Strontium and oxygen isotope evidence for the origin and movement of cattle at Late Neolithic Durrington Walls, UK

The geographic origins of livestock found at the Late Neolithic site of Durrington Walls (Wiltshire, UK) is explored using strontium (87Sr/86Sr) and oxygen (δ18OcarbVSMOW) isotope analysis of tooth enamel as an archive of lifetime movement. The analysis of 49 cattle is augmented with data for small numbers of animals from the contemporaneous monumental centres of West Kennet Palisade Enclosures (4), Stonehenge (1), and Marden (1). Unburnt human remains are scarce at these sites and the suite of biomolecular analyses that can be undertaken on cremated remains is limited. Therefore, these animals provide the best proxy for the origins of the people who raised them and give key information on livestock management. This builds on the Sr isotope analysis of 12 animals previously published from Durrington Walls and complements recent research on pig remains from the same sites, providing further evidence for the scale of human and animal movement and the catchment of these sites. The strontium isotope signatures from the animals’ teeth range between values that are consistent with local chalkland grazing to radiogenic values typical of granites and older rock types. The oxygen isotope data, coupled with the strontium results, provide new geographic resolution and indicate that the majority of the animals come from southern and western areas of Britain.


Archaeological context and previous work
During the Late Neolithic, Durrington Walls was the focus of feasting activities that included the slaughter and consumption of a substantial number of animals (Albarella and Serjeantson 2002;Richards and Thomas 1984). This was evidenced by the work carried out on the considerable quantity of faunal remains associated with the henge, its internal features, and extensive middens excavated in the 1960s (Wainwright and Longworth 1971) and, between 2004and 2007, by the Stonehenge Riverside Project (Parker Pearson et al. 2011). Among the animals consumed at the site, pigs (Sus domesticus) were predominant, but the remains of cattle (Bos taurus) were also abundant in the zooarchaeological assemblage. Evidence from pig age-at-death analysis suggests that the intensity of feasting varied seasonally, with a peak during the winter months (Albarella and Payne 2005;Wright et al. 2014). The question of how, and from where, such largescale consumption was provisioned was one of the central research foci of the Feeding Stonehenge Project, funded by the Arts and Humanities Research Council (AHRC).
A pilot study carried out in 2009-2010 hinted that the provisioning of livestock to the site was complex. Using strontium isotope analysis of cattle teeth, the research suggested that livestock found at the site were from diverse geographical origins. While some of the cattle were likely to be local, a proportion of the animals had 87 Sr/ 86 Sr values that were inconsistent with the Cretaceous chalk on which the site is located (Viner et al. 2010). These animals must have been raised away from Durrington Walls and transported to the site after their teeth had formed.
The aim of this paper is to provide detail to the story of cattle origins in the Late Neolithic. A larger dataset, the integration of strontium and oxygen isotope analysis, and the inclusion of a small number of teeth from other Late Neolithic sites (West Kennet Palisade Enclosures, Marden and Stonehenge) in the study area were all necessary to provide greater resolution (Fig. 1). This approach has the potential to determine the extent of cattle movement to Durrington Walls, the likely origins of the livestock that had been moved, and whether the movement of cattle was part of a wider phenomenon of livestock mobility.

Tooth formation
Tooth enamel does not undergo any remodelling and hence its isotope composition is fixed during the period of formation. This means that analytical results from cattle third molar enamel will provide a snapshot of the prevailing conditions during the period from approximately 9 to 30 months of a cattle's life (Balasse 2002). Unlike tooth enamel, both tooth dentine and bone are susceptible to diagenetic alteration due to their more porous and less crystalline structure (Trickett et al. 2003); hence, bone and dentine provide information about the burial environment rather than a lifetime signature. In addition, these tissues remodel throughout life so, even if they were not diagenetically affected, they would provide a blended isotope signal for the years before death that can be difficult to interpret.

