Sandy Beach Ecology — A Review

  • Anton McLachlan
Conference paper
Part of the Developments in Hydrobiology book series (DIHY, volume 19)


Sandy beaches dominate most temperate and tropical coastlines where they represent both important recreational assets and buffer zones against the sea. In some areas they are very productive and are exploited commercially. However, they have been regarded as marine deserts by many biologists and were largely neglected until Remane (1933) began studies on the coasts of Germany. The work of Pearse et al (1942) was also pioneering and represented the first qualitative attempt to evaluate a whole beach system. Since then sandy beach ecology has advanced considerably, though it has always lagged behind other aspects of coastal marine ecology in the attention it has enjoyed. During this period biological research on beaches has spread from early taxonomic and qualitative studies through quantitative ecology and physiology of important species towards a more holistic systems approach today.


Sandy Beach Sand Beach Surf Zone Filter Feeder Harpacticoid Copepod 
These keywords were added by machine and not by the authors. This process is experimental and the keywords may be updated as the learning algorithm improves.


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  1. Achutankutty CT (1976) Ecology of sandy beach at Sancoale, Goa: Part 1. Physical factors influencing production of macrofauna, Indian J. mar. Sci. 5, 91–97.Google Scholar
  2. Alikunhi NH (1944) The zonal distribution of the mole crab (Emerita asiatica) on the Madras coast, J. Bombay Nat. Hist. Soc. 45, 94–96.Google Scholar
  3. Albeit J (1978) Distribution of the polychaete genus Nephtys: a stratified random sampling survey, Kieler Meeresforsch 4, 61–67.Google Scholar
  4. Alheit J and Naylor E (1976) Behavioral basis of intertidal zonation in Eurydice pulchra Leach, J. exp.mar. Biol. Ecol. 23, 135–144.CrossRefGoogle Scholar
  5. Alheit J and Scheibel W (1982) Benthic harpacticoids as a food source for fish. Mar. Biol. 70, 141–147.CrossRefGoogle Scholar
  6. Amspoker, A.M. (1977) The distribution of intertidal epipsammic diatoms on Scripps Beach, La Jolla, California, U.S.A. Bot. Mar. 20, 227–232.Google Scholar
  7. Anderson JG and Meadows PS (1979) Micro-environments in marine sediments, Proc. Roy. Soc. Edinburgh 76B, 1–16.Google Scholar
  8. Anderson JG, Boonruang P and Meadows PS (1981) Interrelationships between cholrophylls, carbon, nitrogen and heterotrophic bacteria in an intertidal sediment transect, Mar. Ecol. Prog. Ser. 6, 278–284.Google Scholar
  9. Andrews AR, Floodgate GD and Pugh KB (1976) An annual cycle at constant temperature in a model sandy beach. Part 2. Microbial biol. J. exp. Mar. Biol. Ecol. 24, 61–72.Google Scholar
  10. Angus IS (1979) The macrofauna of intertidal sand in the outer Hebrides, Proc. Roy. Soc. Edin. 77B, 155–171.Google Scholar
  11. Ansell AD (1961) Reproduction, growth and mortality of Venus striatula (Da Costa) in Kames Bay, Millport, J.mar. biol. Ass. U.K. 41, 191–215.CrossRefGoogle Scholar
  12. Ansell AD (1983) The biology of the genus Donax. In McLachlan A and Erasmus T,/ eds. Sandy beaches as ecosystems, The Hague, Junk.Google Scholar
  13. Ansell AD and Trevallion A (1969) Behavioral adaptations of intertidal molluscs from a tropical sandy beaches, J. exp. mar. Biol. Ecol. 4, 9–35.CrossRefGoogle Scholar
  14. Ansell AD and Trueman ER (1973) The energy cost of migration of the bivalve Donax on tropical sandy beaches, Mar. Behay. Physiol. 2, 21–32.CrossRefGoogle Scholar
  15. Ansell AD and Lagardére F (1980) Observations on the biology of Donax trunculus and D. vittatus at Ile d’ Oleron (French Atlantic Coast), Mar. Biol. 57, 287–300.Google Scholar
  16. Ansell AD, McLusky DS, Stirling A and Trevallion A (1978) Production and energy flow in the macrobenthos of two sandy beaches in South West India, Proc. Roy. Soc. Edinburgh 76B, 269–296.Google Scholar
  17. Ansell AD, Sivadas P, Narayanan B, Sankaranarayanan VN and Trevallion A (1972e) The ecology of two sandy beaches in South West India I. Seasonal changes in physical and chemical factors and in the macrofauna, Mar. Biol. 17, 38–62.Google Scholar
  18. Ansell AD, Sivadas P, Narayanan B and Trevallion A (1972b) The ecology of two sandy beaches in South West India 1II: Observations on the populations of Donax incarnatus and Donax spiculum Mar. Biol. 17, 318–332.Google Scholar
  19. Armitage TM and Alevizon WS (1980) The diet of the Florida pompano (Trachinotus carolinus) along the East Coast of Central Florida, U.S.A. Fla. Sci. 43, 19–26.Google Scholar
  20. Bacescu M (1975) Contributions to the knowledge of the mysids (Crustacea) from the Tanzanian water, Univ. Dar es Salaam Sci. J. 1, 39–61.Google Scholar
  21. Bally R (1981) The ecology of three sandy beaches on the west coast of South Africa, Ph.D. thesis, University of Cape Town.Google Scholar
  22. Barnard LJ (1963) Relationship of benthic amphipoda to invertebrate communities of inshore sublittoral sands of Southern Carolina, Pacific Naturalist 1963, 3. 439–467.Google Scholar
  23. Bartsch 1 and Schmidt p (1979) Distribution and ecology of some halacarid mites (Halacaridae) in sandy beaches of the Baltic, Kiel Bay, the North Sea, Sylt, West Germany and the Norwegian Sea Tromso area Norway, Mikrofauna Meeresbodens 0 (74), 163–197.Google Scholar
  24. Bascom W (1964) Waves and beaches, New York, Doubleday and Co Inc.Google Scholar
  25. Boaden PJS (1968) Water movement–a dominant factor in interstitial ecology, Sarsia 34, 125–136.Google Scholar
  26. Bolter M. Meyer-Reil L, Dawson R, Liebezeit G, Wolter K and Szwerinski H (1981) Structure analysis of shallow water ecosystems: interaction of microbial, chemical and physical characteristics measured in the overlying waters of sandy beach sediment, Estuar. cstl shelf Sci. 13, 579–589.Google Scholar
  27. Bonnet DD (1946) The Portuguese man-of-war as a food source for the sand crab (Emerita pacifica) Science 130, 148–149.Google Scholar
  28. Boucher G and Chamroux S (1976) Bacteria and meiofauna in an experimental sand ecosystem I. Material and preliminary results, J. exp. mar. Biol. Ecol. 24, 237–249.CrossRefGoogle Scholar
  29. Bousefield EL (1970) Adaptiver adiation in sand burrowing amphipod crustaceans, Cheseapeake Sci. 11, 143–154.CrossRefGoogle Scholar
  30. Bowman TE (1971) The distribution of calanoid copepods off the south eastern United States between Cape Hatteras and Southern Florida, Smithsonian Contrib. Zool. 96, 58 pp.Google Scholar
  31. Brattegard T (1969) Mysidacea from shallow water in the Bahamas and Southern Florida. Part I. Sarsia 39, 17–105.Google Scholar
  32. Brown AC (1964) Food relationships on the intertidal sandy beaches of the Cape Peninsula,, S. Afr. J. Sci. 60, 35–41.Google Scholar
  33. Brown AC (1971a) The ecology of the sandy beaches of the Cape Peninsula, South Africa Part I: Introduction, Trans. Roy. Soc. S. Afr. 39, 247–279.CrossRefGoogle Scholar
  34. Brown AC (1971b) The ecology of the sandy beaches of the Cape Peninsula, South Africa Part 2: The mode of life of Bullia (Gastropoda; prosobranchiata), Trans. Roy. Soc. S. Afr. 39, 281–319.CrossRefGoogle Scholar
  35. Brown AC (1973) The ecology of the sandy beaches of the Cape Peninsula, South Africa Part 4: Observation on two inter-tidal isopoda, Eurydice longicornis (Studer) and Exosphaeroma truncatitelson (Barnard), Trans. Roy. Soc. S. Afr. 40, 381–404.CrossRefGoogle Scholar
  36. Brown AC (1982) The biology of sandy beach whelks of the genus Bullia (Nassariidae), Oceanogr. Mar. Biol. Ann. Rev. 20, 309–361.Google Scholar
  37. Brown B (1982) Spatial and temporal distribution of a deposit-feeding polychaete on a heterogeneous tidal flat. J.exp. mar. Biol. Ecol. 65, 213–227.CrossRefGoogle Scholar
  38. Brown R (1981) Saccocirridae (Annelida: Archiannelida) from the central coast of New South Wales, Aust. J. Mar. Freshwat. Res. 32, 439–456.CrossRefGoogle Scholar
  39. Brown AC and Talbot MS (1972) The ecology of the sandy beaches of the Cape Peninsula, South Africa Part 3: A study of Gastrosaccus psammodytes Tattersall (Crustacea: Mysidacea), Trans. Roy. Soc. S. Afr. 40, 309–333.CrossRefGoogle Scholar
  40. Brunton PM (1978) Toheroa predation by black-backed gulls on Dargaville beach North Auckland, New Zealand, Notornis 25, 128–140.Google Scholar
  41. Buchanan JB (1966) The biology of Echinocardium cordatum (Echinerdermata: Spatangoidea) from different habitats. J. mar. biol. Ass. U.K. 46, 97–114.CrossRefGoogle Scholar
  42. Burger J, Howe MA, Caldwell D and Chase J (1977) Effects of tide cycles on habitat selection and habitat partitioning by migrating shore birds, Auk 94, 743–758.CrossRefGoogle Scholar
  43. Burger J, Haan DC and Chase J (1979) Aggressive interaction in mixed species flocks of migrating shore birds, Anim. Behay. 27, 459–469.Google Scholar
  44. Bush LF (1966) Distribution of sand fauna in beaches at Miami, Florida, Bull. Mar. Sci. 16, 58–75.Google Scholar
  45. Cadbury CJ (1978) The beached bird survey and other sea bird surveillance, Ibis 120, 119–120.Google Scholar
  46. Cahet G (1976) Biogeochimie du carbone et du soufre en sediments litteraux Mediterraneens,(Golfe du Lion): Modèlles biopedologiques, Vie Milieu 26, 1–25.Google Scholar
  47. Caine EA (1974) Feeding of Ovalipes guadulpensis (Saussure)(Decapoda: Brachyura: Portunidae) and morphological adaptations to a burrowing existence. Biol. Bull. 147, 550–559.CrossRefGoogle Scholar
  48. Carlisle JG, Schott JW and Abrahamson NJ (1960) The barred surfperch in southern California, Calif. Dept. Fish Game. Bull. 109.Google Scholar
  49. Cassie RM and Cassie V (1960) Primary production in a New Zealand west coast phytoplankton bloom, New Zealand J. Sci. 3, 173–199.Google Scholar
  50. Chamroux S (1965) Contribution a l’etude du pouvoir denitrifiant des sediments marins, Instit. Pasteur. Anal. 109, 424–442.Google Scholar
