids, laminar alveolitids, and solitary Rugosa are associated. Bioclasts can reach a size up to several cm (Fig. 7d). Abundant fragmented crinoids and rare calcispheres occur in interspaces. The matrix consists of peloids, coarse sparite ( Fig. 7e), or small-sized detritus formed by unidentifiable bioclasts and peloids. The sorting of the components is poor. Crinoid remains are partly surrounded by syntaxial overgrowth (see Fig. 7e). Interpretation: MF-A5 includes a higher diversity of organisms than the previous MFs. We assume a more open environment, from the reef core to the outer reef edge, as the bioclast source. Their brecciated, non-sorted arrangement, poor preservation, the lack of micrite, and the mixture of fauna typical for different settings, points to high-energy deposition by storms and waves. In comparison to the directly underlying back-reef facies types, the arrival of material from the seaward side indicates reef (inner platform) backstepping but channels through the reef core may have facilitated the material transport. MF-A5 can be assigned to FZ 4 (slope) to 5 (platform margin) sensu Wilson (1975). In the scheme of Flügel (2004), it can be classified as SMF 6 and resembles the detrital stromatoporoid-echinoderm facies of Krebs (1974). Roemerolites sp. and with stylolithic dissolution surfaces; slightly darker packstone with small-sized, undefined bioclasts at the base (HON_1101, 78.00-78.15m, Bed -20) g MF-B5i, rudstone with bulbous (left, not clearly identifiable), laminar (Schistodictyon sp., lower centre, Salairella buecheliensis, right), and dendroid (Stachyodes) stromatoporoids, rare thamnoporid fragments, subordinate crinoid debris, and grainstone matrix of peloids, fine bioclasts, and sparite (HON_1101, 83.30-83.51 m, top Bed -22) h MF-B5i, Alveolitid-dominated rudstone (Av Alv. (Alv.) ex gr. edwardsi) with overgrown thamnoporid (Th. ex gr. polyforata) and diagenetic dissolution seams that intrude significantly into bioclasts (HON_1101, 79.12-79.32 m, Bed -20, Eisborn Member) Distribution: This facies type is common in the Rhenish Massif (e.g. Krebs 1974). Machel and Hunter (1994) assigned it to the transition from the reef core to the fore-reef (their zones IV/V to IIIf). In the Moroccan Meseta, Eichholt and Becker (2016) described detrital coral-stromatoporoid floatrudstones as fore reef breccias; MF-A5 is similar to their subtype A.

MF-B: Eisborn Member (biostromal facies types)
The characteristic feature of the Eisborn Member is the combined presence of subordinate open-water biota (conodonts, brachiopods, trilobites), bioturbation, dominance of fine debris (mud-to pack-and grainstones), and of isolated, variably sized fragments of reef builders (dominant tabulate or rugose corals, some stromatoporoids), giving various types of floatstones. They represent "reef gardens" characterised by widely spaced reef builders grading into biostromes (rudboundstones of HON_1101 and of the backside of the Beul outcrop). The faunal spectrum (Table 1) is restricted. Comparisons can be made with initial reef phases (Schwelm Facies, e.g. Koch-Früchtl and Früchtl 1993;Löw et al. 2022, this issue) and Rhenish reefs that remained in the biostrome phase, since both show open water influence. Examples come from the Eifel Mountains (Faber 1980) and small reefs intercalated within siliciclastic units of the Sauerland. The latter have been well-studied for their faunas (e.g. May 1983May , 1994bMay , 2003May and Marks 2014) but not for their microfacies spectrum. Comparable conodont-bearing reef limestones were also described by Oetken (1997) from the Lahn Syncline in the southern Rhenish Massif. Distinctive for the Eisborn Member is the low diversity of microfauna in conodont samples and the absence of microbialites or of coarse slope debris that characterise in other German reef complexes the Iberg/Schlupkothen facies. In the Elbingerode Reef Complex of the Harz Mountains, Weller (1991) assigned some comparable facies types to a "demergence stage".
Interpretation: The dominance of moderately fine reef debris in combination with the occasional larger clasts of reef builders and open-water organisms suggests the setting of a drowned back reef that received with minor currents or during storms the distal talus of the submerged reef margin or from flooded patch reefs. Episodic bottom currents reworked clotted micrite as peloids. Deposition took place well below the fair-weather wave base. Distribution: Equivalents of MF-B1 were included by Koch-Früchtl and Früchtl (1993) in their MF-Typ 4 ("microbioclastic-peloidal wackestone with packstone parts") but the latter occurred in the still open, initial reef phase at Hagen-Hohenlimburg (Steltenberg Reef, Fig. 1), not in the final drowning phase. MF-B1 can also be compared with the peloid-bearing Microfacies I1 ("micritic-arenitic crinoid facies" with reef builder debris) of Rieck and Stritzke (1999) from the atoll of the Messinghausen Anticline. MF-B1 differs from the peloid-rich back-reef Facies Zone IIB ("sparsely fossiliferous packstones and wackestone") of Machel and Hunter (1994) in the presence of open-water biota.

MF-B2: Peloidal grainstone (Fig. 8b)
Examples: Beul, beds B1 (samples from the base and from 110 cm above base) and B2 (partly); B102, higher parts of Bed -27. Description: Macroscopically, it is a light-grey, massive, fine-grained, and macrofossil-poor limestone. In thin-sections, peloids dominate and are embedded in a sparitic matrix. Bioclasts consist of echinoderm or tabulate coral debris (B102, Bed -27) and ostracods, which are partly coated (Beul, Bed 1). Diagenetic overprint led to stylolites. MF-B2 grades into MF-B5g (Sample top reef 1D). Interpretation: MF-B2 differs from MF-B1 in displaying evidence for more constant bottom water agitation, leading to a more complete reworking of micrite to peloids ("pseudopeloids"). It is a drowned variant of the lagoonal SMF 16 of Flügel (2004). Distribution: Equivalents of MF-B2 are known from forereef settings of Eifel biostromes (Faber 1980) but rarely occur in bioherm/lagoon settings. Exceptions were found in reefs of the Harz Mountains (Franke 1973;Weller 1991)  Interpretation: In relation to MF-B1, the higher amount of bioclasts, while the micrite has not been washed out, is somewhat contradictive. There must have been weaker bottom currents but a higher influx of reef debris originating probably from episodic storms. In modern reef environments, fragmentation of reef builders is caused to a large extent by recurrent hurricanes (e.g. Scoffin 1993). Coarse material may remain on the platform while much debris is transported towards the outer slope and lagoon, where it is deposited as tempestites with characteristic features, such as cross-bedding and grading (Aigner 1985). In the Hagen-Balve Reef Complex, typical tempestites are only known from outer slope settings (Eder 1971), not from lagoons (see Schudack 1993). Distribution: From the initial phase of the Hagen-Balve Reef, Koch-Früchtl and Früchtl (1993) described a variant rich in siliciclastic detritus (MF-Typ 1); there is no previous documentation from terminal reef stages. MF-B3 differs from the back-reef Zone IIB of Machel and Hunter (1994) in the presence of open water biota.

MF-B5: Coral float-rudstone
Description: Macroscopically, these are light-to dark-grey, thinbedded limestones with moderately common to very abundant reef builders, mostly tabulate and rugose corals, sometimes with stromatoporoids. In many variants, reef builders float in a finegrained matrix, representing destructed "coral-stromatoporoid gardens". Alveolites is mostly preserved in situ and shows only few signs of damage, whereas dendroid Tabulata are heavily broken. Floatstones can change laterally and vertically to rudboundstones, which are autoparabiostromes and parabiostromes sensu Kershaw (1994 Interpretation: In relation to MF-B1 to MF-B4, MF-B5 is characterised by a higher content of reef builders leading up to biostromal rud-boundstones, which were strongly influenced by episodic storms. Deposition took place on the drowned platform below the fair-weather wave base and within the mesophotic zone. The micrite was mostly not washed out by permanent Fig. 9 Microfacies types of the Eisborn Member of the Beul outcrop. a MF-B5c, organic-rich alveolitid-gastropod floatstone with Stromatoporella and commensalic Syringopora, laminar Alveolites (Alveolites) (top right), small gastropods, coral and shell debris in a bioturbated matrix with some micrite (Bed B11, thin-section I) b MF-B5d, alveolitid-stromatoporoid floatstone, contact of fragmented Thamnopora polyforata branches, Stachyodes (Keega) australe, and Alveolites sp. (lower part), surrounded by variably dense, organic-rich, mud-wackestone matrix with crinoid and shell debris (Bed B13) c MF-B5e, alveolitid-thamnoporid floatstone with Alveolites (Alveolites) sp. (upper part), Th. ex gr. polyforata (larger branches), and Th. ex gr. micropora (smaller branches) in a bioturbated, organic-rich mudwackestone matrix with partly washed out micrite (Bed B13e = Top 13) d MF-B5f, organic-rich rugose floatstone with Thamnophyllum, crinoid and shell debris, and ostracods in a bioturbated, fine bioclastic wacke-packstone matrix (Bed B7) e MF-B5g, rugose-thamnoporid floatstone with branches of Th. ex gr. polyforata and Disphyllum breviseptatum in micrite-rich, dense wackestone matrix with poorly preserved crinoid debris and several generations of calcite veins, including short sigmoidal cracks (lower left half) (Bed D) f MF-B5g, detail of a rugose-thamnoporid floatstone with a Th. ex gr. polyforata branch next to a spirally ribbed, pleurotomariid gastropod with geopetal sparite filling, surrounded by dense, organic-rich, micritic mudstone (Bed D, Sample B-D) g MF-B5h, alveolitid-rugose boundstone with Alveolites (Crassialveolites) sp., overlain by wave-laminated mudstone, embedding Disphyllum rugosum, and overlain by peloidal mud-grainstone (top biostrome, Sample top 1C) h MF-B5i, tabulate-rugose rudstone with alveolitids (lower right and upper left), fragmented Thamnopora sp., and Disphyllum rugosum in a bioturbated to brecciated, wacke-packstone matrix with fine debris (top biostrome, Sample top 1A) currents. Alveolitids and rugose corals are common, unlike as in typical back-reef facies of the biohermal Dorp Facies. The different subtypes suggest a small-scale lateral ecological differentiation, with patches dominated variably by thamnoporids, alveolitids, or dendroid rugose corals. The three organic-rich subtypes (MF-B5b, B5c, and B5f) reflect phases of increased primary productivity, obviously a favourable condition for gastropods, as it is known from gastropod-rich back-reef facies (e.g. Malmsheimer et al. 1991;Weller 1991: Type Ha5-2;Schudack 1993). The rud-boundstones indicate slightly shallower conditions with improved light supply for improved growth of frame builders. This is supported by the parallel decline of conodont faunas. Persisting storm influence prevented a preservation as insitu autobiostromes (Kershaw 1994). The growth of tabulate corals outpaced that of stromatoporoids, perhaps due to the recurrent influx of detritus as a limiting factor. It is not uncommon, that Alveolites overgrew other taxa or rather their fragments, such as laminar stromatoporoids or thamnoporid branches (Fig. 7h). Distribution: Givetian equivalents can be found within the initial, biostromal Schwelm Facies of Krebs (1974). Faber (1980)
Interpretation: As typical for (hemi-)pelagic environments, photic zone organisms, such as hermatypic corals, stromatoporoids, algae, and other calcimicrobes, are absent. Instead, there is a dominance of pelagic plankton (dacryoconarids, entomozoans, unilocular foraminifers) and nekton (cephalopods). The pure micrite accumulation suggests that the current energy was low, but high enough to prevent the settling of large amounts of clay, which deposited at the same time in the adjacent Flinz basins (Fig. 3). Bioturbation and abundant agglutinating foraminifers reveal fully oxic, good living conditions on and within the upper substrate, sufficient for burying endobenthonic organisms of unknown affinities. Episodic mass occurrences of dacryoconarids suggest eutrophic facies, with relative shell enrichments during reduced micrite supply and deep-water bottom turbulence. The classical microfacies model sensu Wilson (1975), which is based on the concept of Flügel (1972), was developed for Upper Triassic carbonate platforms. Therefore, we use the differentiated classification of Devonian (hemi-)pelagic to neritic carbonates of Hartenfels (2011). MF-C1 was deposited after the reef platform was transformed into a low-relief submarine rise. The calcisphere-rich variant MF-C1* indicates relative enrichments because of low sedimentation rates. Distribution: Similar dacryoconarid packstones are known from pelagic ramp settings of Morocco (e.g. Aboussalam 2003;Rytina et al. 2013;Ward et al. 2013;Becker and Aboussalam 2013a), the Montagne Noire, southern France (e.g. Aboussalam 2003), and from pelagic submarine rises of the eastern Rhenish Massif (e.g. Stritzke 1989Stritzke , 1990, his crinoid-tentaculite micrite facies; Aboussalam 2003).