Strontium ( 87 Sr/ 86 Sr) isotope analysis
Strontium isotope analysis is an effective tool for identifying mobility in the past. It has been used successfully by researchers interested in tracking animal migration (Britton et al. 2009(Britton et al. , 2011Hoppe et al. 1999;Julien et al. 2012), for the study of trade networks (Madgwick et al. 2012(Madgwick et al. , 2019aMadgwick and Mulville 2015;Minniti et al. 2014;van der Jagt et al. 2012), and patterns of animal movement Bendrey et al. 2009;Bentley 2006;Bogaard et al. 2013;Evans et al. 2007;Sykes et al. 2006;Viner et al. 2010). The method is based on the principle that rubidium 87 Rb decays to 87 Sr over time, changing the ratio of 87 Sr to 86 Sr. As a result, older rocks and Rb-rich rocks have higher ratios of 87 Sr/ 86 Sr, with the effect that biologically available 87 Sr/ 86 Sr varies spatially according to the age and chemistry of the underlying geology. The link between biologically available Sr and underlying geology is well documented (Ericson 1985), but other factors, such as the 87 Sr/ 86 Sr ratio in ground and river water, and in some cases sea-spray (Bentley 2006), may also contribute.
All the sites in this study are located on Cretaceous chalk, which has a relatively well-defined strontium isotope biosphere range of 0.7083 ± 0.0006 (1SD, n = 85; Evans et al. 2018). Dentine and bone samples from Durrington Walls give a similar result of 0.7086 ± 0.0004 (1SD, n = 11; Viner et al. 2010), which is to be expected for tissues that absorb the burial environment strontium isotopic signal. These data represent the predicted Sr isotope composition for tooth enamel of animals that graze on a chalk-founded terrain.

Oxygen (δ 18 O) isotope analysis
Oxygen isotope analysis has also been successfully applied to the investigation of animal mobility (Britton et al. 2009;Henton et al. 2010;Madgwick et al. 2019a;Towers et al. 2011Towers et al. , 2017. Environmental oxygen isotope ratios are dependent on the hydrological system and reflect the fractionation of 18 O compared with 16 O. In general, precipitation is increasingly depleted in 18 O at high latitude (Bentley and Knipper 2005). In Britain, because our weather systems are predominantly from the Atlantic, the oxygen isotope zonation in Britain is predominantly west to east; the lowest values were recorded in the eastern Highlands of Scotland (Darling et al. 2003). Oxygen isotope ratios in tooth enamel have a linear relationship with water ingested during the period of tooth formation and can therefore provide information about the geographical origin of animals from archaeological sites. However, using calibrating equations to 'map' enamel values on to geographically determined water values introduces additional uncertainty onto the data, and their application is therefore questionable . In this study, we have used data from animals from central and eastern England, which represent the British drinking water zone of − 7 to − 8‰, as defined by Darling et al. (2003). These provide reference data against which we compare the Late Neolithic data. These comparative datasets come from the feral cattle from modern Chillingham Castle in Northumberland: (Towers et al. 2014(Towers et al. , 2017 and Bronze Age cattle from Irthlingborough, Northamptonshire, and Gayhurst, Buckinghamshire (Towers et al. 2011).

Carbon (δ 13 C) isotope analysis
Carbon isotopes in tooth enamel record the δ 13 C of plants consumed by cattle. These data are of limited use for addressing the core aims of this research, centering on origins and movement, but are presented here to augment the oxygen isotope data and to explore varied husbandry strategies. Plants are divided into two main groups based on their photosynthetic pathways: C3 and C4 (Schwartz and Shoeninger 1991), and these two groups have substantially different δ 13 C isotope ranges. However, C4 plants, which are common in more arid environments, are rare in the indigenous flora of the British Isles and northern Europe. Therefore, British studies are restricted to variations in terrestrial C3 plants, and the transmission of the carbon δ 13 C composition to the animals that graze on them. This range has been defined for terrestrial grazing cattle as δ 13 C apatiteVPDB between − 15 and − 9‰ for pre-industrial British animals (Gan et al. 2018). The range can be extended to > − 8‰ for animals that graze on seaweed (Balasse et al. 2005(Balasse et al. , 2006. Temperature, altitude, latitude, canopy, and mean annual rainfall (MAR) can all affect the δ 13 C of terrestrial plants (Hare et al. 2018;Kohn 2010). While carbon isotope composition predominantly relates to diet, variation has the potential to shed light on the environments where the cattle were raised.