  51. Chamroux S, Boucher G and Bodin P (1977) Etude experimental d’un ecosystem sableux. II. Evolution de populations de bacteries et de meiofaune, Helgolander wiss. Meeresunters. 30, 163–177.Google Scholar
  52. Chelazzi G and Ferrara F (1978) Research on the coast of Somalia. The shore and dune of Sar Uanle. 19. Zonation and activity of terrestrial isopods (Oniscoida). Monitore Zool. ital. Suppl. 9, 189–219.Google Scholar
  53. Chelazzi G, Chelazzi L and Focardi S (1983) Dynamic zonation of staphylinoid beetles (Coleopatera: Staphylinoidea) on a sandy beach in East Africa. In McLachlan A and Erasmus T, eds. Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  54. Christie ND (1976) A numerical anaylsis of the distribution of a shallow sublittoral sand macrofauna along a transect at Lamberts Bay, South Africa, Trans. Roy. Soc. S. Afr. 42, 149–172.CrossRefGoogle Scholar
  55. Clark RB, Alder JR and McIntyre AD (1962) The distribution of Nephytys on the Scottish coast, J. Anim. Ecol. 31, 359–272.Google Scholar
  56. Clark RB and Milne A (1955) The sublittoral fauna of two sandy bays on the Isle of Cumbrae, Firth of Clyde, J. mar. Biol. Ass. U.K. 34, 161–180.CrossRefGoogle Scholar
  57. Clutter RJ (1967) Zonation of nearshore mysids, Ecology 48, 200–208.CrossRefGoogle Scholar
  58. Cockcroft A (1979) Observed trends in the surf zone zooplankton of Kings Beach, Unpubl. Honour Project, University of Port Elizabeth.Google Scholar
  59. Cockcroft A (1983) Aspects of the biology of the swimming prawn Macropetasma africanus (Balss). M.Sc dissertation, University of Port Elizabeth.Google Scholar
  60. Coe WC (1955) Ecology of the bean clam Donax gouldi on the coast of Southern California, Ecology 36, 512–514.CrossRefGoogle Scholar
  61. Collos Y and Lewin J (1974) Blooms of surf zone diatoms along the coast of the Olympic Peninsula, Washington. IV Nitrate reductase activity in natural populations and laboratory cultures of Chaetoceros armatum and Asterionella socialis, Mar. Biol. 25, 213–221.Google Scholar
  62. Collos Y and Lewin J (1976) Blooms of surf-zone diatoms along the coast of the Olympic peninsula, Washington 7: Variation of the carbon to nitrogen ratio in field samples and laboratory cultures of Chaetoceros armatum Limnol. Oceanogr. 21, 219–225.Google Scholar
  63. Corey S (1970) The quantitative distribution of Cumacea (Crustacea: Peracarida) in Karnes Bay, Scotland, Can. J. Zool. 48, 925–930.Google Scholar
  64. Coull BC (1970) Shallow water meiobenthos of the Bermuda Platform, Oecologia (Berl) 4, 325–357.CrossRefGoogle Scholar
  65. Craig PC (1970) Behaviour and distribution of the intertidal sand beetle Tinopinus pictus (Coleoptera: Staphylinidae). Ecol. 51, 1012–1017.CrossRefGoogle Scholar
  66. Craig PC (1973) Behaviour and distribution of the sand beach amphipod Orchestoidea corniculata Mar. Biol. 23, 101–109.Google Scholar
  67. Crane J (1941) On the growth and ecology of brachyuran crabs of the genus Ocypode Zoologica 26, 298–309.Google Scholar
  68. Crisp DJ and Williams R (1971) Direct measurement of pore-size distribution on artificial and natural deposits and prediction of pore space accessible to interstitial organisms, Mar. Biol. 10, 214–226.Google Scholar
  69. Croker RA (1967a) Niche diversity in five/ sympatric species of intertidal amphipods (Crustacea: Haustoriidae), Ecol. Monogr. 37, 173–199.CrossRefGoogle Scholar
  70. Croker RA (1967b) Niche sepcificity of Neohaustorius schmitzi and Haustorius sp. (Crustacea: Amphipoda) in North Carolina, Ecology 48, 971–975.CrossRefGoogle Scholar
  71. Croker RA (1968) Distribution and abundance of some intertidal sand beach amphipods accompanying the passage of two hurricanes, Chesapeake Sci. 9, 157–162.CrossRefGoogle Scholar
  72. Croker RA (1970) Intertidal sand macrofauna from Long Island, New York, Chesapeake Sci. 11, 134–137.CrossRefGoogle Scholar
  73. Croker RA, Hager RP and Scott KJ (1975) Macroinfauna of northern new England marine sand II. Amphipod dominated intertidal communities, Can. J. Zool. 53, 42–51.Google Scholar
  74. Cubit J (1969) Behaviour and physical factors causing migration and aggregation of the sand crab Emerita analoga (Stimpson), Ecology 50, 118–123.CrossRefGoogle Scholar
  75. Culter JK and Mahadevan S (1982) Long term effects of beach nourishment on the benthic fauna of Panama City beach, Florida, U.S. Army Corps Engineers, Misc. Rep. 82–2, 94 pp.Google Scholar
  76. Cupka DM (1972) A Survey of the icthyofauna of the surf zone in South Carolina. S. Carolina Wildlife Mar. Res. Dept. Tech. Rep 4, 14 pp.Google Scholar
  77. Dahl E (1952) Some aspects of the ecology and zonation of the fauna on sandy beaches, Oikos 4, 1–27.CrossRefGoogle Scholar
  78. Dale NG (1974) Bacteria in intertidal sediments: factors related to their distribution, Limnol. Oceanogr. 19, 509–516.Google Scholar
  79. Dauer DM and Simon JL (1975) Lateral or along-shore distribution of the polychaetons annelids of an intertidal sandy habitat, Mar. Biol. 31, 363–370.Google Scholar
  80. Davenport CB (1903) The animal ecology of the Cold Spring sand spit, with remarks on the theory of adaptation, Univ. Chicago Decennial Publ. 1G, 157–176.Google Scholar
  81. Davies JL (1972) Geographical variation in coastal development, London: Longman.Google Scholar
  82. Davies S (1981) Development and behaviour of little tern chicks, British Birds 74, 291–298.Google Scholar
  83. Davis N and Von Blaricom GR (1978) Spatial and temporal heterogeneity in a sand bottom epifaunal community of invertebrates in shallow water, Limnol. Oceanogr. 23, 417–427.Google Scholar
  84. Day JH, Field J and Montgomery MP (1971) The use of numerical methods to determine the distribution of the benthic fauna across the continental shelf of North Carolina. J.anim. Ecol. 40, 93–125.Google Scholar
  85. Dean HK (1981) Effects of the degree of wave action upon intertidal meiofaunal abundances, Am Zool. 21, 972.Google Scholar
  86. Delamare-Demoutteville C (1960) Biologie des eaux souterraines littorales et continentales, Paris: Hermann 740 pp.Google Scholar
  87. De Villiers G (1974) Growth, population dynamics, a mass mortality and arrangement of white sand mussels, Donax serra Röding, on beaches in the South-Western Cape Province, Invest. Rep. Div. Sea Fish. S. Afr. 109, 1–31.Google Scholar
  88. Dexter DM (1967) Distribution and niche diversity of haustoriid amphipods in North Carolina, Chesapeake, Sci. 8, 187–192.Google Scholar
  89. Dexter DM (1969) Structure of an intertidal sandy beach community in North Carolina, Chesapeake Sci. 10, 93–98.CrossRefGoogle Scholar
  90. Dexter DM (1971) Life history of the sandy beach amphopod Neohaustorius schmitzi (Crustacea, Haustoriidae), Mar. Biol. (Berl.) 8, 232–237.CrossRefGoogle Scholar
  91. Dexter DM (1972) Comparison of the community structures in a Pacific and an Atlantic Panamanian sandy beach, Bull. mar. Sci. 22, 449–462.Google Scholar
  92. Dexter DM (1974) Sandy beach fauna of the Pacific and Atlantic coasts of Costa Rica and Columbia, Rev. Biol. Trop. 22, 51–66.Google Scholar
  93. Dexter DM (1976) The sandy beach fauna of Mexico, Southwest Naturalist 20, 479–485.CrossRefGoogle Scholar
  94. Dexter DM (1977a) Natural history of the Pan-American sand beach isopod, Excirolana braziliensis (Crustacea: Malacostraca), J. Zool. Lond. 183, 103–109.CrossRefGoogle Scholar
  95. Dexter DM (1977b) A natural history of the sand dollar Encope stokesi L Agassiz in Panama, Bull. Mar. Sci. 27, 544–551.Google Scholar
  96. Dexter DM (1978) The infauna of a subtidal, sand-bottom community in Imperial Beach, California, Calif. Fish Game 64, 268–279.Google Scholar
  97. Dexter DM (1979) Community structure and seasonal variation in intertidal Panamanian sandy beaches, Estuar. cstl. mar. Sci. 9, 543–558.Google Scholar
  98. Dexter DM (1981) Intertidal fauna of Israeli and Sinai sandy beaches of the Mediterranean and Red Sea, Bull. Mar. Sci. 31, 812.Google Scholar
  99. Dexter DM (1983) Community structure of intertidal sandy beaches in New South Wales, Australia. In McLachlan A and Erasmus T, eds. Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  100. Dexter DM and Ebert TA (1975) A natural history study of Encope grandis and Mellita grantii, two sand dollars in the northern Gulf of California, Mexico, Mar. Biol. 32, 397–407.Google Scholar
  101. Dillery DG and Knapp LV (1970) Long-shore movements of the sand crab Emerita analoga (Decapoda, Hippidae). Crustaceana 18, 233–240.CrossRefGoogle Scholar
  102. Donn TE (1980) Population dynamics of the sandy beach amphipod Haustorius canadensis in Southern Maine U.S.A., Am. Zool. 20, 812.Google Scholar
  103. Du Preez HH (1981) The biology of the three-spot swimming crab, Ovalipes punctatus (de Haan) (Brachyura: Portunidae) with special reference to feeding. M.Sc. dissertation, University of Port Elizabeth.Google Scholar
  104. Dwivedi SN, Ayyappan S and Rahim A (1973) Ecology and production of intertidal macrofauna during monsoon in a sandy beach at Calangute, Goa, J. Mar. Biol. Ass. India 15, 274–284.Google Scholar
  105. Dye AH (1979a) Measurement of biological oxygen demand in sandy beaches, S.Afr. J. Zool. 14, 55–60.Google Scholar
  106. Dye AH (1979b) Quantitative estimation of Protozoa from sandy substrates, Estuar. cstl. mar. Sci. 8, 199–204.Google Scholar
  107. Dye AH (1980) Tidal fluctuations in biological oxygen demand in sandy beaches, Estuar. cstl. mar. Sci. 11, 1–8.Google Scholar
  108. Dye AH (1981) A study of benthic oxygen consumption on exposed sandy beaches, Estuar. cstl. Shelf. Sci. 13, 671–680.CrossRefGoogle Scholar
  109. Dye AH (1983) A synopsis of benthic oxygen consumption studies on sandy beaches. In McLachlan A and Erasmus T, eds. Sandy beaches as ecosystems, The Hague, Junk.Google Scholar
  110. Dye AH, McLachlan A and Wooldridge T (1981) The ecology of sandy beaches in Natal, South Africa, S. Afr. J. Zool. 16, 200–209.Google Scholar