A variant without goniatites and with only very rare, probably reworked homoctenids occurs in the basal Famennian of HON_1101 (Bed -8). It is characterised by abundant gastropods, ostracods, fragmented mollusk shells and a few, sparitefilled, small orthocones. Interpretation: MF-C2 represents an open marine, subphotic environment with abundant (hemi-)pelagic fauna (cephalopods, dacryoconarids, thin-shelled bivalves, conodonts, and entomozoans) deposited well below the storm wave base and under eutrophic conditions. Pyrite, which coats some bedding planes, and the alternating bioturbation suggest fluctuations between episodes of dysoxic and fully anoxic conditions. Some endofauna exploited the buried organic matter of underlying strata. The dominant shell fragmentation, lack of sorting by bottom currents, and original micrite matrix, normally evidence for calm deposition, are in contrast. Shell crushing occurred outside the area of deposition, with subsequent transport by occasional bottom currents. Frutexites-type structures have been regarded as typical for dysaerobic conditions (Préat et al. 2008) but Koptíková et al. (2010) and Hartenfels (2011) showed that they can also co-occur with a normal benthic fauna. Distribution: MF-C2 differs from typical Rhenish Upper Kellwasser facies (Schindler 1990;Gereke 2007) but shows some similarity with the more peloidal, shallower Kellwasser facies developed on top of the microbial Wülfrath Reef ).
The microfacies of the basal Upper Kellwasser level of HON_1101 (base of Bed -9b) is transitional towards MF-C2 and MF-C5. The marly level with goniatites and large ostracods, sandwiched between black shales in B102 within Bed -18 (Fig. 11e), is intermediate between MF-C3 and MF-C6. Interpretation: Recrystallization and the (hemi-)pelagic faunal assemblage, especially the entomozoans, cephalopods, and buchiolids, are typical for Kellwasser limestones of other sections (Schindler 1990). The local rarity of dacryoconarids in the Hönne Valley is distinctive. The high C org content, abundance of fossils, and some pyrite indicate eutrophic conditions but the absence of lamination/weak bioturbation suggests that the facies was dysoxic, not anoxic. This agrees with the reworking of micrite as peloids, a sign of weak bottom turbulence. Distribution: Gereke (2007) illustrated a Lower Kellwasser example with even higher fossil content from section Volkersbach in the southern Rhenish Massif ("Frankenbacher Schuppenzone"). There are also similarities with the very fossiliferous Kellwasser facies developed on the Tafilalt Platform of southern Morocco (Wendt and Belka 1991). Description: Macroscopically, this facies type occurs as light-to middle-grey, beige, reddish, or yellowish-brown nodular and flaserlimestone, partly with cephalopods (orthocones) and fragmented shells. There is no debris of reefal fauna (in contrast to bioclastic wackestones of MF-B3). The colour reflects fluctuating contents and oxygenation levels of very fine ironmanganese pigments that occasional led to the formation of dendrites (Beul: Sample top 3). The dense, bioturbated micrite may contain thin, fragmented shells, orthoconic cephalopods, goniatites (gephuroceratids), gastropods, smooth ostracods, rare entomozoans, calcispheres/unilocular foraminifers (very abundant in Beul, Sample top 3, including Parathurammina and Archaesphaera), trilobites, gastropods (HON_1101, top Bed -16; Beul, Sample top 2), rare, small-sized deep-water Rugosa (HON_1101, , and crinoid debris. Subordinate styliolinids and rare ribbed homoctenids occur only in Frasnian beds (e.g. HON_1101, Bed -14; B102, lower part Bed -18); these are transitional towards MF-C1. Especially larger shells show borings and a beginning micritization. Some thin-sections display Frutexites-type encrustations, thin silt layers (as a transition towards MF-C7), and mudclasts. The matrix is either micritic or microsparitic, with recrystallization spreading from bed boundaries or dissolution seams. Wide calcite veins were filled by blocky cement and were locally dolomitized. They are crossed by dark brownish stylolites, which may be associated with accumulations of pyrite grains, and younger, smaller calcite veins.
As an exception, the lower part of Bed -14 of HON_1101 is characterised by several irregular, dense accumulations of sponge spicule networks and calcispheres; the former are interpreted as remnants of deeper-water hexactinellid sponges. Interpretation: This facies type represents a quiet and deep (hemi-)pelagic, subphotic palaeoenvironment well below the storm wave base. This is supported by bedding-parallel geopetals. Frutexites-type encrustions indicate very low sedimentation rates . The presence of authigenic pyrite suggests local hypoxic conditions within the substrate, which, however, did not prevent the bioturbation. As normal in the Devonian, the burrowing organisms are not preserved; the very fine mud reflects an originally very soft substrate ("soupground", Etter 1995;Wetzel and Uchmann 1998). Mudclasts (e.g. Fig. 11b, lower right) can be formed by burrowing or by gravitational transport of weakly lithified lime mud on slopes. Distribution: Hartenfels (2011) described equivalents of MF-C4 from the Famennian of adjacent Rhenish localities (Effenberg Quarry and Oese) as well as from Thuringia and the Wildenfelser Zwischengebirge (Saxothuringia), the Holy Cross Mountains (Poland), and the Maider and Tafilalt of SE Morocco. It corresponds to the "low component micrite facies" of Stritzke (1989Stritzke ( , 1990: facies type II5), which was deposited on the submarine rise formed by the drowned Brilon Reef.
Description: MF-C5 is restricted to a thin, dark-grey layer of marly limestone at the base of the Upper Kellwasser level in core B102. Below, in the upper part of Bed -15 (Fig. 10d), there is a transition from MF-C4 by bioturbated ostracod wackestones. Ostracods include ribbed entomozoans as well as smooth forms and are preserved as single or complete shells, the latter partly with geopetal fillings. Additional fauna ist represented by subordinate bivalves and orthocones. Interpretation: An increased occurrence of ostracods instead of dacryoconarids or homoctenids and a higher amount of C org differentiates MF-C5 from the dacryoconarid wackepackstone of MF-C1. Ostracod blooms indicate eutrophication processes that favoured selective organism groups, which has previously been recognised within the Kellwasser Crisis Interval (Schindler 1990). But episodic primary condensation cannot be ruled out (Brett and Allison 1998). A slight increase in bottom currents may have reduced the clay and micrite accumulation (Aigner 1985;Schülke and Popp 2005;Hartenfels 2011), which is supported by partial shell disarticulation. Piper and Stow (1991) interpreted this kind of preservation as a result of distal turbidites or tempestites, respectively, but the pelagic setting was well below the storm wave base and, therefore, minor contourites intersecting the sea floor should be considered (see Hüneke et al., in prep.).
MF-C8: Peloidal and bioclastic packstone (Fig. 11b  (top part)) Examples: HON_1101, within Bed -13 (at 60.40 and ca. 60.48-60.49 m). Description: Thin intervals of abundant mud peloids with very fine skeletal detritus and some larger shells, such as smooth and ribbed bivalves. In the shown example (top Fig.  11b), the base is relatively sharp but upwards there is a gradation into bioclastic wackestone with trilobites and a juvenile, evolute goniatite with depressed cross-section (?Ponticeras), which conch is filled by fine micrite. Laterally, there is a 10 mm large mudclast that differs in its lithology from the rest of the unit. A second layer of MF-C8, again with a welldefined base, occurs at the top of Bed -13. Interpretation: The sharp base, peloid formation by mud reworking, and detrital content suggest short intervals of slightly increased bottom currents. The mudclast may have been moved at the peak of turbulence. Since the biofacies below and above, supported by the pelagic goniatite, indicates overall conditions well below the storm wave base, we suggest deposition by minor bottom currents. Conodont stratigraphy (Fig. 12) Drill core HON_1101 (Figs. 13, 14, 15, 16 and 17) The lagoonal lower ca. 50 m of the borehole (Fig. 13, Bed -39 to top of Bed -31) were barren (11 samples taken). Above a deep weathered marly interval (Bed -30), which probably represents a fault zone, a productive sample was taken from near the top of a ca. 2 m thick interval of flaserlimestone (top Bed -29, at 107.20-107.42 m, Fig. 13). It yielded a deeper-water assemblage with Pa. jamieae savagei M2 (Fig. 14b), Pa. ljaschenkoae M2 (Fig.  14a), Pa. hassi, Ag. cf. triangularis (Fig. 14c), Ad. nodosa (Fig.  14d), Po. paradecorosus, I. symmetricus (Fig. 14e), and "Oz." aff. nonaginta (Fig. 14f). Palmatolepis jamieae savagei M2 enters in the Pa. feisti Zone/Subzone (MN Subzone 11a) of Martenberg (Saupe and Becker 2022, this issue.), where Ag. triangularis becomes common. Typical "Oz." nonaginta are not known from above the Pa. housei Zone (MN Zone 8, see Klapper et al. 1996).
The fault interval may also encompass Bed -28, which was not sampled. Above follow more than 20 m of typical reef limestones (Fig. 13, beds -27 (Fig. 14g) and Ad. rotundiloba fragments; the latter are a clear indicator for the basal Frasnian (MN zones 1 or 2). Three samples from 78.00-78.15 m) contained each different species (Fig. 13) that are all typical for lower Frasnian shallow-water facies. Icriodus subterminus, found in Sample -20c, does not range above the top of the lower Frasnian (Narkiewicz and Bultynck 2010).
From Bed -18 on, conodont faunas become diverse, belong to the pelagic Palmatolepis biofacies, and are partly so rich that we concentrate our report on the successive entry of biostratigraphic markers and unusual forms. At the base of Bed -18 (77.56-77.71 m), poorly preserved Ad. nodosa (Fig. 14h) and narrow relatives of Pa. transitans, possibly Pa. keyserlingi, appear. The first indicates the Ad. nodosa Zone of Becker and Aboussalam (in Pisarzowska et al. 2020), which base correlates with the top of the Pa. transitans Zone (top MN Zone 4). Palmatolepis keyserlingi was described by Kuz´min (1998) from the Lower Member of the Domanik Formation in the Timan, which falls in the Pa. punctata punctata Zone (MN Zone 5, House et al. 2000; regional Po. efimovae-Pa. punctata Zone of Ovnatanova and Kononova 2008). The next sample from the upper part of Bed -18 yielded Pa. transitans, juvenile Ad. nodosa that could be mistaken as representatives of older ancyrodellids (Fig. 14i), and both Pa. punctata punctata (Fig. 14j) and Pa. punctata bohemica (Fig. 14k). The latter is an alternative marker for the Ag. primus Zone (MN Zone 6; e.g. Klapper et al. 1996;Klapper 1997). Associated are a possibly new species of Palmatolepis (Fig. 14l) and a distinctive variant of Z. ovalis (Fig. 14m).