Materials
The dataset comprises intra-tooth strontium ( 87 Sr/ 86 Sr), oxygen (δ 18 O carbVSMOW ) and carbon (δ 13 C apatiteVPDB ) values for 55 cattle teeth. The vast majority of teeth (n = 49) derive from Durrington Walls, with small numbers of specimens analysed from broadly contemporaneous deposits at West Kennet Palisade Enclosures (n = 4), Marden (n = 1), and Stonehenge (n = 1). The presented data are augmented by 12 from a previous pilot study on Durrington Walls (Viner et al. 2010), to make a total number of 67 specimens.
Durrington Walls is the largest henge monument in Britain, covering around 17 ha. It is located on the chalk downlands of southern Britain and is just 3 km from Stonehenge on the west bank of the River Avon (Parker Pearson et al. 2011). Excavations during the Stonehenge Riverside Project found the remains of nine houses, as well as a variety of other features including middens, one of which is very substantial, in addition to the henge itself. An extensive programme of radiocarbon dating has narrowed the main period of settlement activity at the site to around half a century, beginning in 2525-2470 cal BC and ending in 2480-2440 cal BC (Marshall In prep;Parker Pearson et al. 2011). The main period of settlement at the site overlaps with the erection of the sarsen circle and trilithons at Stonehenge (Parker Pearson et al. 2011). All of the specimens used in this study came from the recent excavations at Durrington Walls, undertaken as part of the Stonehenge Riverside Project between 2004 and 2007. Postexcavation analysis of the animal bone assemblage was carried out at the University of Sheffield, initially as part of the Stonehenge Riverside Project, and then the Feeding Stonehenge Project between 2005 and 2013. The single Stonehenge cattle specimen (SH01) was excavated in 1924 (Cleal et al. 1995, pp. 88, 442, fig. 247). The Marden specimen derives from the excavations described in Wainwright et al (1971), but the site has been subject to new excavations in recent years (Leary et al. 2016). Marden is the second largest henge enclosure in Britain covering an area of 14 ha and is located at the edge of the chalkland c. 14 km north of Durrington Walls, in the Vale of Pewsey, Wiltshire. West Kennet Palisade Enclosures is a double enclosure site located c. 27 km north of Durrington Walls and is part of the Avebury complex (Whittle 1997;Bayliss et al. 2017). All sites are founded on chalk lithology.
All the analysed specimens derive from secure contexts and can be confidently defined as of Late Neolithic date, the majority deriving from the rich midden deposits at Durrington Walls (Viner et al. 2010). For oxygen and carbon isotope analysis, the number of incremental samples varied from five to 13 depending on dental attrition. Two samples were analysed from each of the newly selected cattle for strontium. Details of the sampling methods are provided below. In addition, a single human tooth (DUR50) from Durrington Walls was analysed. This tooth (an upper second premolar) derived from the surface of a ceremonial avenue (context 585) leading from the timber Southern Circle to the River Avon.

Analytical methods
Cattle mandibular third (M 3 ) molars were selected for analysis. The lingual cusp of the anterior pillar of each tooth was removed, abraded to remove calculus and debris, and sliced transversally at approximately 3-mm intervals (Appendix Figure 1 in the ESM). Between five and 13 enamel slices were obtained per tooth depending on the degree of dental wear. The sequential numbering of the slices was from root-enamel junction to the occlusal surface. The slice numbers are given in the tables. Two slices, one from close to REJ and one from the centre of slice sequence, were used for Sr isotope analysis. Oxygen isotope analysis was undertaken on alternate slices taken down length of the tooth. Up to six slices per tooth were analysed to represent the range of oxygen compositions within the tooth. Enamel was separated from dentine mechanically at the University of Sheffield and then transferred to the NERC Isotope Geosciences Laboratory at Keyworth to complete the process. The single human sample was removed from the root-enamel junction on the buccal side of the tooth and treated in the same way as the cattle enamel samples.
For Sr isotope analysis, the enamel samples were cleaned ultrasonically in high purity water, then rinsed twice in high purity water and high purity acetone. They were then weighed into pre-cleaned Teflon beakers and mixed with 84 Sr tracer solution and dissolved in Teflon distilled nitric acid (8 M HNO 3 ). Strontium was collected using Eichrom AG50 X8 resin columns and then loaded onto single rhenium filaments following the method of Birck (1986). The isotope composition and concentrations were determined by thermal ionisation mass spectrometry (TIMS). The international standard NBS 987 for 87 Sr/ 86 Sr gave a value of 0.710253 ± 0.00006 for static analysis (1SD, n = 350).
For the isotopic analysis of carbonate oxygen, the 10-mg enamel sample was reduced to a fine powder using an agate mortar and pestle. Approximately 3 mg of the enamel powder was loaded into a glass vial and sealed with septa. The vials were transferred to a hot block at 90°C on the GV Multiprep system. The vials were evacuated and four drops of anhydrous phosphoric acid were added. The resultant CO 2 was collected cryogenically for 14 min and transferred to a GV IsoPrime dual inlet mass spectrometer. The isotope values are treated as a carbonate. δ 18 O is reported as per mil (‰) ( 18 O/ 16 O) normalised to the PDB scale using a within-run calcite laboratory standard (KCM) calibrated against SRM19 and NIST reference material and were converted to the VSMOW scale using the published conversion equation of Coplen (1988): VSMOW = (1.03091 × δ 18 O VPDB ) + 30.91. Analytical reproducibility for laboratory standard calcite (KCM) is for δ 18 O VSMOW = ± 0.05‰ (1σ, n = 20) and δ 13 C VPDB is ± 0.03‰ (1σ, n = 20) and analytical reproducibility for an inhouse tooth enamel standard is δ 18 O VSMOW = ± 0.32‰ (1σ, n = 9) and δ 13 C VPDB is ± 0.17‰ (1σ, n = 9).