  111. Edgren RE (1959) Coquinas (Donax variabilis) on a Florida Leach, Ecology 40, 498–502.CrossRefGoogle Scholar
  112. Edwards RRC (1973a) Production ecology of two Caribbean marine ecosystems. II. Metabolism and energy flow, Estuar. cstl. mar. Sci. 1, 319–333.Google Scholar
  113. Edwards RRC (1973b) Production ecology of two Caribbean marine ecosystems. 1 Physical environment and fauna, Estuar. cstl. mar. Sci. 1, 303–318.Google Scholar
  114. Edwards RRC and Steele JH (1969) The ecology of 0-group plaice and common dabs at Loch Ewe. I. Population and food, J. exp. mar. Biol. Ecol. 2, 215–238.CrossRefGoogle Scholar
  115. Edwards RRC, Steele JH and Trevallion A (1970) The ecology of 0-group plaice and common dabs at Loch Ewe. III. Prey-predator experiments with plaice, J. exp. mar. Biol. Ecol. 4, 156–173.Google Scholar
  116. Efford IE (1965) Aggregation in the sand crab Emerita analoga (Stimpson) J. Anim. Ecol. 34, 63–75.CrossRefGoogle Scholar
  117. Efford IE (1966) Feeding in the sand crab Emerita analoga (Decapoda, Hippidae) Crustaceana 10, 167–182.CrossRefGoogle Scholar
  118. Efford IE (1972) The distribution of the sand crabs, Hippa strigillata (Stimpson) and Hippa pacifica (Dana) in the Eastern Pacific Ocean (Decapoda, Anomura), Crustaceana 23, 119–122.CrossRefGoogle Scholar
  119. Efford IE (1976) Distribution of the sand crabs in the genus Emerita (Decapoda, Hippidae), Crustaceana 30, 169–183.CrossRefGoogle Scholar
  120. Eleftheriou A and Nicholson MD (1975) The effects of exposure on beach fauna, Cah. Biol. Mar. 16, 695–710.Google Scholar
  121. Eleftheriou A and McIntyre AD (1976) The intertidal fauna of sandy beaches–a survey of the Scottish Coast, Scottish Fisheries Research, Report No. 6. 1–61.Google Scholar
  122. Eleftheriou A and Jones DA (1976) The genus Erydice on the west coast of India, J. Zool. Lond. 178, 385–394.CrossRefGoogle Scholar
  123. Eleftheriou A, Holdich DM and Harrison K (1980) The systematics and ecology of a new genus of isopod (Sphaeromatidae) from the West coast sandy beaches of India, Est. cstl. mar. Sci. 11, 251–262.Google Scholar
  124. Eltringham SK (1971) Life in mud and sand, London, English Universities Press.Google Scholar
  125. Emberton KC (1981) Season depth relations in subtidal meiofauna of Cape Cod Bay, Estuaries 4, 121–126.CrossRefGoogle Scholar
  126. Enright JT (1963) The tidal rhythm of activity of a sand beach amphipod, Z. vergl. Physiol. 46, 276–313.Google Scholar
  127. Enright JT (1965) Entrainment of a tidal rhythm, Science 147, 864–867.PubMedCrossRefGoogle Scholar
  128. Enright JT (1972) A virtuoso isopod: circa-lumar rhythms and their tidal fine structure, J. comp. Physiol. 77, 141–162.CrossRefGoogle Scholar
  129. Epelde-Aguire A and Lopez MT (1975) Zonacion en el sustrato arenoso de Playa Blance, Bahia de Coronel y observaciones sobre crustaceos poco frecuentes, Biol. Soc. biol. de Concepcion 49, 161–170.Google Scholar
  130. Escofet A, Gianuca N, Maytia S and Scarabino V (1979) Playas arenosos de Atlantico Sudoccidental entre los 29°y 4J°LS: Consideraciones generates y esquena biocenologico, Unesco seminar, 9–12 May 1978, pp 245–258.Google Scholar
  131. Faubel A (1976) Population dynamics and life cycles of interstitial Acoela and Macrostomida (Turbellaria). Investigations of a middle lotic sandy beach of the Island of Sylt, Microfauna Meeresbodens 56, 1–107.Google Scholar
  132. Feder HM and Paul AJ (1980) Seasonal trends in meiofaunal abundance on two beaches in Port Valdaz, Alaska, Syesis 13, 27–36.Google Scholar
  133. Feller RJ (1980) Quantitative cohort analysis of a sand-dwelling meiobenthic harpacticoid copepod, Est. Cstl. mar. Sci. 11, 459–476.CrossRefGoogle Scholar
  134. Fellows DP (1975) On the distribution of the Hawaian ghost crab Ocypode laevis Dana, Pac, Sci. 29, 257–258.Google Scholar
  135. Fenchel T (1972) Aspects of decomposer food chains in marine benthos, Deutsche Zoologische Geselschaft Verhandlungen 65, 14–23.Google Scholar
  136. Fenchel TM (1978) The ecology of micro-and meiobenthos, Ann. Rev. Ecol. Syst. 9, 99–121.CrossRefGoogle Scholar
  137. Fenchel T and Jansson B-0 (1966) On the vertical distribution of the microfauna in the sediments of a brachish-water beach, Ophelia 3, 161–177.CrossRefGoogle Scholar
  138. Fenchel TM and Riedl RJ (1970) The sulfide system: a new biotic community under the oxidized layer of marine sand bottoms, Mar. Biol. 7, 255–268.Google Scholar
  139. Ferrara F (1974) Researches on the coast of Somalia. The shore and the dune of Sar Uanle 3: Terrestrial isopods, Monitore Zool. ital., Suppl 5, 191–220.Google Scholar
  140. Field JG (1971) A numerical analysis of changes in the soft-bottom fauna along a transect across False Bay, South Africa, J. Exp. Mar. Biol. Ecol. 7, 215–253.CrossRefGoogle Scholar
  141. Figueras A (1956) Moluscos de las playas de la Ria de Vigo. 1. Ecologia y distribution, Inv. Pesq. 5, 51–88.Google Scholar
  142. Fincham AA (1970) Amphipods in the surf plankton. J. mar. biol. Ass. U.K. 50, 177–198.CrossRefGoogle Scholar
  143. Fincham AA (1971) Ecology and population studies of some intertidal and sub-littoral sand-dwelling amphipods, J. mar. biol. Ass. U.K. 51, 471–488.CrossRefGoogle Scholar
  144. Fincham AA (1973) Rhythmic swimming behaviour of the New Zealand sand beach isopod Pseudaega punctata J. exp. mar. Biol. Ecol. 11, 229–237.CrossRefGoogle Scholar
  145. Fincham AA (1974) Intertidal sand-dwelling paracarid fauna of Stewart Island, N.Z. J. mar. Freshwat. Res. 8, 1–14.CrossRefGoogle Scholar
  146. Fincham AA (1977) Intertidal sand dwelling peracarid fauna of North Island, N.w Zealand, N.Z. J. mar. Freshwat. Res. 11, 677–696.CrossRefGoogle Scholar
  147. Fish S (1970) The biology of Eurydice pulcha (Crustacea: Isopoda), J. mar. biol. Ass. U.K. 50, 753–7585.CrossRefGoogle Scholar
  148. Fish JD and Fish S (1972) The swimming rhythm of Eurydice pulchra Leach and a possible explanation of intertidal migration, J. exp. mar. Biol. Ecol. 8, 195–200.CrossRefGoogle Scholar
  149. Fishelson L and Loya Y (1968) Preliminary observations on a population of Gastrosaccus sanctus (Mysidacea, Gastrosaccinae) on a Mediterranean sandy beach of Israel, Crustaceana 15, 149–152.CrossRefGoogle Scholar
  150. Fitch JE (1950) The pismo clam, Calif. Fish Game 36, 285–312.Google Scholar
  151. Fricke AH and Fleming B (1983) Selective microhabitat coolonisation by interstitial meiofauna as a function of grain size. ln McLachlan A and Erasmus T eds. Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  152. Fuchs E (1977) Predation and antipredator behaviour in a mixed colony of terns Sterna sp. and Black-headed gulls Larus ridibundus with special reference to the Sandwich tern Sterna sandvicensis Ornis Scand. 8, 17–32.Google Scholar
  153. Fusaro C (1980) Diel distribution differences in the sand crab, Emerita anagola (Simpson) (Decapoda, Hippidea), Crustaceana 39, 287–300.CrossRefGoogle Scholar
  154. Gauld DT and Buchanan JB (1956) The fauna of sandy beaches in the Gold Coast, Oikos 7, 293–301.CrossRefGoogle Scholar
  155. George RW and Knott ME (1965) The Ocypode ghost crabs of Western Australia (Crustacea, Brachyura), Trans. Roy. Soc. West. Austr. 48, 15–21.Google Scholar
  156. Gerlach SA (1971) On the importance of meiofauna for benthos communities. Oecologia (Berl) 6, 176–190.CrossRefGoogle Scholar
  157. Gianuca NM (1983) A preliminary account of the ecology of sandy beaches in Southern Brazil. In McLachlan A and Erasmus T, eds. Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  158. Gibson RN (1973) The intertidal movements and distribution of young fish on a sandy beach with special reference to the plaice Pleuronectes plateisa J. exp. mar. Biol. Ecol. 12, 79–102.CrossRefGoogle Scholar
  159. Giere 0 (1975) Population structure, food relations and ecological rôle of marine oligochaetes, with special reference to meiobenthic species, Mar. Biol. 31, 139–150.Google Scholar
  160. Giere 0 (1982) Habitat conditions and distribution pattern of the gutless oligochaete Phallodrillus leukodermatus Mar. Ecol. Prog. Ser. 8. 291–299.CrossRefGoogle Scholar
  161. Glynn PW, Dexter DM and Bowman TE (1975) Excirolona braziliensis and a Pan-American sand beach isopod: taxonomic status, zonation and distribution, J. Zool. Lond. 175, 509–521.Google Scholar
  162. Gnaiger E, Gluth G and Wieser W (1978), pH fluctuation in an intertidal beach in Bermuda, Limnol. Oceanogr. 23, 851–857.Google Scholar
  163. Gray JS (1971) The effects of pollution on sand meiofauna communities, Thalassia Jugoslovica 7, 79–86.Google Scholar
  164. Gray JS and Johnson RM (1970) The bacteria of a sandy beach as an ecological factor affecting the interstitial gastrotich Turbanella lyalina Schultse. J. exp. mar. Biol. Ecol. 4, 119–123.CrossRefGoogle Scholar
  165. Gray JS and Rieger RM (1971) A quantitative study of the meiofauna of an exposed sandy beach, at Robin Hood’s Bay, Yorkshire, J. Mar. Biol. Ass. U.K. 51, 1–19.CrossRefGoogle Scholar
  166. Griffiths CL and Stenton-Dozey J (1981) The fauna and rate of degradation of stranded kelp, Estuar. cstl shelf, Sci. 12, 645–653.Google Scholar
  167. Griffiths CL, Stenton-Dozey J and Koop K (1983) Kelp wrack and energy flow through a sandy beach. In McLachlan A and Erasmus T eds. Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  168. Grindley JR and Nel E (1968) Mussel poisoning and shellfish mortality on the west coast of South Africa. S. Afr. J. Sci. 64, 420–422.Google Scholar
  169. Gunter G (1958) Population studies of shallow water fishes of an outer beach in S. Texas, Texas Inst. Mar. S.i. Publ. 5, 186–193.Google Scholar
  170. Gunter G (1979) Notes on sea beach ecology. Food sources on sandy beaches and localized diatom blooms bordering gulf beaches, Gulf Res. Rep 6, 305–307.Google Scholar
  171. Gunter G and Lyles CH (1979) Localized plankton blooms and -jubilees on the Gulf Coast, Gulf Res. Rep. 6, 297–299.Google Scholar