Sample -12a is poor in Palmatolepis but rich in icriodids (sudden surge of I. symmetricus, I. praealternatus praealternatus, I. vitabilis, and I. alternatus alternatus), and polygnathids (sudden late appearances of juvenile Po. capollocki and Po. zinaidae), an ecostratigraphic pattern first noted in Lower Kellwasser beds of the Frasnian/Famennian boundary GSSP section at Coumiac (Klapper et al. 1993). We did not find the Lower Kellwasser level index species Ag. asymmetricus, but a late variant of Ad. hamata in which the secondary carinas of the side lobes rise and develop short free blades (Fig. 15k). The same feature is long-known from late forms of Ad. curvata (see Fig.  15l). In the upper part of Bed -12 (Sample -12b, at 59.33-59.42 m), there are abundant Pa. bogartensis (Morphotype B, Fig.  15m-n), the index species of the Pa. bogartensis Zone (MN Zone 13a). The thin interval of Bed -11 (with Sample -11a from the base and Sample -11b from 59.13-59.22 m) to Bed -10 represents the middle and upper parts of the Pa. bogartensis Zone (MN Zone 13a). Sample -9a (at 58.97-59.08 m) has a sparse fauna. Apart from several forms that are difficult to assign to one of the common terminal Frasnian species, we recognised Pa. boogardi and Pa. beckeri in a very rich assemblage of Sample -9b (at 58.80-58.94 m). Palmatolepis beckeri enters very late in the Pa. bogartensis Zone (top of MN Zone 13a) in its type section of Australia (Klapper 2007), just below the regional Upper Kellwasser equivalent (Manticoceras guppyi Bed, Becker et al. 1991). Sample -9c (at 58.68-58.80 m) yielded Pa. beckeri variants that are somewhat homeomorphic to the older Pa. semichatovae. Palmatolepis linguiformis, the index species of the top-Frasnian Pa. linguiformis Zone (= MN Zone 13b/c), was not found. It is long-known that it was rather facies sensitive and absent or very rare in several regions (Girard et al. 2005).
Subsequently, two samples from the lower, ca. 5.5 m thick red marls (Bed -1, Hemberg Formation) were not productive. The upper ca. 43 m of the borehole consist of basinal marls and shales that are not calcareous enough for conodont sampling.
Drill core B102 (Figs. 18, 19, and 20) The lagoonal lower ca. 87 m of core B102 have not been sampled for conodonts (Fig. 18). A sample from the debris interval at the top of the Hagen-Balve Formation, the local Eisborn Member interval (upper part of Bed -27), was barren and beds -26 and -25 could not be sampled.
Accordingly, the small amounts of dissolved core material of most B102 samples from the Beul Formation yielded only few conodonts. At the base, Sample -24f produced an assemblage with a fragmentary Ag. amplicavus (Fig. 19a), a specimen identified preliminarily as Pa. cf. housei (Fig. 19b), Po. paradecorosus (Fig. 19c), and Ad. lobata. The first enters at the base of the "Oz." nonaginta Zone (MN Zone 7, Klapper et al. 1996) while typical Pa. housei define MN Zone 8. Faunas representing the lower half of the middle Frasnian are missing. In Sample -24e, Ad. lobata (Fig. 19d) is associated with a Pa. cf. plana that appears to be transitional from Pa. housei (Fig. 19f). This gives some uncertainty whether the Pa. plana Zone (MN Zone 10) level has been reached. In Sample -24d, Pa. hassi with a short side-lobe (Fig. 19e) enters, associated with Ad. gigas. The middle/upper part of Bed -24 is conodont-poor. Sample -24c yielded another Ad. gigas (Fig. 19g), Sample -24b an incomplete specimen with short side lobe identified as Ag. aff. barbus; typical Ag. barbus are not known from above the Pa. housei Zone (MN Zone 8; Klapper 1997).
All of the subsequent Famennian to upper Tournaisian is missing in a long gap below cherts of the Hardt Formation (beds -1 and 0).
Beul outcrop (Figs. 21, 22, and 23) The preliminary outcrop conodont biostratigraphy was outlined in Becker et al. (2016c) but further sampling requires updates, especially a shift of the position of the Givetian/Frasnian boundary. Samples from the Eisborn Member mostly have a very low conodont yield, often less than 5 Pa elements (mostly polygnathids) per kg of limestone. A few richer samples (ca. 15-20 Pa elements/kg) indicate deepening intervals. The nodular limestones of the Beul Formation are much richer, partly with several hundred of Pa elements in a normal-sized (2-3 kg) sample, but many specimens are juveniles. The complete local species ranges are given along the section log in Fig. 21. Since published conodont data are very scarce for Frasnian Rhenish reef facies and directly overlying strata, abundances are compiled in Tables 2 and 3. Four samples from the thick lowest bed (Bed B1) yielded almost exclusively small numbers of Po. alatus, which normally does not occur before the topmost Givetian Sk. norrisi Zone (Aboussalam and Becker 2007). The associated single Po. paradecorosus represents a species that enters at the same level. Loose slabs with brachiopods, laminar stromatoporoids, and rare trilobites yielded the same almost monotypic fauna. A slight increase of polygnathid diversity in beds B3 to B5 is caused by local late entries of long-ranging middle/upper Givetian to lower Frasnian species, such as Po. varcus and Po. dubius, in association with Po. alatus (Fig. 22c). Two Ad. rotundiloba pristina date Bed B3 as basal Frasnian Ad. rotundiloba pristina Zone (MN Zone 1). Previously, the first record was from Bed B11 (Becker et al. 2016c), resulting in a significant downwards shift of the local Upper Devonian base. Since the larger specimen ( Fig. 22de) displays an advanced morphology (high number of platform nodes combined with a very large basal cavity), the Frasnian base may lie even slightly lower. Beds B6 to B10b have mostly the same poor assemblages as at the section base, apart from curved variants of Po. dengleri sagitta. The subspecies enters in southern Morocco in the upper Givetian but ranges into the basal Frasnian (Aboussalam and Becker 2007). A more advanced ancyrodellid with 28 platform nodes, identified as Ad. rotundiloba soluta, enters in Bed B10c (Fig. 22f-g) and indicates the base of the Pa. rotundiloba rotundiloba Zone (Pa. rotundiloba soluta Subzone, MN Zone 2a). It is associated with Po. pollocki, which is a typical lower Frasnian species. Entries of Po. pennatus and Po. webbi in Bed B11 represent further facies-controlled delayed FODs. A single Zieglerina from Bed B11 (Fig. 22h-i) represents a new species (Zieglerina n. sp. B).
Icriodids, which normally characterise shallow-water limestones, appear suddenly in Bed B13a. The locally oldest I. subterminus and I. symmetricus represent delayed entries (see icriodid zonation of Narkiewicz and Bultynck 2010). Subsequent beds were difficult to sample in the steep cliff; the small-sized samples retreived proved to be conodont-poor. In thin-bedded limestones at the top (beds E1 and E3), faunas become richer. Specimens of Po. dengleri dengleri from Bed E1 (Fig. 22j) with an incipient anterior platform collar are transitional towards Po. cf. dengleri sensu Aboussalam et al. (2020, fig. 20.10) that occurs in the top-lower Frasnian of Morocco. There is the late record of an ancestral looking anyrodellid (Fig. 22k) that is morphologically intermediate between Ad. rotundiloba pristina and Ad. binodosa. Some Po. dubius (Fig. 22o), Po. paradecorosus (Fig. 22p) subperlobata with some nodes on the inner anterior platform, transitional from Pa. ultima (see Klapper et al. 2004, p. 382), top Bed -8, x 65 this issue), and I. subterminus (Fig. 22m) are associated with a single, last I. difficilis (Fig. 22n). This species is normally restricted to the Givetian (e.g. Narkiewicz and Bultynck, 2010, fig. 10) but Bultynck (2003, fig. 2) noted a questionable lower Frasnian range in North America.
Several Ad. recta (Fig. 22r) date Bed E3 as upper part of the Ad. rotundiloba rotundiloba Zone (MN Subzone 2b, Table 3; see lower range of Ad. recta in Kralick 1994). Associated are six species of Polygnathus, including a re-appearance of Po. webbi (Fig. 22q). The final, poorly exposed coral biostrome yielded hardly any conodonts during several initial sampling trials, a typical feature of reefal facies. A single Mes. guanwushanensis (Fig. 22s) represents a new morphotype, M3 (see taxonomic notes). In southern Morocco, the same form reaches the basal middle Frasnian Pa. punctata punctata Zone (MN Zone 5, Becker and Aboussalam 2013b) but it is more common in the lower Frasnian. A later sample from coral floatstone (Sample "top reef") turned out to be relatively rich in Pa. paradecorosus, including specimens with rather long free blade (compare Po. aff. paradecorosus of Aboussalam et al. 2020). Associated are a single I. cedarensis, a longranging species (Narkiewicz and Bultynck 2010), and Po. dengleri dengleri. The latter taxon is not known from above the top of the lower Frasnian (top of Pa. transitans Zone, see composite range in Klapper 1997;Gouwy et al. 2007). This constrains the youngest age.
Apart from typical I. symmetricus, there is a curved related form, named as I. cf. tafilaltensis (Fig. 23b), and a polygnathid with flat, weakly nodose platform, a rare morphology in middle Frasnian strata (Polygnathus sp., Fig. 23c). Although only two species of Palmatolepis are identified, there are three types of Nothognathella ("sp. 1-3", Fig. 23d-f), the supposed Pb element of the genus. A distinctive feature of the fauna is the relative abundance of a form resembling the mostly Givetian Po. varcus, preliminary identified as Po. aff. varcus (Fig. 23g).

Other stratigraphic markers of the Beul Formation
Shark teeth of the basal Beul Formation of the outcrop belong to Phoebodus latus, which is known from the higher middle Frasnian of the Urals, and Ph. fastigatus (Fig. 22a-b), which ranges from the Givetian to the upper Frasnian (Ginter and Ivanov 2000). Both species have previously not been reported from the Rhenish Massif (see recent review by Ivanov 2021).
The presence of mesobeloceratids in the oldest Beul Formation of the outcrop, typical for UD I-G 2 , agrees with the middle Frasnian conodont age (Becker and House 2009). The slightly younger Beloceras is long-known from slightly higher cliffs (e.g. Becker et al. 2016c). Beloceratidae are usefor for water depth calculations (Hewitt 1996).
Based on additional thin-sections, some previous identifications of supposed solitary Rugosa (Temnophyllum, Chostophyllum) from the Eisborn Member of the Beul outcrop (Becker et al. 2016c) have to be revised. These represent branches of the dendroid genus Disphyllum. Disphyllum breviseptatum (Frech, 1886) is known from the lower Frasnian of Refrath in the western Rhenish Massif, while Dis. rugosum (Wedekind, 1922) Fig. 24b), which is a Frasnian genus. Due to the strictly straight polypar walls and strong carinae, the sectioned hexagonariid colony is not Hexagonaria hexagona, but a relative of Hex. davidsoni (Milne-Edwards and Haime, 1851), identified here as Hex. aff. davidsoni (Fig. 24a). The typical form is cosmopolitan in the Frasnian (compare description by Falahatgar et al. 2018).
Based on the new material, the Stachyodes (Keega) specimens from Beul are now assigned to St. (K.) australe Wray, 1967, not to St. (K.) jonelrayi. The first is known globally, including the eastern Sauerland (May 1993b). Other stromatoporoids are rare and not well preserved; they are assigned to ?Euryamphipora, ?Clathrocoilona, and Stromatoporella.
A rare new odontopleurid trilobite is described in a separate note (Helling and Becker 2022, this issue). It was found on a slab with Po. alatus, St. (Keega) australe, and Th. ex gr. polyforata.