Results and discussion
Oxygen and carbon isotope data The stable δ 18 O carbVSMOW and δ 13 C carbVPDB isotope data for the incremental enamel samples are given in Table 1 and summary statistics are given in Table 2 and displayed using box and whisker charts in Figs. 2 and 3.  The median δ 18 O carbVSMOW intra-tooth values range between 23.74‰ (DUR 10) and 26.77‰ (DUR 31). The interquartile range (IQR: the central 50% of the data range) of the δ 18 O carbVSMOW values ranges widely. Of the animals that have three or more slices measured per tooth, DUR 31 has the smallest IQR of 0.33 and DUR 7 has the largest IQR of 2.97. The typical IQR seasonal range in British cattle, as represented by the Chillingham herd, is 1.23. This value can be taken to represent the natural variation in drinking water in a static herd. This suggests that animals with a substantially lower IQR (i.e. less variation) probably relied on a more seasonally stable water source, such as aquifer-sourced rivers or deep, low evaporation lakes. Alternatively, they could have been subject to seasonal movement that would dampen the impact of intra-annual variation in water oxygen (e.g. summer upland and winter lowland pasture). Conversely, those animals with a large range of δ 18 O carbVSMOW in their teeth might be enhancing the range of water oxygen they are exposed to through shallow evaporated ponds and lakes, or may be incorporating a wider range of water oxygen through movement to varied pasturage, as part of either seasonal management regimes or population movement.
The range of carbon isotope values between animals also varies widely. Five samples (DUR 35, DUR 36, DUR 38, DUR 45, and WK3) have δ 13 C carbVPDB median values above − 13‰, the highest being − 11.89‰ (DUR 45). The lowest median value is − 14.8‰ (DUR 24). As in the oxygen data, there are considerable differences in the magnitude of intratooth variation, as reflected in the IQR values. The smallest IQR is 0.04 from three slices (DUR 18) and the highest 1.5 from four slices (DUR 8). The Chillingham animals record an average IQR of 0.4. These values are within the range − 9 and − 15 for pre-industrial terrestrial grazing British cattle (Gan et al. 2018). The differences in interquartile ranges may reflect variation in animal management, those with a small IQR having more constant annual environments, and those with a larger IQR potentially grazing in areas of more variable canopy cover and/or rainfall levels.
The intra-tooth variations in oxygen and carbon reflect a heterogeneity in the manner in which these animals accessed water and plants. This is strongly suggestive of disparate origins with respect to drinking and grazing habits. This withintooth diversity extends, in some degree, to the strontium data described below.

Strontium isotope data
The Sr isotope variations within the teeth (Tables 2 and 3) are plotted in Fig. 4 in ascending value of the samples from occlusal surfaces (the earliest developing sample), showing the rest of the incremental data relative to this.
Of the 64 cattle teeth that were analysed, only 11 have data that fall at least partially in the range defined for chalk and, of these, only three animals have values that are all consistent with chalk (Table 4). Therefore, results indicate that the  The Sr data are consistent with the vast majority of individual animals being raised in a single location, though it is clear that the animals derived from a wide-ranging area and were brought to the Stonehenge landscape later in life. This suggests that intra-tooth variation in oxygen is most likely to relate to seasonal variation in water oxygen availability or management strategies, rather than long-distance movement of animals in early life. In summary, the isotope composition of the cattle provides evidence for wide-ranging origins, but no indication of substantial movement in early life. Some intra-tooth oxygen and carbon values are wide-ranging and others are tightly constrained, indicating that management strategies for the cattle were variable, with some showing far more marked seasonal effect on their biogenic values. The next section will focus on inter-tooth, rather than on intra-tooth, values, in an attempt to explore the origins of the animals.