  172. Hager RP and Croker RA (1980) The sand-burrowing amphipod Amphiporeia virginiana Shoemaker 1933 in the tidal plankton, Can. J. Zool. 58, 860–864.Google Scholar
  173. Hails AJ and Yaziz S (9182) Abundance, breeding and growth of the ocypodid crab Dotilla myotiroides (Milne-Edwards) on a west malaysian beach, Estuar. cstl shelf Sci. 15, 229–230.Google Scholar
  174. Haley SR (1982) Zonation by size of the pacific mole crab, Hippa pacifica Dana (Crustacea: Anomura: Hippidae), in Hawii, J. exp. mar. Biol. Ecol. 58, 221–231.Google Scholar
  175. Hall JR and Hessler RR (1971) Aspects in the population dynamics of Derocheilocoris typica (Mystacocorida, Crustacea), Vie Mileu 22, 305–326.Google Scholar
  176. Hammer WM, Smyth M and Mulford LD (1968) Orientation of the sand beach isopod Tylos punctatus Anim. Behay. 16, 405–409.CrossRefGoogle Scholar
  177. Hammer WM, Smyth M and Mulford ED (1969) The behaviour and life history of a sand-beach isopod Tylos punctatus Ecology 50, 442–453.Google Scholar
  178. Harris RP (1972a) Horizontal and vertical distribution of the intertidal harpacticoid colepods of a sandy beach, J. Mar. Biol. Ass. U.K. 52, 375–387.CrossRefGoogle Scholar
  179. Harris RP (1972b) Seasonal changes in the meiofauna population of an intertidal sand beach, J. Mar. Biol. Ass. U.K. 52, 382–403.Google Scholar
  180. Hartnoll RG (1983) The species area relationship on a sandy beach. In McLachlan A and Lrasmus T, eds. Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  181. Hartwig E (1982) Effects of temperature and oxygen on the distribution of Diophrys scutium (Ciliophora, Hypotrichida) on a temperate sand beach, an ecophysio-logical study. Cab. Biol. Mar. 23, 227–242.Google Scholar
  182. Hartwig E and Parker JG (1977) On the systematics and ecology of interstitial ciliates of sandy beaches in North Yorkshire, J. mar. biol. Ass. U.K. 57, 735–760.CrossRefGoogle Scholar
  183. Hartwig E, Gluth T and Wieser W (1977) Investigations on the ecophysiology of Geleia migriceps Kahl (Ciliophora, Gymnostomata) inhabiting a sandy beach in Bermuda, Oecologia 31, 159–175.Google Scholar
  184. Hayes WB (1974) Sand-beach energetics: importance of the isopod Tylos punctatus, Ecology 55, 838–837.CrossRefGoogle Scholar
  185. Hayes WB (1977) Factors affecting the distribution of Tylos punctatus (Isopoda, Oniscoideo) on beaches in Southern California and Northern Mexico, Pacific Science 31, 165–187.Google Scholar
  186. Hedgpeth JW (1957) Sandy beaches, In Hedgpeth JHW, ed. Treatise on Marine Ecology and Palaeociology Geol. Soc. America Memoir 67 1, 587–608.Google Scholar
  187. Hill GW (1982) Orientation of ghost crab Ocypode quadrata burrows as an indicator of shoreline position and wind direction. Tex. J. Sci. 34, 23–34.Google Scholar
  188. Hill GW and Hunter RE (1973) Burrows of the ghost crab Ocypode quadrata (Fabricius) on the barrier islands, south-central Texas coast, J. Sedim. Petrol. 43, 24–30.Google Scholar
  189. Hill GW and Hunter RE (1976) Interaction of biological and geological processes in the beach and nearshore environments, Northern Poole Island, Texas, In Davis and Ethrin, eds. Beach Nearshore Sedimentation. 169–187.Google Scholar
  190. Hockey PAR, Siegfried WR, Crowe AA and Cooper J (1983) Ecological structure and energy requirements of the sandy-beach avifauna of Southern Africa. In McLachlan A and Erasmus T, eds. Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  191. Hockin DC (1982) The effects of sediment particle diameter upon the meiobenthic copepod community of an intertidal beach. A field and laboratory experiment. J. Anim. Ecol. 51, 555–572.CrossRefGoogle Scholar
  192. Hogue EW and Miller CB (1981) Effects of sediment microtopography on small-scale spatial distributions of meiobenthic nematodes, J. exp. mar. Biol. Ecol. 181–191.Google Scholar
  193. Holanov SH and Hendrickson JR (1980) The relationship of sand moisture to burroN4ing depth of the sand beach isopod Tylos punctatus J. exp. mar. Biol. Ecol. 46, 81–88.CrossRefGoogle Scholar
  194. Holdich DM and Jones DA (1973) The systematics and ecology of a new genus of sand beach isopod (Sphaeromatidae) from Kenya, J. Zool. Lond. 171, 385–395.CrossRefGoogle Scholar
  195. Holland AF and Polgar TT (1976) Seasonal changes in the structure of an intertidal community, Mar. Biol. 37, 341–348.Google Scholar
  196. Holme NA (1954) The ecology of the British species of Ensis J. mar. biol. Ass. U.K. 33, 145–172.CrossRefGoogle Scholar
  197. Hulings NC (1971) A comparative study of the sand beach meiofauna of Lebanon, Tunisia and Morocco, Thalassia Jugosl. 7, 117–122.Google Scholar
  198. Hulings NC (1974) A temporal study of Lebanese sand beach meiofauna, Cah. Biol. Mar. 15, 319–335.Google Scholar
  199. Hulings NC and Gray JS (1971) A manual for the study of meiofauna, Smithsonian Contrib. Zool. 78, 1–84.Google Scholar
  200. Hulings NC and Gray JS (1976) Physical factors controlling abundance of meio fauna on tidal and atidal beaches, Mar. Biol. 34, 77–83.Google Scholar
  201. Hutchings L, Nelson G, Hortsman D and Tarr R (1983) Interactions between coastal plankton and sand mussels along the Cape coast, South Africa. In McLachlan and Erasmus T, eds. Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  202. Hummon WD, Fleeger JW and Hummon MR (1976) Meiofauna–macrofauna interaction 1. Sand beach meiofauna affected by maturing Limulus eggs, Chesapeake Sci. 17, 297–299.CrossRefGoogle Scholar
  203. Inman DL and Brush BM (1973) The Coastal challenge, Science 181, 20–32. Irwin TH (1973) The intertidal behaviour of the bean clam, Donax gouldii Dall 1921, Veliger 15, 206–212.Google Scholar
  204. Jansson BO (1967) The significance of grain size and pore water content for the interstitial fauna of sandy beaches, Oikos 18, 311–322.CrossRefGoogle Scholar
  205. Jansson BO (1968a) Quantitative and experimental studies of the interstitial fauna in four Swedish sandy beaches, Ophelia 5, 1–71.CrossRefGoogle Scholar
  206. Jansson BO (1968b) Studies on the ecology of the interstitial fauna of marine sandy beaches, Rep. Monogr. non Serials 16.Google Scholar
  207. Jaramillo E (1978) Zonacion estructura de la comunidad macrofaunistica en player de arena de Sur de Chile (Mehuin, Valdivia), Studies on Neotropical Fauna and Environment 13, 71–92.CrossRefGoogle Scholar
  208. Jaramillo E, Stotz W, Bertran C. Navarro J, Roman C and Varela C (1980) Actividad locomotrix de Orchestoidea turberculata (Amphipoda, Talitidae) sobre la superficie de una playa arenosa del sur de Chile ( Mehuin, provincia de Valdivia ), Studies on Neotropical Fauna and Environment 15, 9–33.Google Scholar
  209. Jijina JL and Lewin J (in prep, a) Persistent blooms of surf diatoms along the Pacific cost, USA. I. Physical characteristics of the coastal region in relation to the distribution and abundance of the species.Google Scholar
  210. Jijina JL and Lewin J (in prep, b) Persistent blooms of surf diatoms along the Pacific coast, USA. H. Use of regression analysis to assess the relationships between environmental variables and cell populations along Oregon and Washington beaches (1977 and 1978 ).Google Scholar
  211. Johannes RE (1980) The ecological significance of the submarine discharge of groundwater, Mar. Ecol. Prog. Ser. 3, 365–373.CrossRefGoogle Scholar
  212. Johnson WS (1976a) Biology and population dynamics of the intertidal isopod Cirolana harfordi Mar. Biol. 36, 343–350.Google Scholar
  213. Johnson WS (1976b) Population energetics of the intertidal isopod Cirolana harfordi Mar. Biol. 36, 351–357.Google Scholar
  214. Johnston, PT (1966) Mass mortality in a bivalve mollusc. Limnol. Oceanogr. 1, 429–431.Google Scholar
  215. Johnston PT (1968) Population crashes in the bean clam, Donax gouldi and their significance to the study of mass mortality on other marine invertebrates. J. Invest. Pathol. 12, 349–358.CrossRefGoogle Scholar
  216. Jones A (1973) The ecology of young turbot Scopthalmus maximus at Barth, Cardiganshire, Wales, J. Fish Biol. 5, 367–383.CrossRefGoogle Scholar
  217. Jones DA (1971) The systematics and ecology of some sand beach isopods (Crustacea, Eurydicidae) from the coast of Kenya, J. Zool. (Lond). 165, 201–227.CrossRefGoogle Scholar
  218. Jones DA (1972) Aspects of the ecology and behaviour of Ocypode ceratophthalmus (Pallas) and O. kuhlii (de Haan) (Crustacea, O.ypodidae), J. exp. mar. Biol. Ecol. 8, 31–43.CrossRefGoogle Scholar
  219. Jones DA (1974) The systematics and ecology of some sand beach isopods (family Cirolanidae) from the coasts of Saudi Arabia, Crustaceana 26, 202–211.CrossRefGoogle Scholar
  220. Jones DA (1979) The ecology of sandy beaches in Penang Malaysia with special reference to Excirolana orientalis Estuar. cstl. mar. Sci 9, 677–682.Google Scholar
  221. Joubert CSW (1981) The density of shore birds on a Natal sandy beach, Ostrich 52, 190–192.Google Scholar
  222. Kamihira Y (1979) Ecological studies of macrofauna on a sandy beach of Hakodate Japan. 2. Distribution of peracarids and the factors influencing their distribution, Bull. Fac. Fish, Hokkaido Univ. 30, 133–143.Google Scholar
  223. Kamihira Y (1981) Life history of sand burrowing amphipod Haustorioides japonicus (Crustacea, Dogielinotidae), Bull Fac. Fish. Hokkaido Univ. 32, 338–348.Google Scholar
  224. Kastendiek J (1982) Factors determining the distribution of the sea pansy, Renilla kollikeri in a subtidal sandbottom habitat, Oecologia 52, 340–347.CrossRefGoogle Scholar
  225. Kensley BF (1972) behavioural adaptations of the isopod Tylos granulatus Krauss, Zool. afr. 7, 1–4.Google Scholar
  226. Kensley BF (1974) Aspects of the biology and ecology of the genus Tylos Ann. S. Afr. Mus. 65, 401–471.Google Scholar
  227. Khiyama HM and Makemson JC (1973) Sand beach bacteria: numeration and characterization, Appl. Mircobiol. 26, 293–297.Google Scholar
  228. King CAM (1972) Beaches and coasts, London, Arnold.Google Scholar