Facies developments
Drill core HON_1101 (Figs. 4,13,16) Asbeck Member: In drill core HON_1101, the upper ca. 80 m of the Asbeck Member (excluding a ca. 8 m interval of intercalated post-reefal strata) were penetrated between ca. 78 and 165 m. In its lower ca. 7 m (beds -39 to -36, 158.15-165.00 m), the palaeoenvironment is characterised by shallow lagoonal conditions (MF-A1, micritic, dendroid stromatoporoid float-rudstone, Fig. 6b). Amphipora and Stachyodes (represented by St. (St.) costulata) are variably dominant and associated with bulbous stromatoporoids. This suggests the lateral growth of delicate Amphipora and more robust Stachyodes thickets depending on the level of water agitation. The setting was relative stable for some time, presumably caused by a carbonate production that kept up with subsidence. Amphipora (MF-A3, represented by Am. ex gr. laxeperforata) became dominant in Bed -35, indicating calm episodes. In the more than 4 m thick Bed -34 (Fig. 6f), the change to the high-energy conditions of MF-A2 (detrital stromatoporoid grain-rudstone, including Parallelopora sp.) reflects storm events or the lateral shift of a channel adjacent to a patch reef.  Fig. 7c), the sporadic appearance of MF-A4 (fenestral pack-grainbindstones) indicates a shift from subtidal to intertidal conditions and, therefore, a minor sea level fall. As conodonts avoided back reefs and lagoons, it was not possible to date this phase.
Between 113.00 and 105.50 m (Fig. 13), the carbonate succession is interrupted by strongly weathered clayey sediment, which probably belongs to a fault zone. An intercalated, ca. three-metres thick limestone unit (Bed -29) is in bioturbated and bioclastic wackestone facies (MF-C2) revealing open shelf conditions, which is supported by pelagic palmatolepid biofacies. The dating as upper Frasnian proves that it is an intercalation of Beul Formation that does not belong to the reefal succession.
The "normal" reef development continues at 105.50 m (Bed -27) with lagoonal stromatoporoid float-rudstones (MF-A1, Fig. 6d, with Actinostroma clathratum) of the Asbeck Member. As typical for back-reef settings, there is no conodont record. At the top of Bed -25 (102.80-103.00 m), MF-A1 yielded, as below, the dendroid St. (St.) costulata (Fig. 6a). A short sea level fall is noticeable at 102.00 m by MF-A4b, indicating an intertidal to supratidal environment. At 94.98 m, peloidal stromatoporoid rudstones (MF-A5, Fig. 7d) occur, which yielded dendroid St. (St.) costulata and Thamnopora sp., but also Alveolites (Crassialveolites) sp. This was only a brief episode, directly followed by a return to the lagoonal MF-A4a (fenestral pack-grain-bindstones, Fig. 7a). The close association records either an influx of outside detritus by a major storm or a short transgressive pulse shifting reef facies onto the lagoon. The last MF-A4a is found at 90.00 m.
The final phase of the reef complex (Asbeck Member) is marked in core HON_1101 by a package (ca. 83.40-90.00 m, Bed -22) of MF-A5. Large stromatoporoids, including Schistodictyon sp., Salairella buecheliensis, Actinostroma ex gr. filitextum, and others dominate rudstones with some Stachyodes, Th. ex gr. micropora (as in the Eisborn Member, see below), subordinate solitary rugose corals, and a peloidal grainstone matrix that includes fragmented crinoids. This is clearly a reef core to outer margin facies.
Eisborn Member: Deepening is supported by the incoming of open shelf organisms (basal Frasnian conodonts) in Bed -21 of the basal Eisborn Member. The sudden facies shift above the thick lagoonal sequence reflects a significant reef margin backstepping, which correlates in time with the eustatic floodings of the global Frasnes Events (e.g. House 1985;Ebert 1993;Aboussalam andBecker 2007, 2017;= Mesotaxis Event of Racki 1993). Its severe effect on reefal platforms and its varied biota is long known from the Ardennes, where the thick Fromelennes Reef Complex was flooded at the base of the Frasnian (e.g. Bigey et al. 1982;Devleeschouwer et al. 2010). Far away, in the eastern Taurides of southern Turkey, biostromes were submerged in a similar way by two top-Givetian and lower Frasnian sea level rises that correlate with the Frasnes Events (Özkan et al. 2019). This example underlines the eustatic nature of sea level changes.
In Bed -20, alveolitid-dominated rudstones (Fig. 7h, 79.12-79.32 m) with Alv. (Alv.) ex gr. edwardsi, some Th. ex gr. polyforata, subordinate rugose corals, a small amount of crinoid debris, and rare polygnathids follow. The dominant laminar growth of tabulates and the presence of crinoid debris reveal the transformation into a drowned biostromal platform. The deepening also caused the replacement of most stromatoporoids by corals. The microfacies is assigned to MF-B5h/i of the Eisborn Member, correlating laterally with the top lower Frasnian biostrome of the Beul outcrop (see below).
The drowning of the reef platform involved an incredible decrease (up to 120 times) of carbonate production. The up to 1.000 m thick top-lower to middle Givetian strata of the main Asbeck Member, a time span in the order of 3 Ma , gives an accumulation of ca. 333 m/Ma. This contrast with 6-7 m for all of the lower Frasnian, lasting ca. 2.5 Ma, giving ca. 2.8 m/Ma. This trend is accompanied by the unique pack-rudstones with the auloporid Roemerolites sp. Fig. 7f,. The I. subterminus fauna from the top shows that the reef remained in the open shallow-water realm (see Narkiewicz and Bultynck 2010).
Frasnian Beul Formation: Starting with the thin Bed -19, no reefal fauna is noticeable anymore. The rapid change to  Pisarzowska et al. (2020). In many regions, but not everywhere, the Middlesex Event is characterised by the sudden spread of hypoxia (black shales). In the distal slope setting south of the Brilon Reef, there is geochemical evidence both for enhanced primary production and oxygen deficiency. At the Beul, the event caused extreme condensation, probably with unconformities due to non-deposition, but no increased deposition of C org , and the facies remained oxic. The Pa. punctata punctata and Ag. primus zones (MN zones 5/6, lasting up to 1 Ma; Becker et al. 2020) are represented by just ca. 70 cm of beds -18 and -17. The latter, a solid limestone layer at 77.00 m, is a bioclastic, styliolinedominated, pelagic wacke-packstone (MF-C1, Fig. 10e).
The anoxic sequence ended as sharp as it began. Bed -14, which falls in the higher part of the Pa. feisti Zone (MN Zone 11), consists again of well-oxygenated, bioturbated flaserlimestone. The occurrence of several sponge bodies, indica-ted by spicule arrangements, is unique for the borehole. Within the following Bed -13, there are thin, strongly bioturbated layers of silt (MF-C7) and peloidal packstones (MF-C8) that indicate minor phases of increased turbulence (Fig. 11b). This is the only and rather weak local evidence for equivalents of the regressive Usseln Limestone that characterises just below the Lower Kellwasser interval basinal and submarine rise sections of the eastern and southern Rhenish Massif, Harz Mountains, and Saxothuringian Zone (Gereke 2007;Gereke and Schindler 2012;Gereke et al. 2014). The eutrophic and transgressive subsequent Lower Kellwasser Event is represented by the almost 60 cm thick, darkgrey, organic-rich, recrystallized ostracod-mollusk packgrainstones of Bed -12 (MF-C3, Fig. 10h). As elsewhere, the unit spans the boundary between the Pa. winchelli (MN Zone 12) and Pa. bogartensis (MN Zone 13a) zones. It is remarkable that there are still only a few published illustrations of Rhenish Lower Kellwasser facies types from submarine rises. In the Hönne Valley area, eutrophic conditions did not lead to sea floor anoxia. Ostracods and mollusks benefitted locally much more strongly than dacryoconarids or homoctenids. In the well-studied Schmidt Quarry (Kellerwald, LKW thickness = 50-55 cm), Benner Quarry at Bicken (Dill Syncline, LKW thickness = 41 cm), and in the Kellwasser type-section of the Harz Mountains (LKW thickness = 37 cm), the typical Lower Kellwasser limestones are rich in homoctenids and mostly laminated (Schindler 1990), representing deeper submarine rise facies. The intra-Kellwasser succession is a thin interval (Bed -11 to lower part of Bed -9) characterised by the return to bioturbated, oxygenated, micritic mud-wackestone deposition. At the base of Bed -11, recrystallization and strong fracturing mask the facies transition. However, there is at least one reworked Lower Kellwasser clast indicating a disconformable contact. Above a thin interval with calcisphere wackestones (MF-C1*), light-grey, bioturbated mudstones (MF-C4) prove the return to a quiet and oligotrophic deep-water setting. The main part (level of Sample -9a, 58.97-59.08 m) consists of three layers that are each more bioturbated and nodular at their bases. The recrystallised (microsparitic) mudstone contains some calcispheres, shell debris, rare ostracods and dacryoconarids, as well as the single branch of a bryozoan or algal thallus.
The 14 cm thick level of Sample -9b (58.80-58.93 m) is characterised by the return to (moderately) organic-rich, middle-to dark-grey limestone, interpreted as a more shallowwater, better oxygenated equivalent of the Upper Kellwasser Limestone (MF-C2, Fig. 10b-c). The base is relatively sharp, with the top of the underlying subunit (Sample -9a, 58.93-59.94  Fig. 16) developed as a light-grey, bioturbated mudwackestone (MF-C4) with shell debris, some ostracods, and crinoid ossicles. Right above, at 58.91-58.93 m (base of Fig. 10b), eutrophication caused an ostracod and homoctenid bloom; the microfacies is transitional between MF-C2, C3, and C5. The ostracod bloom occurs also in core B102 (see below). However, the typical Upper Kellwasser ecological succession established for Rhenish basinal and other submarine rise settings by Schindler (1990Schindler ( , 1993 and Gereke and Schindler (2012) is not developed.
Typical for Kellwasser Facies is the rather strong recrystallization. The subsequent middle grey, partly laminated, partly bioturbated mollusk wacke-packstones (MF-C2) are interrupted by two thin layers of light-grey, bioturbated mud-wackestone (MF-C4, Fig. 10b). From these, burrowing organisms dug downwards into the more organic-rich layers. The polyphase deposition of the local Upper Kellwasser unit is underlined by vertical fluctuations between wacke-and packstones. At least ten successive depositional intervals can be recognised, which equals the number of subunits at Steinbruch Schmidt (Schindler 1990(Schindler , 1993. A small-sized, incomplete, pachyconic goniatite cross-section resembles the shell form of early Crickites holzapfeli. Interesting is the abundance of embryonic to early juvenile goniatite shells. These suggest recurrent worsening of environmental conditions, e.g. by changing oxygenation, which killed episodically juvenile goniatite populations before they could grow. The strong fragmentation of most shells indicates allochthonous mollusk debris, with water agitation in the source region and episodic transport by bottom currents, however, without winnowing the micrite. There are clear differences to the typical Upper Kellwasser beds of other Rhenish sections deposited in the shelf basin or on isolated submarine rises (e.g . Schindler 1990;Gereke 2007;Gereke and Schindler 2012). Becker et al. (2016b, fig. 21) illustrated a shallow-water variant, but still within the Palmatolepis Biofacies realm, from the Upper Kellwasser unit that drowned the microbial top part of the Wülfrath Reef. It differs in its more peloidal matrix, resembling the Lower Kellwasser Limestone of HON_1101.
Sample -9c (58.68-58.80 m, Fig. 16) appears to be lightergrey and less organic-rich than Sample -9b but this is mostly a consequence of its much stronger recrystallization, including wide, sparite-filled cracks, iron mineralizations, and beginning dolomitization. The less affected parts of the thinsection suggest the same MF-C2 microfacies as below, with some bioturbation and micritization of larger mollusks shells. This interpretation is supported by partial goniatite cross-sections, which probably belonged to Manticoceras, Crickites, and "Archoceras", a typical Kellwasser assemblage, as well as by the conodont fauna.