Establishing the geographic origins of the animals
Oxygen isotope composition provides a broad-brush method for subdividing the data through comparison with animals of known geographic origin. Figure 5 presents a box and whisker diagram that compares the mean δ 18 O carbVSMOW values for animals from this study with those of animals from eastern and central England (Towers et al. 2010(Towers et al. , 2011(Towers et al. , 2017. In making this comparison, a number of issues have to be considered, including irregular wear patterns, methodological differences in preparation, possible biases to median value where sample numbers are low, and reliability of place of origin. These issues are discussed and addressed in the appendix in the ESM. With these caveats in mind, we present the following observations. There is a clear separation between the data from the two groups, with the animals from this study recording significantly     (Darling et al. 2003).
The strontium data further refines the origins of the cattle. Twenty percent of the animals are compatible with grazing on chalklands, which could infer an origin close to where they were found. The majority (66%) of the Sr data fall between the upper limit of chalk 0.7089 and 0.7110. These values are widely available in Britain and provide little geographical constraint. They could represent origins in the broader region of central southern Britain, but are also commonplace in various areas throughout the British Isles (Evans et al. 2018). However, the higher values between 0.711 and 0.713 are less common in the British biosphere. Fifteen samples from 11 individuals record values in this range. The closest areas to Durrington Walls and the other Late Neolithic sites that record this Sr isotope biosphere data range are in Wales and south west England. Similar values from cremated human remains were recently interpreted as likely to derive from West Wales (Snoeck et al. 2018). Such values are also recorded further north in Scotland. Eight animals (16 samples) record highly radiogenic Sr isotope compositions over 0.713. According to current mapping (Evans et al. 2018), biosphere values of > 0.7132 are virtually absent from southern Britain, though are present in restricted zones around the Malvern Hills (Chenery et al. 2010). They are recorded in small areas around the Lake District in northern England, but are most common across large areas of older geology in Scotland. However, there is growing evidence that, perhaps, there is a source of more radiogenic biosphere values available in England and Wales based on finds that seem unlikely, on archaeological grounds, to have originated in Scotland. Neolithic human data from Penywyrlod, South Wales (0.7132 to 0.7165), have a very similar 87 Sr/ 86 Sr range to the most radiogenic Durrington Walls cattle. These individuals are interpreted as having spent their childhood in Wales (Neil et al. 2017). Similarly, Roman fauna from Caerleon, South Wales (Madgwick et al. 2019b), and Worcester (Gan et al. 2018) have produced highly radiogenic values, 0.71628 and 0.71582 respectively, suggesting that other radiogenic sources may be present in southern Britain.
An overseas origin is highly unlikely for the animals of this study, as the Late Neolithic is distinctive in having no evidence for continental contact (Vanderlinden 2012;Wilkin and Vanderlinden 2015) and therefore origins must be explored in a British context. Increasing evidence of pockets of radiogenic biosphere values may explain the origins of some of these animals. However, there can be little doubt that such values can only be attained in restricted areas of the landscape, and the large range of values (0.7133 to 0.7168) in the dataset suggests that the animals in this study derive from different areas of radiogenic geology, potentially including Scotland.
In summary, very few of the animals could be described as locally raised with respect to a typical chalk-based Sr isotope value. However, large tracts of Britain can accommodate the  Comparison with pig and human data from Durrington Walls Durrington Walls was a major ceremonial site and, while this paper examines cattle data from there, a recent study provides a comparative dataset on pigs, which were the principal feasting animal at the site (Madgwick et al. 2019a). Figure 6 highlights clear similarities in the range of values represented in both datasets, which can be divided into four groups based on breaks in the dataset shown by both pigs and cattle. The majority of animals (group 1) are consistent with an origin on the chalk and other Mesozoic deposits that dominate much of southern England and are common across Britain. Groups 2 and 3 are from more radiogenic terrains, probably characteristic of granitic and Palaeozoic areas, the closest of which are in southwest England and Wales (Evans et al. 2018). A final group (group 4) has distinctive values of > 0.714 and must derive from areas of more radiogenic geology. As discussed, on the basis of current biosphere mapping data,  origins in Scotland cannot be excluded for at least some animals, but others may derive from radiogenic areas yet to be mapped in detail in Wales or England. For both pigs and cattle, the majority of the data plot between 0.708 and 0.7108. The datasets primarily differ in the proportion of animals with lower values associated with chalk: 28% of the pigs are below 0.709, in contrast to 17% of the cattle. This suggests that a greater proportion of the pigs may have derived from chalk based terrain. However, this does not necessarily mean that the animals were raised on the Wessex chalkland surrounding Durrington Walls, as many of the pigs have an elevated sulphur value of > 14‰ that indicates a coastal origin (Madgwick et al. 2019a). The dataset indicates that the people that raised and brought cattle and pigs to Durrington Walls and other Late Neolithic monumental centres came from wide-ranging areas, and that, in some instances, may have travelled with both types of livestock.