  229. Klapow LA (1972) Fortnightly moulting and reproductive cycles in the sand beach isopod Lxcirolana chiltoni Biol. Bull. (Woods Hole) 143, 568–591.CrossRefGoogle Scholar
  230. Koepcke LW (1953) Contribucion al conocimiento de la forma de vida de Ocypode gaudichaudi Milne Edwards Y Lucas (Decapoda, Crust):, Publ. Mus. Hist. Nat. ‘Javier Prado’ ser A 13.Google Scholar
  231. Koepcke HW and Koepcke N (1952) Sobre el processo de transformacion de la materia organica en las playas arenosas del Peru, Revista de Ciencias Univ. Nac. San Marcos 54, 5–29.Google Scholar
  232. Koop K and Field JG (1980) The influence of food availability on population dynamics of a supralittoral isopod, Ligia dilatata Brandt, J. exp. mar. Biol. Ecol. 48, 61–72.CrossRefGoogle Scholar
  233. Koop K and Field JG (1981) Energy transformation by the supralittoral isopod Ligia dilatata Brandt, J. exp. mar. Biol. Ecol. 53, 221–233.CrossRefGoogle Scholar
  234. Koop K and Griffiths CL (1982) The relative significance of bacteria, meioand macrofauna on an exposed sandy beach, Mar. Biol. 66, 295–300.Google Scholar
  235. Koop K, Newell, RC and Lucas MJ (1982a) Biodegradation and carbon flow based on kelp debris (Ecklonia Maxima) in a sandy beach microcosm, Mar. Ecol. Prog. Ser. 7, 315–326.Google Scholar
  236. Koop K, Newell RC and Lucas MI (1982b) Microbial regeneration of nutrients from the decomposition of macrophyte debris on the shore, Mar. Ecol. Prog. Ser. 9, 91–96.Google Scholar
  237. Kuyken E (1978) Beached bird surveys in Belgium, Ibis 120, 122–123.Google Scholar
  238. Lasiak TA (1982) Structural and functional aspects of the surf-zone fish community in the Lastern Cape, Ph.D. thesis, University of Port Elizabeth.Google Scholar
  239. Lasiak TA (1983) The impact of surf zone fish communities on faunal assemblages associated with sand beaches. In McLachlan A and Erasmus T, eds. Sandy beaches as Ecosystems, The Hague, Junk.Google Scholar
  240. Lasker R, Wells JBJ and McIntyre AD (1980) Growth, reproduction, respiration and carbon utilization of the sand-dwelling harpacticoid copepod, Asellopsis intermedia J. mar. biol. Ass. U.K. 50, 147–160.Google Scholar
  241. Lassere p (1976) Metabolic activities of benthic microfauna and meiofauna. Recent advances and review of suitable methods of analysis, In New York, ‘The Benthic Bounday Layer’. ed. T.N. McCave Plemmum Press.Google Scholar
  242. Lawrence JM and Ferber J (1971) Substrate particle size and the occurrence of Lovenia elongata (Echinodermata: Echinoidea) at Taba, Gulf of Llat (Red Sea), Israel J. Zool, 20, 131–138.Google Scholar
  243. Leber KM (1982a) Seasonality of macroinvertebrates on a temperate, high wave energy sandy beach, Bull. Mar. Sci. 32, 86–98.Google Scholar
  244. Leber KM (1982b) Bivalves (Tellinacea: Donacidae) on a North Carolina beach: contrasting population size structures and tidal migrations, Mar. Ecol. Prog. Ser. 7, 297–301.CrossRefGoogle Scholar
  245. Lee JJ, Tietjen JH, Mastropaola C and Rubin H (1977) Food quality and the heterogeneous spatial distribution of meiofauna, Helgoländer wiss. Meeresunters. 30, 272–282.Google Scholar
  246. Lenanton RCJ, Robertson AI and Hansen JA (1982) Nearshore accumulations of detached macrophytes as nursery areas for fish, Mar. Ecol. Prog. Ser. 9, 51–57.Google Scholar
  247. Lewin J (1974) Blooms of surf zone diatoms along the coast of the olympic Peninsula, Washington. III. Changes in the species composition of the blooms since 1925, Nova Hedwigia 45, 251–256.Google Scholar
  248. Lewin J (1977) Persitant blooms of surf diatoms along the Northwest Coast, The Maring Plant. In R Krauss, ed. Biomass of the Pacific NW Coast, 81–92.Google Scholar
  249. Lewin J (1978) Blooms of surf-zone diatoms along the coast of the Olympic Peninsula, Washington IX: FactorsGoogle Scholar
  250. controlling the seasonal cycle of nitrate in the surf at Copalis Beach (1971 through 1975 ), Est. cstl. mar. Sci. 7, 173–183.Google Scholar
  251. Lewin J and Norris RE (1970) Surf-zone diatoms of the coasts of Washington and New Zealand (Chaetoceros armatum) T. West and Asterionella spp., Phycologia 9, 143–149.CrossRefGoogle Scholar
  252. Lewin J and Mackas D (1972) Blooms of surf-zone diatoms along the coast of the Olympic Peninsula, Washington. J. Physiological investigations of Chaetoceros armatum and Asterionella socialis in laboratory cultures, Mar. Biol. 16, 171–181.Google Scholar
  253. Lewin J and Hruby T (1973) Blooms of surf zone diatoms along the coast of the Olympic Peninsula, Washington. II. A diel periodicity in buoyancy shown by the surf zone diatom species, Chaetoceros armatium T. West, Estuar. cstl. mar. Sci. 1, 101–105.Google Scholar
  254. Lewin J and Rao VNR (1975) Blooms of surf-zone diatoms along the coast of the Olympic Peninsula, Washington VI: Daily periodicity phenomena associated with Chaeteceros armatum in its natural habitat, J. Phycol. 11, 330–338.Google Scholar
  255. Lewin J and Schaefer CT (1983) The rôle of phytoplankton in surf ecosystems. In McLachlan A and Erasmus T, eds. Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  256. Lewin J, Hruby T and Mackas D (1975) Blooms of the surf-zone diatoms along the coast of the Olympic Peninsula. Washington V: Environmental conditions associated with the blooms (1971–1972), Est. cstl mar. Sci. 3, 229–241.Google Scholar
  257. Lewin J, Chen C and Hruby T (1979a) Blooms of surf-zone diatoms along the coast of the Olympic Peninsula, Washington X: Chemical composition of the surf diatom Chaeterceros armatum and its major herbivore, the Pacific razor-clam, siliqua patula, Mar. Biol. 51, 259–265.Google Scholar
  258. Lewin J, Colvin JR and McDonald KL (1980) Blooms of surf-zone diatoms along the coast of the Olympic Peninsula, Washington. XII. The clay coast of Chaetoceros armatum T. West. Biol. Mar. 23, 333–341.Google Scholar
  259. Lewin J, Eckman JE and Ware GN (1979b) Blooms of surf zone diatoms along the coast of the Olympic Peninsula, Washington XI. Regeneration of ammonium in the surf environment by the Pacific razor clam, Siliqua patula Mar. Biol 52, 1–9.Google Scholar
  260. Lockwood SJ (1974) The settlement, distribution and movements of 0-group plaice Pleuronectes platessa (L.) in Filey Bay, Yorkshire, J. Fish Biol. 6, 465–477.CrossRefGoogle Scholar
  261. Loesch HC (1957) Studies of the ecology of two species of Donax on Mustang island, Texas, Texas Univ. Instit. Mar. Science Publ. 4, 201–227.Google Scholar
  262. Longbottom MR (1970) The distribution of Arenicola marina (L.) with particular reference to the effects of particle size and organic matter of sediments. J. exp. mar. Biol. Ecol. 5, 138–157.CrossRefGoogle Scholar
  263. Lopez-Cotelo J. Vieitez JM and DiazPineda F (1982) Tipos de comunidades bentonicas de la Playa de Puntal ( Bahia de Santander ), Cah. Biol. Mar. 23, 53–69.Google Scholar
  264. Macer CT (1967) The food webs in Red Wharf Bay (N. Wales) with particular reference to young plaice, Helgoländer wiss. Meeresunters. 15, 560–573.CrossRefGoogle Scholar
  265. Macquart-Moulin C (1977) Le contrôle de l’émergence et des rages nocturned chez les péracarides des plages de Méditerranée. Eurydice affinis Hansen (Isopoda), Gastrosaccus mediterraneus Bacescu, Gastrosaccus spinifer (Goes) (Mysidacea), J. exp. mar. Biol. Ecol. 27, 61–81.CrossRefGoogle Scholar
  266. Marsh BA and Branch GM (1979) Circadion and circatidal rythms of oxygen consumption in the sandy beach isopod Tyles yranulatus Krauss. J. exp. mar. Biol. Ecol. 37, 77–89.CrossRefGoogle Scholar
  267. Masse h (1963) Quelques donnees sur l’economie alimentaire d’une biocenose infralittorale. Rec. Tran. St. Mar. End. 31, 153–166.Google Scholar
  268. Masse H (1970) Contribution a l’etude de la macrofauna de peupléments des sables fins infralittoraux des Cotes de provence. I. La Baie de Bandol, Tethys 2, 783–820.Google Scholar
  269. Masse H (1971) Etude quantitative de la macrofaune de peupléments des sables fins infralittoraux II. La Baie du Prado (Golfe de Marseille), Tethys 3, 113–158.Google Scholar
  270. Masse H (1972) Quantitative investigations of sand-bottom macrofauna along the Mediterranean north-west coast, Mar. Biol. 15, 209–220.Google Scholar
  271. Matthews DC (1955) Feeding habits of the sand crab Hippa pacifica (Dana) Pac. Sci. 9, 382–386.Google Scholar
  272. Mazure HGF and Branch GM (1979) A preliminary analysis of bacterial numbers and biomass in Langebaan Lagoon, Trans, Roy. Soc. S. Afr. 44, 43–54.CrossRefGoogle Scholar
  273. McDermott J (1983) Food web in the surf zone of an exposed sandy beach along the mid-Atlantic coast of the United States. In McLachlan A and Erasmus T, eds. Sandy beaches as ecosystems, The Hague, Junk.Google Scholar
  274. McFarland WN (1963) Seasonal changes in the numbers and biomass of fishes from the surf at Mustang Is. Texas, Publ. Inst. Mar. Sci. 9, 91–105.Google Scholar
  275. McGwynne LE (1980) A comparative ecophysiological study of three sandy beach gastropods in the Eastern Cape, M.Sc. dissertation, University of Port Elizabeth.Google Scholar
  276. McIntyre AD (1968) The macrofauna and meiofauna of some tropical beaches, J. Zool. Lond. 156, 377–392.CrossRefGoogle Scholar
  277. McIntyre AD (1969) Ecology of marine meiobenthos, Biol. Rev. 44: 245–290.Google Scholar
  278. McIntyre AD (1970) The range of biomass in intertidal sand with special reference to the bivalve, Tellina tenuis J. mar. biol. Ass. U.K. 50, 561–575.CrossRefGoogle Scholar
  279. McIntyre AD (1971) Control factors on meiofauna populations, Thalassia Jugoslavia 7, 209–215.Google Scholar
  280. McIntyre AD (1977) Sandy forshores, In Barnes RSK, ed. The Coastline, 31–47.Google Scholar
  281. McIntyre AD and Eleftheriou A (1968) The bottom fauna of a flatfish nursery ground, J. mar. biol. Assn. U.K. 48, 113–142.CrossRefGoogle Scholar
  282. McIntyre AD, Munro ALS and Steele JH (1970) Energy flow in a sand ecosystem, In JH Steele, ed. Marine Food Chains, 19–31.Google Scholar