Our data prove that both Kellwasser Events extended in the Rhenish Massif as blooms of specific organism groups and increased burial of C org to shallower settings than described in most previous studies. The microfacies evidence does not answer the unsolved questions why groups adapted to Kellwasser conditions (e.g. conodonts, ammonoids, specific ostracods, and bivalves) died out at the top of the crisis interval. For reviews of the many Kellwasser hypotheses and models see Racki (2005), Carmichael et al. (2019), and Becker (in press).
Famennian Beul Formation: The basal Famennian Bed -8 is a ca. 1 m thick, poorly preserved, and deeply weathered unit consisting of heavily fractured to brecciated, middle-grey, marly limestone and flaserlimestone. It is impossible to decide whether the Frasnian-Famennian (F-F) boundary brecciation was a synsedimentary or entirely post-sedimentary, diagenetic feature. However, there is mounting evidence for seismic events right at the stage boundary (e.g. Racki 1999;Becker et al. 2016bBecker et al. , 2018Szrek and Salwa 2020). The poorly preserved interval lies at ca. 59 m depth but deep carstification is common in the Hönne Valley area and led to the formation of its famous caves. The brecciated interval may have been a path for circulating ground water. The microfacies (MF-C2) in the upper part of Bed -8 (58.40-58.50 m) still resembles the top-Frasnian interval but the goniatites, buchiolids, and homoctenids have disappeared. While basal Famennian blooms of ostracod populations are known from other sections (Casier and Devleeschouwer 1995;Becker et al., 2016e), the proliferation of gastropods is a rather unique facies pattern in the basal Famennian.
The following 14 m of the upper part of the drill core consist of nodular limestone and flaserlimestone, rather monotonous MF-C4 (mud-wackestones). The fauna contains ostracods, rare cephalopods, broken mollusk debris, associated with authigenic pyrite, and common traces of bioturbation. The setting was a stable to uniform, (hemi)pelagic, subphotic palaeoenvironment with oxic to dysoxic conditions. Beds -6 and -7 had a richer macrofauna of cephalopods and some trilobite debris. The recognition of Cheiloceras cross-sections at 55.85-56.03 m (Fig. 10d) is important since it confirms the onset of lower Famennian ammonoid radiation in upper parts of the Pa. termini Zone (see Becker 1993, p. 157). The Hönne Valley setting was too distal to clastic shelf regions and too deep to get affected by the two regressive Condroz Events at the top of the lower Famennian (Becker 1993;Becker et al. 2020) that caused the basinward progradation of a sandstone wedge in the west ("Plattensandstein", Fig. 3; Krebs 1979). Based on the conodont stratigraphy, the Condroz Events should occur within Bed -2 (within the Pa. rhomboidea and Pa. gracilis gracilis zones). The change from condensed nodular limestone to marl at the base of Bed -1 suggests a deepening trend in the middle of the middle Famennian (higher Pa. marginifera utahensis Zone) but there is no correlation with the refined eustatic curve . The typical red nodular limestones of the Hemberg Formation are locally not developed.
In the interval from 53.58 to 43.79 m, there are three, variably thick units of macroscopic reefal breccia (Bed -44, 337 cm thick, Bed -42m, 105 cm thick, and Bed -39, 73 cm thick). These can be explained by two possibilities: reef backstepping during sea level rise, resulting in the shedding of reef debris from the retrograding reef core onto the previous lagoon strata, or by shallowing in the lagoon, resulting in the formation of patch reefs that were strongly affected by wave action and recurrent storms. Since the intervening beds and the overlying  are in lagoonal facies, the second interpretation seems more likely. Especially intriguing is the intercalation within Bed -43, at ca. 49 m, of fenestral fabrics (MF-A4) indicating a very shallow back-reef setting and lowstand peak. The number of stromatoporoids decreased but Amphipora bafflestones still occur.
From Bed -31 to Bed -30, there is a gradual increase and rise to dominance of laminar stromatoporoids, which we interprete as the beginning of the terminal reef drowning by backstepping. However, in the middle of Bed -30 (at 30.97-31.12 m), there is a last episode of fenestral and peloidal lagoonal facies (MF-A4), grading upwards (30.85-30.97 m) into fenestral stromatoporoid floatstones with peloidal grainstone matrix (intermediate towards MF-A2). The peloidal stromatoporoid rudstone with crinoid debris of Bed -28 (23.07-23.25 and 24.20-24.38 m), typical MF-A5, shows that a reef core/margin setting was reached, which lasted to the top of the bed, the top of the Asbeck Member.
Beul Formation: At a depth of ca. 18 m, the Beul Formation begins after a phase of extreme condensation or non-deposition, spanning at least the lower part of the middle Frasnian, possibly also parts of the lower Frasnian. The contact to the Eisborn member was obscured by diagenetic overprint (brecciation and flaser-bedding). A significant deepening is marked at the base of Bed -24 by the onset of bioturbated dacryoconarid wackestones (MF-C1, Fig. 10f-g), followed higher by mud-wackestones (MF-C4, Fig. 11c-d). In thin-sections, the fauna consists of typical pelagic organisms, such as cephalopods (orthocones and goniatites), entomozoans, smooth-shelled ostracods, dacryoconarids, rare trilobites, thin, broken bivalve shells, deformed crinoids, and gastropods. Authigenic pyrite and Frutexites-type microbial encrustations suggest dysoxic conditions within the sediment, similar to HON_1101. At the top of the middle Frasnian, in the lower part of Bed -18, bioturbated wackestones include as faunal newcomers various, partly juvenile goniatites and homoctenids. This facies type is sharply interrupted at 7.14-7.24 m by a package of flaser-laminated, organic-rich black shales with a few ostracods (MF-C6; Fig.  11e). The conodont record is not precise (no record of the Pa. feisti Zone = MN Zone 11 below), but by comparison with core HON_1101, we assume that it represents the semichatovae Event Interval. Because it predates the Pa. winchelli Zone (MN Zone 12), it cannot be the Lower Kellwasser level. The shales grade into a non-laminated layer of argillaceous, peloidal, and recrystallized wackestone with partly unusually large ostracods and juvenile to fully grown goniatites (probably Manticoceras), intermediate between MF-C6 and MF-C3 (Fig.  12e). This suggests first anoxic, then dysoxic conditions. Above a thin, marly unit (Bed -17), no unequivocal equivalent of the Lower Kellwasser Limestone can be identified but the core recovery was incomplete (Fig. 18). Within , there are several layers of bioclastic wackestone with bivalves, ostracods, juvenile goniatites, and rare homoctenids, separated by pyrite-enriched, horizontal dissolution seams, which suggest episodic dysoxia. In the higher part, at 5.89 m, and as a filling of a larger orthocone near the top, there are patches of fossiliferous wackestones that are slightly darker and richer in C org and, therefore, transitional towards MF-C2. But the difference to the rest of the bed, also in comparison with HON_1101, is too weak to draw conclusions. The overlying, bioturbated Bed -15 is characterised by a gradual bloom of ostracods (Fig. 10d) culminating in a thin packstone layer (MF-C5) at the top. The conodont residue contained originally pyritic juvenile bivalves and longiorthoconic cephalopods. This indicates hypoxic conditions and a transition towards the partly laminated black, organicrich, marly limestone (Bed -14) that forms the top of the Frasnian and represents the Upper Kellwasser level Fig. 11f). Fully anoxic facies with rather restricted fauna is typical for basinal Upper Kellwasser settings (e.g. Gereke 2007). The lack of lamination in the upper, unfossiliferous part is caused by recrystallization (micritization), not by bioturbation. The absence of ammonoids indicates hostile conditions both at the seafloor and within the water column.
The overlying flaserlimestone (Bed -13, 5.05-5.28 m) yielded a rich basal Famennian conodont assemblage, which confirms that Bed -14 is the Upper Kellwasser level. The poor core preservation prevents to document the contact at the stage boundary. The lower Famennian nodular to flaser-bedded limestones are bioclastic wackestones (MF-C4) that indicate a return to oxic pelagic facies. The local marl-limestone alternations suggest a slightly deeper setting than for HON_1101. The sedimentary and faunal record was cut off still low in the lower Famennian (near the top of the Pa. minuta minuta Zone), probably due to non-deposition since there is no evidence for faulting below the overlying thin Viséan strata (Fig. 5).
Beul section (Fig. 21) Eisborn Member: At its base (beds B1-4, Fig. 21), 2.8 m of thick bedded and medium grey limestone consists of peloidal grainstones (MF-B2) grading upwards into peloidal, bioclastic wackestones (MF-B1). Tabulate corals dominate the dispersed frame builders but additionally there are rare colonial rugose corals (phaceloid Disphyllum), accessory stromatoporoids, crinoid and mollusk debris, as well as ostracods. This detrital variant of the Eisborn Member characterises a drowning of the platform, which caused a significantly reduced carbonate production and flooding of the protecting reef margins. This enabled the subordinate influx of open water organisms, such as conodonts, brachiopods (e.g. the spiriferid Adolfia, Fig. 24c-d), and trilobites (Gondwanaspis, Helling and Becker 2022, this issue). Moderate currents clumped fine micrite to "pseudopellets" via hydraulic forcing at the seafloor (Fåhraeus et al. 1974). The sparse polygnathid fauna supports a correlation of the bioherm drowning with the first pulse of the Frasnes Events in the Sk. norrisi Zone (uppermost Givetian; Becker and Aboussalam 2004). Ebert (1993) coined the term Ense Event for this topmost Givetian eustatic rise.
At the base of Bed B3 (Frasnian base), the peloids become rarer (MF-B3), due to decreased bottom currents and micrite reworking. In the higher part of Bed B4, thamnoporids become abundant in floatstones (MF-B5a) that also include three genera of stromatoporoids, reflecting an episodic "coral-sponge garden" setting disturbed by distal storms. The next lithological change is marked at the base of Bed B5. From there to Bed B11, the percentage of organic content is increased in more thin-bedded limestones, mostly dark, bioclastic mud-wackestones (MF-B4). This suggests a higher rate of C org burial but there is no corresponding rise in carbon isotope values as evidence for high primary productivity (Becker et al. 2016c). Very typical, despite their small numbers, are small-sized, planispiral, multi-chambered, calcareous foraminifers (Nanicella), which had a pantropical distribution in eutrophic reefal facies with moderate water agitation. Solitary, hermatypic Rugosa (Thamnophyllum, Becker et al. 2016c, fig. 13) occur in Bed B7. The gradation of MF-B4 into organic-rich and micritic floatstones with alveolitids and gastropods (Bed 8, MF-B5c) or thamnoporids and stromatoporoids (Bed B9, MF-B5b) confirms that the drowned lagoon stayed in the lower part of the euphotic zone. The increase of conodont abundance in beds 6-7 (Tables 2 and 3, middle part of Ad. rotundiloba pristina Zone = middle MN Zone 1) correlates with the main deepening pulse of the Frasnes Events in the Ardennes and North Africa (Aboussalam and Becker 2007;Narkiewicz and Bultynck 2010). It should be called Basal Frasnes Event in order to enable a clear distinction from the previous top-Givetian deepening.