The human tooth
Human remains from Durrington Walls are very rare, and just one tooth (DUR 50) was available for isotope analysis. This tooth (an upper second premolar) produced an 87 Sr/ 86 Sr value of 0.7126. This value is higher than values (0.7078 to 0.7118) obtained from cremated bone from Stonehenge presented in Snoeck et al. (2018), but should not be treated as directly comparable. Cremated bone gives an average of strontium uptake in the years before death, rather than the more temporally defined value in childhood that the analysis of dental enamel provides. The result is inconsistent with the local chalk range and suggests that the individual moved t o D u r r i n g t o n Wa l l s s o m e t i m e a f t e r t h e f u l l mineralisation of the tooth (around age 6 years; Hillson 1996).  (Towers et al. 2011(Towers et al. , 2014(Towers et al. , 2017. The mean (x) and median (-) are given and the box represents the interquartile range (IQR) or central 50% of the data. The whiskers extend to beyond the IQR box by 1.5 times the IQR. Values beyond this range plot a dot

Conclusions
These results have demonstrated the diversity of cattle origins at Late Neolithic Durrington Walls. The 87 Sr/ 86 Sr isotope data suggest that at least four distinct terrains are represented in the dataset. The majority of animals are consistent with an origin on the chalk and other Mesozoic deposits that dominate much of southern England and are common across Britain. Many of these animals must have been imported to the sites, though not necessarily over long distance. Two distinct groups of cattle are from more radiogenic terrains, probably characteristic of Palaeozoic areas, the closest of which are in southwest England and Wales. A final group has distinctive values of > 0.714 and must derive from areas of even more radiogenic geology. On the basis of current biosphere mapping data, origins in Scotland seem likely for at least some animals, but others may derive from radiogenic areas incompletely mapped in England or Wales. Oxygen isotope data indicate that the majority of the animals are likely to derive from western or southern areas of Britain. Highland areas in the north of England and northeast Scotland are probably not represented in the dataset, but depleted oxygen isotope compositions suggest that some animals came from eastern and/or central areas of England.
Cattle are an important component of animal bone assemblages from Late Neolithic Britain. Remains of cattle were second only to pigs in abundance at Durrington Walls, and the presence of large quantities of cattle remains, along with evidence of butchery and burning on many bones, indicates that they were included in feasting activities (Albarella and Serjeantson 2002). The zooarchaeological evidence is also consistent with an introduction of cattle to Durrington Walls, due to the almost complete absence of neonatal bones. Such remains are expected to occur in breeding areas, because of natural casualties-their absence therefore suggests that husbandry largely occurred off-site.
The movement of cattle over long distances is an example of their importance in Neolithic society. Not only were they a significant source of food, but their role in feasting was important enough to warrant a huge investment of time and energy in herding them over long distances. These animals clearly had a role to play in sustaining long-distance networks in Late Neolithic Britain. As a proxy for human movement, the cattle from Durrington Walls are representative of the human journeys that were undertaken during the period and suggest links between human groups in many different parts of the country, both close and distant. The few cattle teeth from other contemporary sites hint that this phenomenon was more widespread and, perhaps, that Durrington Walls was not unique, but part of a wider network of connections and livestock exchange.
The exogenous origin of the livestock is in contrast with the largely local nature of the material culture (Chan et al. 2016). Animals could be driven on the hoof, while large quantities of objects would have been very onerous to carry. Such practical concern meant, however, that the local and the imported both played a role in the make-up of the Durrington Walls ceremonies, and probably contributed substantially to define the character of the communities occupying-permanently or periodically-the Stonehenge landscape.