  283. McIntyre AD and Murison DJ (1973) The meiofauna of a flatfish nursery ground, J. mar. biol. Ass. U.K. 53, 93–118.CrossRefGoogle Scholar
  284. McIntyre RJ (1963) The supralittoral fringe of New Zealand sand beaches, Trans. roy. Soc. N.Z. Gen. 1, 89–105.Google Scholar
  285. McLachlan A (1977a) Composition, distribution, abundance and biomass of the marcofauna and meiofauna of four sandy beaches, Zool. Afr. 12, 279–306.Google Scholar
  286. McLachlan A (1977b) Studies on the psammolittoral meiofauna of Algoa Bay II. The distribution, composition and biomass of the meiofauna and macrofauna communities, Zool. Afr. 12, 33–60.Google Scholar
  287. McLachlan A (1977c) The larval development and population dynamics of Derocheilocaris algoensis (Crustacea: Mystacocarida) Zool. Afr. 12, 1–14.Google Scholar
  288. McLachlan A (1978) A quantitative analysis of the meiofauna and the chemistry of the redox potential discontinuity zone in a sheltered sandy beach, Estuar. cstl. mar. Sci. 7, 275–290.Google Scholar
  289. McLachlan A (1979a) Volumes of sea water filtered through Eastern Cape Sandy beaches, S.Afr. J. Sci. 75, 75–79.Google Scholar
  290. McLachlan A (1979b) Growth and production of Donax sordidus Hanley (Mollusca, Lamellibranchia) on an open sandy beach in Algoa Bay I., S.Afr. J. Zool. 14, 61–66.Google Scholar
  291. McLachlan A (1980a) The definition of sandy beaches in relation to exposure: a simple rating system, S.Afr. J. Sci. 76, 137–138.Google Scholar
  292. McLachlan A (1980b) Intertidal zonation of macrofauna and stratification of meiofauna on high energy sandy beaches in the Eastern Cape, South Africa, Trans. Roy. Soc. S.Afr. 44, 213–223.CrossRefGoogle Scholar
  293. McLachlan A (1980c) Occurrence of ghost crabs Ocypode spp., in the eastern Cape, S. Afr. J. Sci. 15, 57–58.Google Scholar
  294. McLachlan A (1980d) Exposed sandy beaches as semi-closed ecosystems, Mar. Environ. Res. 4, 59–63.Google Scholar
  295. McLachlan A (1982) A model for the estimation of water filtration and nutrient regeneration by exposed sandy beaches, Mar. environ. Res. 6, 37–47.Google Scholar
  296. McLachlan A (1983) The ecology of sandy beaches in the Eastern Cape, South Africa. In McLachlan A and Erasmus T, eds. Sandy beaches as Ecosystems, The Hague, Junk.Google Scholar
  297. McLachlan A, Erasmus T and Furstenberg JP (1977a) Migrations of sandy beach meiofauna, Zool. Afr. 12, 257–277.Google Scholar
  298. McLachlan A, Winter PED and Botha L (1977b) Vertical and horizontal distribution of sub-littoral meiofauna in Algoa Bay, South Africa, Mar. Biol. 40, 355–364.Google Scholar
  299. McLachlan A and Hanekom N (1979) Aspects of the biology, ecology, and seasonal fluctuations in biochemical composition of Donax serra in the East Cape, S. Afr. J. Zool. 14, 183–193.Google Scholar
  300. McLachlan A and van der Horst G (1979) Growth and production of two molluscs from an open sandy beach, S. Afr. J. Zool. 14, 194–210.Google Scholar
  301. McLachlan A, Dye AH and Van der Ryst P (1979e) Vertical gradients in the fauna and oxidation of two exposed sandy beaches, S.Afr. J. Zool. 14, 43–47.Google Scholar
  302. McLachlan A, Cooper C and van der Horst G (1979b) Growth and production of Bullia rhodostoma on an open sandy beach in Algoa Bay, S.Afr. J. Zool, 14, 49–53.Google Scholar
  303. McLachlan A, Wooldridge T and van der Horst G (1979c) Tidal movements of the macrofauna on an exposed sandy beach in South Africa, J. Zool. Lond. 188, 433–442.Google Scholar
  304. McLachlan A. Wooldridge T, Schramm M and Kühn M (1980) Seasonal abundance, biomass and feeding of shorebirds on sandy beaches in the Eastern Cape, South Africa, Ostrich 51, 44–52.Google Scholar
  305. McLachlan A, Erasmus T, Dye AH, Wooldridge T, van der Horst G, Rossouw G, Lasiak TA and McGwynne L (1981a) Sand beach energetics: an ecosystem approach towards a high energy interface, Estuar. cstl shelf Sci. 13, 11–25.CrossRefGoogle Scholar
  306. McLachlan A. Wooldridge T and Dye AH (1981b) The ecology of sandy beaches in Southern Africa, s. Afr. J. Zool. 16, 219–231.Google Scholar
  307. McLachlan A, Dye AH and Harty B (1981c) Simulation of the interstitial system of exposed sandy beaches, Estuar. cstl. shelf Sci. 12, 267–278.CrossRefGoogle Scholar
  308. McLachlan A and Lewin J (1981) Observation on surf phytoplankton blooms along the coasts of South Africa, Bot. Mar. 24, 553–557.Google Scholar
  309. McLachlan A and Young N (1982) Effects of low temperature on burrowing rates of four sandy beach molluscs. J. exp. mar. Biol. Ecol. 65, 275–284.CrossRefGoogle Scholar
  310. McLusky DS, Nair SA, Stirling A and Bharoava R (1975) The ecology of a Central West Indian beach with particular reference to Donax incarnatus, Mar. Biol. 30, 267–270.Google Scholar
  311. Meadows PS and Anderson JG (1966) Microorganisms attached to marine and freshwater sand grains, Nature 212, 1059–1060.CrossRefGoogle Scholar
  312. Meadows PS and Anderson JG (1968) Microorganisms attached to marine sand grains, J. mar. biol. Ass. U.K. 48, 161–175.CrossRefGoogle Scholar
  313. Meineke T and Westheide W (1979) Tide dependent migrations of the interstitial fauna in a sand beach of the Island of Sylt, North Sea, West Germany, Mikrofauna Meeresbodens 0 (75) 203–236.Google Scholar
  314. Meyer-Reil L-A and Faubel A (1980) Uptake of organic matter by meiofanua organisms and interralationships with bacteria, Mar. Ecol. Prog. Ser. 3Google Scholar
  315. Meadows PS and Anderson JG (1968) Microorganisms attached to marine sand grains, J. mar. biol. Ass. U.K. 48, 161–175.Google Scholar
  316. Meyer-Reil L-A, Dawson R. Liebezeit G and Tiedge H (1978) Fluctuations and intractions of bacterial activity in sandy beach sediments and overlying waters, Mar. Biol. 48, 161–171.Google Scholar
  317. Meyer-Reil L-A, Bolter M, Dawson R, Liebezeit G, Szwerinski H and Wolter K (1980) interrelationships between microbiological and chemical parameters of sandy beach sediments, a summer aspect, Appl. Env. Microbiol. 39, 797–802.Google Scholar
  318. Mielke W (1976) Copepod ecology of a sandy beach on the isle of Sylt West Germany, Mikrofauna Meeresbodens 59, 1–86.Google Scholar
  319. Mock B (1981) Diurodrilus subterraneus (Polychaeta: Dinophylidae) from sandy beaches of the North Sea island of Sylt, West Germany, Helgol. Wiss. Meeresunters. 34, 329–336.Google Scholar
  320. Moddle T and Ross ST (1981) Seasonality of fishes occupying a surf zone habitat in the northern Gulf of Mexico, Fish. Bull. U.S. 78, 911–922.Google Scholar
  321. Moore CG (1979a) The distribution and ecology of psammolittoral meiofauna around The Isle of Nan, Cah. de Biol. Mar. 20, 383–415.Google Scholar
  322. Moore CG (1979b) The zonation of psammolittoral harpacticoid copepods around The isle of Man, J. mar. biol. Ass. U.K. 59, 711–724.CrossRefGoogle Scholar
  323. Moran S (1972) Ecology of distribution of the sand-dwelling mysid Gastrosaccus sanctus (Van Beneden, 1961 ) along the Mediterranean sandy shore of Israel, Crustaceana Suppl. B., 357–361.Google Scholar
  324. Moran S and Fishelson L (1971) Predation of a sand-dwelling mysid crustacean Gatrosaccus sanctus by plover birds (Charadriidae), Mar. Biol. 9, 63–64.Google Scholar
  325. Morgans JFC (1962) The benthic ecology of False Bay. Part 2. Soft and rocky bottoms observed by diving and sampled by dredging, and the recognition of grounds, Trans. Roy. Soc. S.Afr. 36, 288–334.Google Scholar
  326. Mori S (1938) Donax semigranosus Dkr. and the experimental analysis of its behaviour at the flood tide, Zool. Mag. Tokyo 50, 1–12.Google Scholar
  327. Mouëza M and Chessel D (1976) Contribution a l’etude de la biologie de Donax trunculus L. (Mollusque: Lomellibranche) dans l’Algero’s: analyse statisque de la dispersion le long d’une plage en baie de bon Ismail, J. Exp. Mar. Biol. Ecol 21, 211–221.CrossRefGoogle Scholar
  328. Muir DG (1977) The biology of Talorchestia capensis (Amphipoda: Talitridae), including a population energy budget, M.Sc thesis, University 376 of Cape Town.Google Scholar
  329. Munro ALS and Brock TD (1968) Distinction between bacterial and algal utilization of soluble substances in the sea, J. gen. Microbiol. 51, 35–42.PubMedGoogle Scholar
  330. Munro ALS, Wells JBJ and McIntyre AD (1978) Energy flow in the flora and meiofauna of sandy beaches, Proc. Roy. Soc. Edinburgh 76B, 297–315.Google Scholar
  331. Myers JP, Williams SL and Pitelka FA (1980) An experimental analysis of prey availability for sanderlings (Ayes: Scolopacidae) feeding on sandy beach crustaceans, Can. J. Zool. 58, 1564–1574.Google Scholar
  332. Nair A (1978) Production ecology of a tropical sandy beach at Benaulim, Goa, Ind. J. Mar. Sci. 7, 163–167.Google Scholar
  333. New TR (1968) The dispersal of insects across sandy beaches with special reference to the larvae of Neuroptera, Entomol. gaz. 19, 119–125.Google Scholar
  334. Nodot C (1976) Biological cycles of some species of psammic harpacticoid copepods, Tethys 8, 241–248.Google Scholar
  335. Oliff WD, Gardner BD, Turner WD and Sharp JB (1970) The chemistry of the interstitial water as a measure of conditions in a sandy beach, Water Res. 4, 179–188.CrossRefGoogle Scholar
  336. Oliver JS, Darden JM and Slattery PN (1982) Phoxocephalid amphipod crustaceans as predators on larvae and juveniles in marine softbottom communities, Mar. Ecol. Prog. Ser. 7, 179–184.Google Scholar
  337. Orth RE, Moore J and Fisher TW (1977) Year round survey of Staphylinidae of a sandy beach in Southern California, Wasmann J. Biol. 35, 169–195.Google Scholar
  338. Orton JH (1929) Severe environmental mortality among Abra (=Syndosmya) alba, Donax vittatus and other organisms of the Lancashire coast. Nature Lond. 124, 911.CrossRefGoogle Scholar
  339. Pamatmat MM (1977) Benthic community metabolism: a review and assessment of present status and outlook, In B.C. Coull, ed. Ecology of Marine Benthos. University of South Carolina Press, 89–111.Google Scholar