A small peak of conodont diversity, with seven species, including a new form, is reached in Bed 11 (Ad. rotundiloba soluta Subzone, MN Zone 2a). The microfacies is intermediate between the bioclastic MF-B4 and floatstones of MF-B5c/d. There is a gradual return to lighter grey limestone with a higher amount of tabulate corals, such as Alveolites and Platyaxum (Roseoporella) (Becker et al. 2016c, fig. 16), and some stromatoporoids (Stachyodes, Stromatoporella). These indicate a minor shallowing but the "reefal garden" (alveolitid-stromatoporoid floatstone, MF-B5b-d) received a lot debris. In the following three beds (beds B12b-B13a), the microfacies became again more organic rich but otherwise without much change (MF-B4 grading into MF-B5d). This episode may correlate with eutrophication and deepening of the Ad. rotundiloba soluta Subzone known from the onset of dark, organic-rich marls at Giebringhausen in the eastern Sauerland , from the Blauer Bruch section in the Kellerwald (Sandberg et al. 1989, base of Unit H), and from the lower Frasnian maximum of black limestone accumulation in North Africa (Lower Styliolinite, Aboussalam and Becker 2007). The change of conodont biofacies at the top (Bed 13a, sudden income of icriodids) indicates a subsequent minor shallowing, leading to the P-I Biofacies of drowned carbonate platforms of Narkiewicz et al. (2016).
With Bed B13b, another microfacies change occurred. First, there are light-grey alveolitid-thamnoporid floatstones (MF-B5e), an oxygenated "coral garden" setting. After a brief detrital phase (Bed B13c, MF-B3), phaceloid rugose corals (Disphyllum) become more abundant (MF-B5g, beds B13d to D) in association with thamnoporids. This suggests improved living conditions for more fragile reef builders, interrupted by occasional storms. Stromatoporoids and hexagonariids are rare. Interestingly, a short peak of carbon isotope values occurred within this phase (Bed C, Becker et al. 2016c) still within the Ad. rotundiloba soluta Subzone. At the top, the disphyllids become rare (Bed E1, MF-B5a). Above, peloidal matrix re-appears, returning at the level of oldest Ad. recta (oldest MN Zone 2b, Bed E3) to the setting known from the base of the section (mostly MF-B1).
Remnants of the latest reefal stage of the Hönne Valley Reef are preserved among the roots of fallen trees on the backside of the cliff. Limestone plates display abundant, nicely weathered tabulate and rugose corals in light-grey, finegrained matrix. Dominant are large, flat alveolitid colonies and thamnoporids, while phillipsastreids (Haplothecia) and hexagonariids are rare. Among the Rugosa, Disphyllum is abundant and associated with some Stachyodes (Becker et al. 2016c). This "coral garden", grading into a thin coral biostrome (coral rud-boundstones, MF-B5i) represents the top of the lower Frasnian. A range of actinopterygian teeth from conodont residues show that it was a good biotope for early bony fishes. Based on the presence of last lower Frasnian conodonts in biostrome slabs and dates for the oldest Beul Formation of core HON_1101, the final biostrome extinction is correlated with the eustatic rise of the global Middlesex Event (see review in Pisarzowska et al. 2020).
Beul Formation: Exposed under more fallen trees ca. 10 metres beyond, the overlying pelagic nodular limestones (bioclastic wackestones, MF C-4) indicate a significant deepening, resulting in an oxic, subphotic setting well-below the storm wave base. We assume a water depth exceeding 80-100 m based on the lack of euphotic biota (including photosynthetic calcimicrobes, such as Renalcis or Sphaerocodium) and the upper bathymetric estimates of Franke and Walliser (1983) and Wendt and Aigner (1985). This is supported by sections in the Canning Basin (Australia), where one can walk down from reefal carbonates to micritic pelagic limestones of the lower slope (see Playford 1980, fig. 18). Calculation for the implosion depth of beloceratid ammonoids, which occur in the Beul Formation, yielded values between ca. 110 and 170 m (Hewitt 1996).
Conodont dating of the palmatolepid-dominated assemblages as ca. upper half of the middle Frasnian (Pa. housei to Pa. plana zones, MN zones 8-10) proves an outcrop gap that spans the lower part of the middle Frasnian. This is precisely the interval of extreme condensation or non-deposition in the two studied cores. Conodont residues include shark teeth and inarticulate brachiopods, which are typical in perireefal to pelagic facies. The poor macrofauna content suggests that the environment became oligotrophic. More ammonoidrich upper Frasnian and Famennian nodular limestones are exposed between tree roots and in small, scattered natural cliffs higher up on the slope of the Beul hill (e.g . Wedekind 1913a. Wedekind , 1913bLange 1929;Paeckelmann 1938;Price 1982). They included an outcrop of the Upper Kellwasser Limestone with coral-and trilobite-bearing strata just below (Feist and Schindler 1994;Weyer 2016) but details are not known.
Correlation of boreholes and outcrop (Fig. 25) In both cores, the thick, lower parts of the sections (Asbeck Member) consist of relatively monotonous, lagoonal limestones deposited in a variably calm and restricted palaeoenvironment. The microfacies fluctuates between MF-A1 to A4. It is not possible to correlate individual intervals of Amphipora rud-bafflestones (MF-A3) or of fenestral limestones (MF-A4); these record local palaeoecological shifts within the wide platform. This also applies to the three levels of brecciated limestone of core B102 (within the interval from Bed -44 to -39, marked as "br" in Fig. 5). Near the top, the initial drowning and backstepping in the course of the Frasnes Events, with changes from lagoonal to reef core facies (MF-A5), suggest to correlate the upper ca. 6.5 m of Bed -22 in HON_1101 with the ca. 13 m of Bed -30 to Bed -28 in B102. Based on the onset of early Ancyrodella in the overlying Bed -21 of HON_1101 and in Bed B3 of the Beul section, the lagoon drowning began roughly at the same time. Due to similar microfacies, the lower part of the Eisborn Member in the Beul outcrop correlates ca. with Bed -25 in core B102. The final reef phase at the top of the lower Frasnian, a biostrome with abundant tabulate and branching rugose corals, is rather similarly developed in all three sections (B102: Bed -25; HON_1101: Bed -20; Beul: top biostrome). This relative homogenization is a remarkable trend that was obviously caused by external factors, such as eustatic rise.
The base of the (hemi)pelagic Beul Formation appears to be diachronous but this is caused by the extreme condensation (HON_1101: beds -18 and -17) to local non-deposition (B102: between Bed -25 and the base of Bed -24) in the lower middle Frasnian. The discontinuous outcrop prevented sampling of the contact in the Beul section but, based on the conodonts, Sample "top 2" correlates with the sample from the basal Beul Formation in core B102. The higher part of the middle Frasnian is rather uniform but with slightly variable accumulation rates (HON_1101: 16 m, Bed -16; B102: less than 11m, Bed -24 to the lower part of Bed -18; Beul outcrop: thickness unknown). In both boreholes, the basal upper Frasnian semichatovae Event is apparently marked as a sudden intercalation by laminated black shales (MF-C6, HON_1101: within Bed -15; B102: within Bed -18). The overlying upper Frasnian is more condensed than the middle Frasnian (HON_1101: 2.8 m, Bed -14 to B102: ca. 2.15 m,. The Lower Kellwasser Limestone is thick and well-developed in HON_1101 (58 cm, Bed -12) but not identifiable in core B102. The Upper Kellwasser equivalents are also characterised by laterally variably facies (HON_1101: 25 cm, Bed -9b/c, alternating MF-C2 to MF-C4; B102: less than 10 cm, Bed -14, MF-C5 grading upwards into MF-C4). This reflects a small-scale local palaeotopography, with deposition of laminated and marly facies in calmer depressions.
In the lower/middle Famennian, the environment was more stable and monotonous in core HON_1101 (Fig. 16). In core B102, nodular limestones alternate with marls, which caused a more punctuated conodont record that ends much earlier (high in the Pa. minuta minuta Zone; Fig. 18). Schäfer (1978, Sample P250) described from the northern part of the Beul a lower Famennian (Pa. minuta minuta Zone) reworking unit with resedimented upper Frasnian taxa. This shows that at close distance variably scouring or condensed deposition occurred. Such facies differentiation suggests a small-scale palaeorelief with fluctuating bottom currents on the drowned submarine rise.

Discussion: Causes and patterns of the Hagen-Balve Reef Complex extinction
Controls on the reef demise in the studied sections In the classical reviews of Rhenish reefs (Jux 1960a(Jux , 1960bKrebs, 1967Krebs, , 1971Krebs, , 1974, extinctions were not studied in detail, hardly discussed, or attributed to regional subsidence or block faulting. It was recognised that most reefs did not survive until the end-Frasnian Kellwasser Events but that in many cases black shales directly overlie reef carbonates. The concept that "reefs suffocated in mud due to sudden major subsidence" was also adopted for the Ardennes reef belt in the west (Lecompte 1970). Based on the study of six reefs from the Ardennes to the Wuppertal region, Wilder (1989) suggested a different model in which tectonically triggered precipitation changes caused land plant spreading and increased chemical weathering, resulting in a higher terrestrial discharge of fine detritus and nutrients that, again, led to phytoplankton blooms, increased C org deposition, and euxinic conditions. The model lacked any indicators for Givetian-Frasnian humidity, vegetation cover, proxies for weathering and terrestrial runoff, or precise biostratigraphical data. Therefore, it remained highly hypothetical. For example, it did not consider other explanations, such as open marine nutrient recycling and climatically induced thermohaline upwelling (e.g. Murphy et al. 2000), to explain biogeochemical evidence for phytoplankton blooms in black shales that overlie some of the reefs.
Several processes are widely considered to cause potentially the termination of reef growth: high siliciclastic influx, drowning by rapid eustatic rise or strongly increased subsidence, emersion due to eustatic regression or tectonic uplift, the spread of anoxia, and strong fluctuations of temperature or salinity (for Devonian reefs see review of Copper 2002a). Our biostratigraphically constrained data from the Hönne Valley provide a correlation of reef drowning, backstepping and eventual extinction with three eustatic deepening pulses, the Ense Event, Basal Frasnes Event, and Middlesex Event. There are no indicators for accelerated subsidence, the reverse is true (Krebs 1974), and there was no uplift or subaerial exposure. The fully marine and stenohaline faunas indicate persisting normal salinity. Hypoxic conditions may have developed in the Eisborn Member of the Beul outcrop within the sediment due to increased influx and bacterial recycling of organic matter. But the carbon isotope trends (Becker et al. 2016c) of the dark, bituminous limestones give no support for increased primary productivity. Sea floor anoxia of the semichatovae and Kellwasser events arrived long after the reef was dead.
The carbonate production of reefs is normally able to keep up with slow sea level rises, unless its growth is restricted in muddy water, for which there is no regional microfacies evidence. This suggests that the three eustatic pulses in combination with continuing subsidence outpaced the reef builder capacities. This occurred in an interval of pulsed global climatic heating (Joachimski et al. 2009), which probably caused contemporaneous thermohaline upwelling in the adjacent Flinz basins. Kiessling and Simpson (2011) reviewed the potential of ocean acidification as a cause of ancient reef crises but found no evidence in the Upper Devonian.
Comparison with the western Hagen-Balve Reef Complex (Fig. 1) The area from Hagen to Iserlohn (Fig.1) is characterised by much earlier rapid drowning, during the Taghanic Transgression near the top of the middle Givetian (May and Becker 1996), associated with a sudden onset of organic-rich, dark, pelagic facies. The overlying Flinz limestones and black shales may contain reef debris, top-middle Givetian stringocephalids, and maenioceratids (Denckmann 1902(Denckmann , 1905Becker 1985), but also upper Givetian goniatites (Denckmann 1901b(Denckmann , 1903Becker 1985). West of the Lenne river, in the Donnerkuhle Quarry at Hagen-Eppenhausen ( Fig. 1), the sharply overlying black shales at the base of the local Eppenhausen Formation (von Kamp 1972) yielded also a pyritic, upper Givetian pharciceratid fauna (Bockwinkel et al. 2013). There was clearly a combined role of rapid sea level rise, eutrophication and suffocation by anoxia at the middle/upper Givetian boundary in that region, which predated the reef drowning in the Hönne Valley region.
Only small-sized reef caps survived, for example at the Steltenberg of Hagen-Hohenlimburg (von Kamp 1972; Fig.  1). Their reef debris is characterised by tabulates, some chaetetid sponges, and dominant solitary rugose corals that grew within the photic zone, proven by the presence of the calcimicrobes Girvanella and Rothpletzella. The lack of stromatoporoids is conspicuous (May and Becker 1996) and turbiditic limestones occur laterally. This characterises a small-scale, steep slope palaeoenvironment, which was very different from the low-relief final reef phase (Eisborn Member) of the Beul area.
The small-sized caps probably did not survive into the top-Givetian but their extinction is not yet well-dated. Higher up, there is a re-surgence of limestone turbidites ("Flinzkalk-Horizont") with some reef margin debris and intercalated marls with goniatites in the lower Frasnian (Becker 1985). This suggests a top middle Givetian reef backstepping towards the southern core of the Remscheid-Altena Anticline, followed by a lower Frasnian phase of progradation, resulting in a northward shift of debris shedding. This latter pulse correlates in time with the lower part of the Eisborn Member at the Beul.
East of Iserlohn, around Bilveringsen (Figs. 1, 3), brachiopodrich limestones form the famous Schleddenhof Member, which yielded some pre-Taghanic goniatites (Torley 1908(Torley , 1934. It represents a gentle slope setting with extremely diverse neritic assemblages. After the Taghanic deepening, the brachiopod limestones were overlain by turbiditic, dark Flinz limestones. Eastern Hagen-Balve Reef Complex (Figs. 3,25) At Hemer (Figs. 1, 25), the organic-rich Flinz limestones and shales overlying the youngest reef contain upper Givetian to middle Frasnian conodonts (Clausen and Ziegler 1989). However, a 2 m thick reef limestone lense was found south of Hemer-Höcklingsen between basal upper Frasnian beds (Heinke 1978: "Ag. triangularis Zone"; Figs. 3, 25). Its nature, autochthonous reef growth or allochthonous glide block, is unclear. To the southeast, in the Hemer-Apricke region (Figs. 1, 25), organic-rich, basinal, turbiditic Flinz facies directly overlies the last coral limestones and yielded rich basal Frasnian conodont faunas (Ad. rotundiloba pristina Zone, Hoppenberg fauna of Aboussalam and Becker 2009). Therefore, the final reef extinction in that region is also linked with a very sudded change to eutrophic black shales and turbidites, in this case coinciding with the Basal Frasnes Event. It correlates in time with the initial Hönne Valley drowning but pre-dates the final biostrome of the Beul. It seems that there was a younging of the top Hagen-Balve Formation from west to east, interrupted by local cap reefs. The reef breccia noted by Jux (1960a) at the former top of Emil Quarry (Figs. 1, 25) may have been an equivalent of the final alveolitidthamnoporid biostrome of the Beul area. It was shown to be overlain by basinal and organic-rich Flinz facies, which was confirmed during mapping by Kruse (2013). This basinal post-reefal development ends more or less with the Hönne Valley (Fig. 25).  Paeckelmann (1938) gave a section from the final reef limestones through the Frasnian to lower Famennian at the eastern slope of the B515 road that runs through the Hönne Valley (Fig. 2). It was re-studied by W. Ziegler in 1959 but only the Famennian conodont data were published in 1962. Paeckelmann (1938) noted intercalating light-grey "Adorf Limestone" and dark-grey to black Flinz limestones and shales, with black limestones with buchiolids in higher parts, probably evidence for Kellwasser beds. The important section is now overgrown but we sampled at the southern end of Oberrödinghausen, at 31 km, light-grey limestones that yielded deep-water (Palmatolepis biofacies) basal upper Frasnian conodonts of the Pa. nasuta Subzone (MN Zone 11b), as known from the Beul. There is no evidence for Frasnian Flinz facies overlying the reef limestones from there towards the east (e.g. Hacke 1999). Dornsiepen (1973) discussed the considerable decline of reef limestone thicknesses from our study area towards the south and the implications for middle Givetian subsidence variations at short distance. There are no precise data for the reef extinction midway between Oberrhödinghausen and Balve. But Schäfer (1978) emphasised that alternating thin-bedded limestones and basinal shales of Frasnian age occur at the Radeberg NE of Beckum (Fig. 1). This suggests that the post-reefal submarine rise palaeotopography descended to the southeast. Further to the south, from the Balve Cave and adjacent Helle Quarry to Garbeck (Fig. 1), the so-called Garbeck Limestone (Eder 1971; now Garbeck Memer) consists of middle Givetian, thick marginal slope reef talus transported by debris flows and turbidites into a southern basin, where limestone deposition interfingered with the active volcanism (Dobrzinski 2001;Figs. 3, 25). The top of the volcanic succession of the Husenberg at Balve (Figs. 1, 25) was locally covered by iron ore and condensed, upper Givetian goniatite limestones ("Pharciceras Kalk", Paeckelmann 1938; Fig. 25). On the flanks, slightly further to the south, pelagic shales with goniatites deposited from the top-middle Givetian on (Clausen 1989). Interestingly, the nodular, micritic Beul Formation characterises not only the drowned reef submarine rise but it expanded eventually southwards, to the Husenberg area (Figs. 3, 25).

Extinctions of other Rhenish reefs
Combined effects of eustatic and hypoxic events on the Middle/Upper Devonian reef development have previously been recognised in other Rhenish reef complexes. In the Aachen region, the Middle Devonian to lower Frasnian is characterised by a biostromal limestone succession, which is sharply overlain by an argillaceous interval, long-known as "Grenzschiefer" (Holzapfel 1910) and re-named as Inde Member by Aboussalam and Becker (2016). Conodonts date the transgression as Timan Event high in the lower Frasnian (Ad. pramosica Zone = MN Zone 3b of Aboussalam and Becker in Pisarzowska et al. 2020). Interestingly, the Timan Event has not yet been recognised in more eastern Rhenish reef successions. A full recovery of reef growth is marked in the Aachen region by the re-onset of coral and massive stromatoporoid bearing limestone ("Frasne Reef Limestone = Hahn Member). This mostly middle Frasnian reef platform drowned in the course of the basal upper Frasnian semichatovae Transgression .
In the western part of the Velbert Anticline, the Frasnes Events are developed as black shale intercalations within the Hofermühle Reef (Hofermühle South Quarry, Ellerkamp et al. 2018). The last reef caps are overlain by a poorly known middle Frasnian siliciclastic unit ("Hülsbeck-Schluffstein-Folge", Ribbert and Lange 1993). Local reef extinction may have been caused by clastic shedding during regression. Frasnian coarse siliciclastics are restricted to the Hofermühle region. This suggests that fluctuating terrestrial discharge did play a role for reef extinctions in a restricted region and that not a single extinction model can be applied to all Rhenish reef bodies.
At the eastern end of the Velbert Anticline, the Wülfrath Reef was initiated on the siliciclastic shelf (Flandersbach Formation) by the deepening of the Basal Frasnes Event (Ribbert and Lange 1993;Becker et al. 2016b). In its upper part, dominating Stromatactis bindstones form the Schlupkothen Member, a microbial deep-water (dysphotic) reef platform and subtype of the "Iberg Facies" (Krebs 1974). The record of deep-water palmatolepids indicates a platform drowning, with a partly thick intercalation of black marls, before the Pa. winchelli Zone (MN Zone 12) but with an overprint by synsedimentary tectonic tilting (Nowak 2012;Becker et al. 2016b). The Middlesex Event that finally killed the last Hönne Valley biostrome is not yet known to have affected the Wülfrath Reef but the age of a thick black marl unit is not yet constrained. The final drowning and extinction were caused by the transgressive Upper Kellwasser Event ).
Further to the south, basal Frasnian black shales with goniatites directly overlie the youngest reef limestones of the poorly studied Neandertal Reef. This underlines the different significance of the Frasnes Events for major shifts of reef growth throughout the Velbert Anticline region, simultaneously with the initial Hönne Valley drowning.
In the eastern Rhenish Massif, the last marginal slope limestones with abundant reef debris of the Burgberg Atoll (no. 5 in Fig. 1) yielded conodonts from higher parts of the lower Frasnian Duda 2020) but dolomitization masks the final reef extinction. Just to the north, the up to 1200 m thick Brilon reef succession is mostly of Middle Devonian age (no. 4 in Fig. 1). The youngest reefal limestones are younger than in the Hönne Valley (Stritzke 1989(Stritzke , 1990Brinkmann and Stoppel, in press) and were overlain by upper Frasnian (hemi-)pelagic limestones (Grottenberg Member of Burgberg Formation, . A possible role of the semichatovae Transgression requires more data. Comparisons of the Hönne Valley succession with the extinction phases of other Rhenish reefs are hampered by the still incomplete knowledge, especially for the Wuppertal (no. 1 in Fig. 1), Attendorn-Elspe (no. 6), and Warstein regions (no. 3). The combined available data for Rhenish reefs suggest a major role of eustatic and anoxic events. The overprint by local/ regional factors, such as subsidence, block faulting, volcanism and siliciclastic influx, resulted in a regionally variable significance and expression of event signatures.

Conclusions
The analysis of conodont stratigraphy, reef fauna, and carbonate microfacies in two cores and the only currently available outcrop enable a high-resolution reconstruction of initial backstepping, drowning, and final extinction of the thick Hönne Valley Reef. Results suggest general patterns of Rhenish reef extinctions that occurred well before the end-Frasnian Kellwasser Crisis. The post-reefal strata give insights into the sedimentary and palaeoecological effects of Frasnian and Famennian global events on a stable, pelagic submarine rise formed by the drowned reef platform. Main results are as follows:  (Johnson et al. 1985;Becker et al. 2020). Further work should try to find the well-established (summarised in Pisarzowska et al. 2020) global isotope signature. 12. The Beul Formation contains partly very rich pelagic conodont assemblages and consists of eight pelagic microfacies types, four of which are restricted to thin specific event levels. Frasnian samples of core HON_1101 support the refined succession around the global semichatovae Event established by Saupe and Becker (2022, this issue). 13. The global semichatovae Event Interval is developed in the Hönne Valley region as laminated, anoxic black shales, locally (core HON_1101) with four phases. A cyclic development was previously not known from the Rhenish Massif. 14. In the eastern Rhenish Massif, the extremely condensed Usseln Limestone (Gereke 2007) marks a strongly cyclic pre-Lower Kellwasser regressive phase. Only in core HON_1101, much weaker evidence for contemporaneous shallowing comes from the only thin levels of silty limestone (MF-C7) and peloidal packstone (MF-C8). 15. The Lower Kellwasser Event Interval is only in core HON_1101 distinctive. Its dark-grey, ostracod-mollusk pack-grainstones, with indications of bottom currents, differ strongly from the established typical Lower Kellwasser facies (e.g . Schindler 1990;Gereke 2007). The overlying intra-Kellwasser Interval was extremely condensed and characterised by a return to fully oxic, calm, and bioturbated (hemi-)pelagic facies. 16. The Upper Kellwasser Event Interval is characterised in core HON_1101 by recrystallized, polyphase, partly laminated, partly bioturbated, only moderately organicrich mollusk wacke-packstone. Such an event facies was previously unknown from the Rhenish Massif. It demonstrates that faunal blooms (e.g. ostracods) related to eutrophication reached shallow settings. In core B102, the event interval is more typically developed as a laminated, black calcareous mudstone. 17. Despite the limited sample size from half cores, all Famennian conodont zones were recognisable. The lower/middle Famennian facies differed at short distance and the top of the Beul Formation is strongly diachronous, probably due to variable bottom currents. 18. Pulses of reef drowning and final extinction were caused in the Hönne Valley and other parts of the northern Rhenish Massif by well-known eustatic events that involved in adjacent Flinz basins spreads of hypoxia, recorded by polyphase black shale/ limestone deposition. Regional overprint controlled the variable significance of individual events, which were probably linked to global climatic change, but details are still poorly understood. The main phase of the southern Hönne Valley volcanism clearly predates the polyphase reef decline. 19. The identified non-local factors delimiting reef growth and carbonate accumulation make it unlikely that large amounts of upper Givetian to Famennian carbonate can be expected anywhere in the subsurface continuation of the current outcrops. On the other hand, the dominant lagoonal inner platform setting of the thick top-lower/ middle Givetian Asbeck Member, with reef margin deposits mostly known from the south and northwest, suggests a continuation in the underground to the north and east.