  340. Panikkar BM and Rajan S (1970) Observations on the ecology of some sandy beaches of the southwest coast of India, Proc. Indian Acad. Sci. (B) 71, 247–260.Google Scholar
  341. Papi F and Pardi L (1963) On the lunar orientation of sandhoppers. Biol. Bull. 124, 97–105.CrossRefGoogle Scholar
  342. Pawlak R (1969) On the systematics and ecology, life cycles and population dynamics of the turbellarian genus Paromalöstomum. Investigations at the sandy beach of the North Sea Island Sylt, Helgolander wiss. Meeresunters 19, 417–454.CrossRefGoogle Scholar
  343. Pearse AS, Humm HJ and Wharton GW (1942) Ecology of sand beaches at Beaufort, N.C., Ecol. Monogr. 12, 135–190.Google Scholar
  344. Penchaszadeh PL (1971) Observaciones cuantitatives pre1iminares en playas arenosas de la coast central del Peru, con especial referencia a las poblaciones de Muy-Muy (Emerita analoga)–(Crustacea, Anomura, Hippidae), Contr. Inst. biol. mar. Mar. de Plata 177, 3–19.Google Scholar
  345. Penchaszadeh P (1983) Subtidal sandy beach trophic structure in the area of Punta Moron, Venezuela. In McLachlan A and Erasmus T, eds. Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  346. Penchaszadeh PE and Olivier SR (1975) Ecologia de una poblacion de “berberecho” (Donax hanleyanus) en Villa Gesell, Argentina, Malacologia 15, 133–146.Google Scholar
  347. Philip KP (1972) The inter-tidal fauna of the sandy beaches of Cochin, Proc. Ned. Sci. Acad. India, 38B, 317–328.Google Scholar
  348. Philips T (1970) Notes on the predation of the asteroid Luidia on Donax navicula at the Plaza Almejas, Baja California del Norte, Mexico, Tabulata 2, 16–18.Google Scholar
  349. Pichon M (1967) Contribution to the study of populations of tropical sandy beaches in the vicinity of Tulear, Madagascar, Recl. Tray. Sta. Mar. Endoume Suppl 7, 57–100.Google Scholar
  350. Pienkowski MW (1982) Diet and energy intake of grey and ringed plovers, Pluvialis squatarola and Charadius hiaticula in the non-breeding season, J. Zool. Lond. 197, 511–549.Google Scholar
  351. pike RB and Le Sueur RF (1958) The shore zonation of some Jersy Cumacea, Ann. May. Nat. Hist., Ser ‘13, 1, 515–523.Google Scholar
  352. Pollock LW and Hummon WD (1971) Cyclic changes in interstitial water content, atmospheric exposure and temperature in a marine beach, Limnol. Oceanogr. 16, 522–535.Google Scholar
  353. Poxton MG, Eleftheriou A and McIntyre AD (1982) The population dynamics of 0-group flatfish on nursery grounds in the Clyde Sea area, Estuar, cstl, shelf. Sci. 14, 265–282.Google Scholar
  354. Prater AH and Davies M (1978) Wintering sanderlings in Britain, Bird Studies 25, 33–38.CrossRefGoogle Scholar
  355. Pringle JS and Cooper J (1977) Wader populations (Charadrii) of the marine littoral of the Cape Peninsula, South Africa, Ostrich 48, 98–105.CrossRefGoogle Scholar
  356. Prosch R and McLachlan A (1983) Regeneration of surf zone nutrients by the sand mussel, Donax serra. In prep.Google Scholar
  357. Pugh KB (1976) An annual cycle, at constant temperature, in a model sandy beach. I Nutrient chemistry, J. exp. mar. Biol. Ecol. 22, 179–192.CrossRefGoogle Scholar
  358. Quinlan AV, Lewis T and Hoyt JK (1983) Fouling of the sandy beaches of Nahant Bay (Massachusetts, USA.) by an abnormal free-living form of the macroalgaGoogle Scholar
  359. Pilayella littoralis (Phaeophyta). I. Habitat characteristics. In McLachlan A and Erasmus T, eds, Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  360. Randall R and McLachlan A (1982) Damara terms Sterna balaenarum in breeding the Eastern Cape, South Africa, Ostrich 53, 50–51.CrossRefGoogle Scholar
  361. Rapson AM (1952) The Toheroa, amphidesma ventricosum Gray (Eulamellibranchiata), development and growth, Aust. J. Mar. Freshwat. Res. 3, 170–198.CrossRefGoogle Scholar
  362. Rapson AM (1954) Feeding and control of Toheroa (Amphidesma ventricosum Gray) (Eulamellibranchiata) populations in New Zealand, Aust. J. Mar. Freshwat. Res. 5, 486–512.Google Scholar
  363. Rees CB (1939) Notes on the ecology of the sandy beaches of North Donegal, Proc. roy. Irish Acad. 45, 215–229.Google Scholar
  364. Reilly FJ and Bellis VJ (1978) A study of the ecological impact of beach nourishment with dredged materials on the intertidal zone. U.S. Army Corps. Eng. Ins. Cstl. and Mar. Res. Tech. Rep 4, 107 pp.Google Scholar
  365. Reilly FJ (1979) Energy flow between biological systems on exposed sandy beaches, J. Elisha Mitchell Soc. 95, 125.Google Scholar
  366. Reilly FJ and Bellis VJ (1978) Trophic dynamics of sandy beach macrobenthos, Assoc. Southeast. Biol. Bull. 25, 34–35.Google Scholar
  367. Reise K (1979) Moderate predation on meiofauna by the macrobenthos of the Wadden Sea, Helgolander wiss. Meeresunters 32, 453–465.CrossRefGoogle Scholar
  368. Remane A (1933) Verteilung und Organisation der benthonischen Mikrofauna der Kieler Bucht, Wiss. Meeresunters. 21, 161–221.Google Scholar
  369. Renaud-Debyser J (1963) Recherches ecologiques sur la faune interstitielle des sables-Bassin d’Arcachon Ile de Bimini, Bahamas, Vie Milieu, Suppl. 15, 1–157.Google Scholar
  370. Rheinheimer G (1977) Investigations on the bacterial ecology in sand beaches of the North Sea and the Baltic, Bot. Mar. 20, 385–400.Google Scholar
  371. Rieper M (1978) Bacteria as food for marine harpacticoid copepods. Mar. Biol. 45, 337–345.CrossRefGoogle Scholar
  372. Riedl RJ (1971) How much seawater passes through sandy beaches ? Int. Rev. ges. Hydrobiol. 56, 923–946.CrossRefGoogle Scholar
  373. Riedl RJ and MacMahan E (1969) High energy beaches In Odum and Copeland, eds. Coastal Ecological Systems of the United States.Google Scholar
  374. Riedl RJ and Machan R (1972) Hydrodynamic patterns in lotic intertidal sands and their bio-climatological implications, Mar. Biol. 13, 179–209.Google Scholar
  375. Riedl RJ, Huang N and Machan R (1972) The subtidal pump: a mechanism of interstitial water exchange by wave action, Mar. Biol. 13, 210–221.Google Scholar
  376. Rieger R and Ott T (1971) Gezeitenbedingte Wanderungen von irubellarien und Nematoden eines Nordadriatischen Sandstrandes, Vie Milieu 22 (Suppl) 425–447.Google Scholar
  377. Riley JD and Corlett J (1566) The number of 0-group plaice in Port Grin Bay, 1964–1966. Dept. mar. biol. Stn. Port Erin, 78, 51–56.Google Scholar
  378. Robertson JR and Pfeiffer WJ (1981) Deposit feeding by the ghost crab Ocypode quadrata, J. exp. mar. Biol. Ecol. 56, 165–178.CrossRefGoogle Scholar
  379. Robertson S and Lewin J (1976) Blooms of surf-zone diatoms along the coast of the Olympic Peninsula, Washington. VII. Effect of temperature and oxygen concentration of respiration of Chaeteceras armatum in the natural environment, Int. Revue, ges. Hydrobiol. 61, 201–210.CrossRefGoogle Scholar
  380. Rossouw G (1983) The importance of non-teleost fishes in the surf zone with special reference to Rhinobatos Google Scholar
  381. annulatus. ln McLachlan A and Erasmus T, eds, Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  382. Round FE (1979) A diatom assemblage living below the surface of intertidal sand flats, Mar. Biol. 54, 219–223.Google Scholar
  383. Saloman CH and Naughton SP (1978) Benthic macroinvertebrates inhabiting the swash zone of Panama City Beach, Florida, Northeast Gulf Science 2, 65–72.Google Scholar
  384. Saloman CH, Naughton SP and Taylor JL (1982) Benthic community response to dredging borrow pits, Panama City Beach, Florida, U.S. Army Corps Engineers, Misc. Res. 82–3, 138 pp.Google Scholar
  385. Salvat B (1964) Les conditions hydrodynamiques interstitielles des sediment menbles intertidaux et la repartition verticale de la jenne endogee, C.R. 259, 1576–1579.Google Scholar
  386. Salvat B (1966) Eurydice pulchra Leach 1815 et Eurydice affinis Hansen 1905 (Isopodes: Cirolanidae). Taxonomie, ethologie, repartition verticale et cycle reproduction, Acta Soc. Limn. Bordeaux A 193, 1–77.Google Scholar
  387. Salvat B (1967) La macrofaune carinologique endogue des sediments meubles intertidaux (Tanaidaces, isopodes et Amphipodes): ethologie, bionomie et cycle biologique, Mem. Mus. natn. Hist. nat. Ser. A 45, 1–275.Google Scholar
  388. Sanchez M, Castilla JC and Mena 0 (1982) Summer and winter variations of the macrofauna of a sandy beach in Morrillas Norte-Chico, Chile (In Spanish) Studies on Neotropical Fauna and Environment, 17, 31–50.CrossRefGoogle Scholar
  389. Sameoto DD (1969a) Comparative ecology, life histories, and behaviour of intertidal sand-burrowing amphipods (Crustacea: Haustoriidae) at Cape Cod., J. Fish. Res. Bd. Canada 26, 361–388.CrossRefGoogle Scholar
  390. Sameoto DD (1969b) Some aspects of the ecology and life cycle of three species of subtidal sand-burrowing amphipods (Crustacea: Huastoriidae), J. Fish. Res. Bd. Canada 26, 1321–1345.CrossRefGoogle Scholar
  391. Sander F and Moore E (1978) A comparative study of inshore and offshore copepod populations at Barbados, West Indies, Crustaceana 35, 225–240.CrossRefGoogle Scholar
  392. Scapini F and Ugolini A (1981) Influence of landscape on the orientation of Talitus saltator (Crustacea, Amphipoda), Monit. Zool. Ital. 15, 324–325.Google Scholar
  393. Schaeffer RH (2967) Species composition, size and seasonal abundance of fish in the surf waters of Long Island, N.Y. Fish Game J. 1, 1–46.Google Scholar
  394. Schaeffer RH (1970) Feeding habits of striped bass (Morone saxatilis) from the surf waters of Long Island, N.Y. Fish Game J. 17, 1–17.Google Scholar
  395. Scheibling RE (1982) Habitat utilization and bioturbation by Oreaster reticulatus (Asteroidea) and Meona ventricosa (Echinoicea) in a subtidal sand patch. Bull. mar. Sci. 32, 624–629.Google Scholar
  396. Schmidt P (1970) Zonation of the interstitial polychaete Stygocapiletta subterranea (Stygocapitellidae) in European sandy beaches, Mar. Biol. (Berl.) 7, 319–323.CrossRefGoogle Scholar