Taxonomic notes
Ancyrognathus cf. triangularis Youngquist, 1945 (Fig. 14c) Description: In core HON_1101, the top of Bed -29 yielded a distinctive variant of Ag. triangularis with rather wide platforms and without free blade. Discussion: The many previous illustrations of the species suggest considerable variation concerning platform shapes and ornamentation. Typical Ag. triangularis possess a short free blade. Our form resembles the specimen figured by Klapper (1990, fig. 11.12). Provisionally, we name such representatives as Ag. cf. triangularis until formal morphotypes or subspecies are established. Age: Basal upper Frasnian, Pa. feisti Zone (MN Zone 11).
Icriodus cf. alternatus Branson and Mehl, 1934 (Fig. 20i) Description: Juvenile icriodid from Bed -5 of core B102 with very large, asymmetric basal cavity, three, very small nodes in the longitudinal side rows, and five medium-sized nodes in the median denticle row, with the last bigger one not protruding beyond the posterior end of the cusp. Discussion: Typical I. alternatus alternatus are slender, with a very narrow basal cavity, especially in early stages, and a median row of nodes that are smaller and lower than the side row nodes. A second morphotype of the subspecies is characterised in median to adult stages by a wide cavity that extends laterally. The subspecies I. alternatus helmsi and I. alternatus mawsonae differ in their denticulation. They were recently studied in terms of morphometrics by Girard et al. (2022) but did not include juveniles with very large and asymmetric cavity. The B102 specimen resembles the juvenile I. alternatus figured by Matyja (1993, pl. 24, fig. 2). We identify such forms as I. cf. alternatus until the morphotype nomenclature is further refined.
Icriodus aff. expansus Branson and Mehl, 1938 (Fig. 22l) Description: Spindle weakly curved, with seven transverse rows of denticles, where median and side row denticles alternate apart from the two posterior rows. Denticles are not connected by longitudinal or transverse ridges; the median row ones are partly larger and as high as the somewhat spinose side row nodes. At the posterior end, the median row continues with two elongate, ascending denticles, the last of which projects as a spine beyond the posterior platform end. The basal cavity is narrow and subsymmetric, without a spur. Discussion: In typical I. expansus (see Narkiewicz and Bultynck 2010), the side and median denticles do not alternate so strongly, and the side denticles are often somewhat elongate in transverse direction. In I. symmetricus, the denticles tend to be connected by ridges and typically also do not alter strongly. Preliminarily, we name our form I. aff. expansus. Age: Lower Frasnian, Ad. rotundiloba soluta Subzone (MN Subzone 2a), Beul outcrop, Bed E1.
Icriodus cf. tafilaltensis Narkiewicz and Bultynck, 2010 ( Fig. 23b) Description: Spindle markedly curved (> 10°), especially in the posterior third, with ten transverse rows of elongated nodes or ridges that are mostly connected to the median denticle row (not in the transverse rows 3-5), and with two large middle row denticles at the posterior end, the last one projecting beyond the large, strongly asymmetric basal cavity. Discussion: Typical I. tafilaltensis are characterised by the merging of the two posterior nodes, which are well separated in our specimen; hence the cf. identification. In addition, I. tafilaltensis has not yet been described from the middle Frasnian. The number of transverse nodes and shape of the basal cavity differ from I. symmetricus. Age: Middle Frasnian, Pa. housei Zone (MN Zone 8), Beul outcrop, Sample top 2.
Mesotaxis guanwushanensis Tian in Hou, 1988 Morphotype 3 (Fig. 22s) Description: Platform leaf-shaped, subsymmetric, with a short free blade and a straight, long carina. The basal pit is slightly enlarged and elongate. Discussion: Aboussalam and Becker (2007) regarded M. falsiovalis as a senior synonym of the Chinese Mes. guanwushanensis. However, there is a small difference that enables a separation of both taxa. In Mes. falsiovalis, the minute basal pit is subcircular while it is narrow and elongate in Mes. guanwushanensis. Based on this criterion, three morphotypes are separated in the latter. M1, which includes the holotype, lacks a short free blade and the carina does not reach the posterior platform end. In M2, the carina is also short but there is a free blade. Specimens with a free blade and isolated carina nodes at the posterior end are assigned to M3. In the latter, a basal pit that is slightly wider than the aboral carina, as in the Beul specimen, suggests transition towards Z. ovalis. The stratigraphic ranges of the three guanwushanensis morphotypes are not yet fully established. Age: Top-lower Frasnian, Beul outcrop, Sample top reef.
"Ozarkodina" aff. nonaginta Klapper, Kuz´min and Ovnatanova, 1996 (Fig. 14f) Description: Specimens from HON_1101, Bed -29, differ from typical "Oz." nonaginta by subdued denticles at the rectangular junction of the two blade parts. Discussion: "Ozarkodina" nonaginta was placed by Dzik (2002) in his prioniodinid genus Pluckidina. However, the type-species of this genus is a ramiform element, and the supposedly associated Pa element lacks the typical bending of the "Oz." nonaginta Group. Most recently, Kotik et al. (2021) used the generic name "Lagovina" (not Lagovidina, a Famennian prioniodinid genus), which appears to be a nom. nud. Therefore, there is still no valid genus name for the nonaginta Group, left here in "Ozarkodina" until further revision. The Beul form may represent a new subspecies that is stratigraphically younger than the established upper range of "Oz." nonaginta (see Klapper 1997). Open nomenclature is used until the variability of the latter is better documented. Age: Basal upper Frasnian, Pa. feisti Zone (MN Zone 11).
"Ozarkodina" cf. plana (Bischoff and Ziegler, 1957) (Fig. 23h) Description: Blade with high, alternating large and smaller teeth anterior of the elongated, slightly laterally expanded, shallow basal cavity, a minor incision above the cavity end, and four isolated posterior teeth that gradually become much shorter, with the tips forming a line that arches downwards. Discussion: The most similar "ozarkodinid" is the Givetian "Oz." plana but our top-middle Frasnian specimen may belong to an un-named homeomorphic taxon. Age: Top-upper Frasnian, Pa. plana Zone (MN Zone 10), Beul outcrop, Sample top 3-1.
"Ozarkodina" aff. sannemanni (Bischoff and Ziegler, 1957) ( Fig. 23i) Description: Blade straight and with very narrow, short platforms with one or two small marginal nodes, developed on both sides above the small, elongated basal cavity.
Discussion: The only very incipient development of platforms with few nodes differs from typical "Oz." sannemanni sannemanni, which normally does not occur above the lower Frasnian (Klapper 1997). The "Oz." aff. sannemanni from the Frasnian of Brazil described by Cardoso et al. (2015) is curved and displays a very different type of basal cavity and asymmetric minor platforms. Age: Top-upper Frasnian, Pa. plana Zone (MN Zone 10), Beul outcrop, Sample top 3-1.
Palmatolepis cf. housei Klapper, 2007 (Figs. 15a, 19b) Description: Core HON_1101 yielded a unique Palmatolepis with broad, lappet-like side lobe, orientated slightly anteriorly, and with two sini of the posterior platform margin, followed by an incurved posterior apex. The carina is first straight, then sigmoidal, and becomes straight and very fine after the central node. Core B102 produced a very different form with wide platform, subcircular posterior platform margin, and slightly curved carina that terminates with the central node.
Discussion: The HON_1101 specimen is interpreted as an atypical variant of Pa. housei that shows morphological trends towards both Pa. proversa (side lobe orientation) and Pa. semichatovae (lappet shape of side lobe). The B102 specimen differs from Pa. housei in the lack of a posterior carina and of a pointed posterior platform tip but the wide platform and fine anterior carina are similar: In the absence of any other similar species, it is also interpreted as an atypical Pa. housei variant. Age: Middle Frasnian, ?Pa. housei Zone (MN Zone 8), core B102, Sample -24f; basal upper Frasnian, Pa. feisti Zone/ Subzone (MN Zone 11), core HON_1101, Sample -16c.
Palmatolepis cf. plana Ziegler and Sandberg, 1990 (Fig. 19f) Description: Sample -24e of borehole B102 yielded a palmatolepid with a very fine posterior carina and strongly asymmetric, anteriorly positioned, wide side lobe, which margin runs almost straight towards the posterior tip. Discussion: We interprete the specimen as transitional from Pa. housei towards Pa. plana but closer to the latter because of the subrectangular anterior demarcation of the side lobe. Therefore, we record it in open nomenclature as Pa. cf. plana. Age: Top-middle Frasnian, Pa. plana Zone (MN Zone 10).
Palmatolepis n. sp. (Fig. 14l) Description: Several small, narrow palmatolepids with straight carina, and well-defined side lobe differ from named lower to basal middle Frasnian palmatolepids by their chagrin platform.
Polygnathus aff. varcus Stauffer, 1940 (Fig. 23g) Description: Free blade very long (ca. 60 % total length), platform short, rather flat, smooth apart from two incipient nodes where it is widest, margins not upturned and without anterior collar, platform carina with four larger merged nodes and small nodes at the posterior end, basal cavity just anterior of the platform. Polygnathus sp. (Fig. 23c) Description: A fragmentary polygnathid from the lower Beul Formation of the Beul outcrop is characterised by a flat, weakly nodose platform and an unusually fine, strongly curved carina that reaches the posterior end. Discussion: Polygnathids with such platform morphology are uncommon in the middle Frasnian pelagic facies of the Rhenish Massif. Ziegler et al. (2000) included similar forms in Po. foliatus, which lectotype, however, is an objective junior synonym of Po. dubius (Huddle, 1970)  Zieglerina n. sp. B (Fig. 22h-i) Description: Platform narrow, elongate, subsymmetric, and with pointed posterior tip (spear-shaped). Ornament with short ribs and minor nodes along the platform margins that is characterised by a net-like microstructure. Free blade with six merged denticles ascending in height anteriorly, continuing as a straight carina consisting anteriorly of merged denticles, then of six isolated, gradually smaller denticles that reach the posterior end. Basal cavity centred at ca. the middle of the entire length (0.92 mm) and at ca. 1/3 under the anterior platform, very wide (> 60 % platform width), asymmetric, narrowing but extending far towards the posterior end, filled by concentric growth lamellae. Discussion: The extended, asymmetric basal pit clearly places the new form in Zieglerina, which is partly homeomorphic to the upper Givetian genus Klapperina. The narrow platform and pit shapes do not resemble that in any named or figured species of both genera. This would justify the introduction of a new species but since re-sampling failed to provide additional representatives and since specimen B9A.14.73 is fractured, we currently apply open nomenclature (with B for Beul).
There is no evidence of pathological morphology. The netlike microstructure underlines the good preservation. Age: Lower Frasnian, Ad. rotundiloba soluta Subzone (MN Subzone 2a), Beul outcrop, Bed B11. for their help and support during the preparation of thin-sections. We are grateful to Eva Kuropka, Davina Mattijssen, and Lukas Afhüppe (Münster) for their assistance with the conodont samples. Traudel Fährenkemper (Münster) put considerable efforts into our diagrams. We appreciate the careful reviews by Peter Königshof and of an anonymous reviewer.
Funding Open Access funding enabled and organized by Projekt DEAL. The Ph.D. project of the first author was funded by the Lhoist Germany Rheinkalk GmbH.
Data availability All data generated during or analysed during the current study are included in this published article.

Declarations
Conflict of interest The authors declare that they have no conflict of interest.
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