  397. Schmidt P (1972a) Sonierung und jahreszeitliche fluktuationen de interstitiellen fauna in sandstränden des gebiets von Tromso (Norwegen), Mikrofauna Meeresbodens 12, 81–164.Google Scholar
  398. Schmidt P (1972b) Zonation and annual fluctuation of the mesopsammon in the sandy beach of Schilksee Bay of Kiel, West Germany, Mikrofauna Meeresbodens 10, 3–60.Google Scholar
  399. Schneider AC (1981) Size selective predation on mysids by birds (Plovers), Mar. Ecol. Prog. Ser. 5, 223–224.CrossRefGoogle Scholar
  400. Schneider D (1982) Predation by ruddy turnstones Arenaria interpres on a polymorphic clam Donax variabilis at Sanibel Island, Florida, Bull. Mar. Sci. 32, 341–344.Google Scholar
  401. Schneider DC and Harrington BA (1981) Timing of shorebird migration in relation to prey depletion. Auk, 98, 801–811.Google Scholar
  402. Schuster-Diederichs 0 (1956) Die macrofauna am sandingen Brandungsstrand von El Salvador, Senckenberg. biol. 37, 1–56.Google Scholar
  403. Schwinghamer P (1981) Characteristic size distributions of integral benthic communities, Can. J. Fish. Aquatic Sci. 38, 1255–1263.CrossRefGoogle Scholar
  404. Scott A (1960) The fauna of the sandy beach, Village Bay, St. Kilda: a dynamic relationship, Oikos 11, 153–160.CrossRefGoogle Scholar
  405. Scott KJ and Croker RA (1976) Macroinfauna of northern New England marine sand. III. The ecology of Psammonyx nobilis (Stimpson), 1853 (Crustacea: Amphipoda ), Can. J. Zool. 54, 1519–1529Google Scholar
  406. Seed R and Lowry BJ (1973) The intertidal macrofauna of seven sandy beaches of County Down, Proc. Roy. Irish Acad, B. 73, 217–230.Google Scholar
  407. Shelton CR and Robertson PB (1981) Community structure of intertidal macrofauna on two surf exposed Texas sandy beaches, Bull. Mar. Sci. 31, 833–842.Google Scholar
  408. Shin PKS (1981) Temporal variations in a shallow subtidal community in North Bay, Inner Galway Bay, West Coast of Ireland, Ir. Nat. J. 20, 321–324.Google Scholar
  409. Shin PKS (1982) Multiple discriminant anaylsis of macrobenthic infaunal assemblages, J. exp. mar. Biol. Ecol. 59, 39–50.CrossRefGoogle Scholar
  410. Smith DAS (1971) Polymorphism and population density in Donax rugosus (Lame11ibranchiata: Donacidae), J. Zool. Lond. 164, 429–441.CrossRefGoogle Scholar
  411. Sourie R (1957) Etude ecologique des playes de la eote Senegalaise aux environs de Dakar, Ann. Ec. sup. Sci. Dakar 3, 1–110.Google Scholar
  412. Southward AJ (1953) The fauna of some sandy and muddy shores in the south of the Isle of Man, Proc. Trans. Liverpool. biol. Soc. 59, 51–71.Google Scholar
  413. Steele 3H (1976) Comparative studies beaches, Phil. Trans. Roy. Soc. Edinburgh B 274, 401–415.CrossRefGoogle Scholar
  414. Steele JH and Baird JE (1968) Production ecology of a sandy beach, Limnol. Oceanogr. 13, 15–25.Google Scholar
  415. Steel JH and Edwards RRC (1970) The ecology of 0-group plaice and common dabs in Loch Ewe. IV. Dynamics of the plaice and dab populations, J. exp. mar. Biol. Ecol. 4, 174–187.CrossRefGoogle Scholar
  416. Steele JH, Munro ALS and Giese GS (1970a) Environmental factors controlling the epipsommial flora on beach and sub-littoral sands, J. mar. biol. Ass. U.K. 50, 907–918.Google Scholar
  417. Steele JH, McIntyre AD, Edwards RRC and Trevallion A (1970b) Interrelations of a young plaice population with its invertebrate food supply. In Watson, A., ed. Animal Populations in Relation to their food Resources. British Ecological Society Symposium 10, 375–388, Oxford, Blackwell.Google Scholar
  418. Stenton-Dozey J and Griffiths CL (1980) Growth, consumption and respiration by larvae of the kelp fly Fucellia capensis (Diptera: Anthomyiidae), S. Afr. J. Zool. 15, 280–283.Google Scholar
  419. Stenton-Dozey J and Griffiths CL (1983) The fauna accosiated with kelp stranded on a sandy beach. In McLachlan A and Erasmus T, eds. Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  420. Stephen AC (1928) Notes on the biology Tellina tenuis da Costa J. mar. biol. Ass. U.K. 15, 683–702.CrossRefGoogle Scholar
  421. Stephen AC (1931) Notes on the biology of certain lamellibranchs on the Scottish coast, J. mar. biol. Ass. U.K. 17, 277–300.CrossRefGoogle Scholar
  422. Stephen AC (1932) Notes on the biology of some lamellibranchs in the Clyde Sea area, J. mar. biol. Ass. U.K. 18, 51–68.CrossRefGoogle Scholar
  423. Subramoniam T (1979) Some aspects of reproductive ecology of a mole crab Emerita asiatica Milne Edwards, J. exp. mar. Biol. Ecol. 36, 259–268.CrossRefGoogle Scholar
  424. Summers RW and Cooper J (1977) The population ecology and conservation of the Black Oystercatcher Haematopus moquini Ostrich 48, 28–40.Google Scholar
  425. Summers RW, Cooper JZ and Pringle JS (1977) Distribution and numbers of coastal waders (Charadrii) in the south western Cape South Africa, summer 1975–76, Ostrich 48, 85–97.CrossRefGoogle Scholar
  426. Summers RW and Waltner M (1979) Seasonal variations in the mass of waders in southern Africa, with special reference to migration. Ostrich 50, 21–37.CrossRefGoogle Scholar
  427. Summers RW and Hockey PAR (1980) Breeding biology of the white-fronted plover Charadrius marginatus in the south western Cape, South Africa, J. Nat. Hist. 14, 433–446.CrossRefGoogle Scholar
  428. Swedmark B (1964) The interstitial fauna of marine sand, Biol. Rev. 39, 1–42.Google Scholar
  429. Thayer LA (1935) Diatom water-blooms on the coast of Washington. Proc. Louisiana Acad. Sci. 2, 67–71.Google Scholar
  430. Thijssen R, Lever AJ and Lever J (1974) Food composition and feeding periodicity of 0-group plaice Pleuronectes platessa in the tidal area of a sandy beach, Neth. J. Sea Res. 8, 369–377.Google Scholar
  431. Thomassin B (1969) Indentification, variabilité èt écologie des Hippidea (Crustacea, Anomura) de la règion de Tulear, S.W. de Madagascar, Rec. Tray. Stn. Mar. Endoume-Marseille Fasc. Aovs. Ser. Suppl. 9, 135–177.Google Scholar
  432. Thum AB and Allen JC (1975) Distribution and abundance of the lamp urchin Echinolampas crassa (Bell) 1980 in False Bay, Cape Trans. Roy. Soc. S.Afr. 41, 359–373.CrossRefGoogle Scholar
  433. Tietjen JH (1980) Microbial-meiofauna interrelationships: a review, Microbiology, 1980, 335–338.Google Scholar
  434. Trevallion A (1971) Studies on Tellina tenuis da Costa. III. Aspects of general biology and energy flow, J. exp. mar. Biol. Ecol. 7, 95–122.CrossRefGoogle Scholar
  435. Trevallion A, Ansell AD, Sivadas P and Narayanan B (1970) A preliminary account of two sandy beaches in South West India, Mar. Biol. 6, 268–279.Google Scholar
  436. Trueman LR (1971) The control of burrowing and the migratory behaviour of Donax denticulatus (Bivalvia: Tellinacea), J. Zool. Lond. 165, 453–469.CrossRefGoogle Scholar
  437. Turbeville DB and Marsh GA (1982) Benthic fauna of an offshore burrow area in Broward Country, Florida, U.S. Army Corps Engineers Misc. Reo. 82–1, 42 pp.Google Scholar
  438. Turner JH and Belding DL (1957) The tidal migrations of Donax variabilis (Say), Limnol. Oceanogr. 2, 120–125.Google Scholar
  439. Upadhyay RS, Singh DB and Bharat Rai (1978) Ecology of microfungi in a tropical coastal sand belt, Indian J. Mar. Sci. 7, 187–190.Google Scholar
  440. Vader W (1982) Pied wagtails catching young ghost crabs, Ostrich 53, 205.CrossRefGoogle Scholar
  441. Vannini M (1976) Research on the coast of Somalia. The shore and the dune of Sar Uanle. Part 10. Sandy beach decapods, Monit. Zool. ital. Suppl. 8, 255–286.Google Scholar
  442. Veitch CR (1980) Sea birds found dead in New Zealand in 1970, 1971 and 1972, Notornis 27, 369–385.Google Scholar
  443. Vernberg WB and Coull BC (1974) Respiration of an interstitial ciliate and benthic relationships, Oecologia (Berl.) 16, 259–264.CrossRefGoogle Scholar
  444. Virnstein RW (1977) The importance of predation by crabs and fishes on benthic infauna in Chesapeake Bay, Ecology 58, 1199–1217.CrossRefGoogle Scholar
  445. Vohra FC (1971) Zonation on a tropical sandy shore, J. Anim. Ecol. 40, 679–708.CrossRefGoogle Scholar
  446. Wade BA (1967) Studies on the biology of the West Indian beach clam Donax denticulatus Linne. 1. Ecology, Bull. mar. Sci. 17, 149–174.Google Scholar
  447. Wade BA (1968) Studies on the biology of the West Indian beach clam, Donax denticulatus (Linne) 2. Life-history, Bull. Mar. Sci. 18, 877–899.Google Scholar
  448. Wade K (1982) Wandering in Scopimera globosa (Crustacea, Ocypodidae) Publ. Seto. Mar. Biol. Lab. 26, 447–454.Google Scholar
  449. Watkin BE (1942) The macrofauna of intertidal sand at Karnes Bay, Millport, Buteshire, Trans. Roy, Soc. Edinburgh 60, 543–561.CrossRefGoogle Scholar
  450. Webb JE (2958) The ecology of Lagos Lagoon V: Some physical properties of Lagoon deposits, Phil. Trans. Roy. Soc. Lond. Ser B 241, 393–419.CrossRefGoogle Scholar
  451. Wenner AM (1977) Food supply, feeding habits, and egg production in Pacific mole crabs (Nippa pacifica Dana) Pac. Sci. 31, 39–47.Google Scholar
  452. Wenner AM and Fusaro C (1979) An analysis of population structure in Pacific mole crabs (Nippa pacifica Dana), Biol. Bull. 157, 205–220.CrossRefGoogle Scholar
  453. Wicks SR (1974) Presence of Acotobacter in marine sand beaches, Fla. Sci. 38, 167–169.Google Scholar
  454. Wieser W (1959) The effect of grain size on the distribution of small invertebrates inhabiting the beaches of Puget Sound, Limnol. Oceanogr. 4, 181–194.Google Scholar
  455. Wieser W (1975) The meiofauna as a tool in the study of habitat heterogeneity: ecophysiological aspects. A review, Cah. Biol. Mar. 16, 647–670.Google Scholar
  456. Wieser W (1960) Benthic studies in Buzzard’s Bay II: The meiofauna, Limnol. Oceanogr. 5, 121–137.Google Scholar
  457. Wieser W and Scheimer F (1977) The ecophysiology of some marine nematodes from Bermuda: Seasonal espects, J. exp. mar. Biol. Ecol. 26, 97–106.CrossRefGoogle Scholar
  458. Wieser W, Ott J, Schiemer F and Gnaiger E (1974) An ecophysiological study of some meiofauna species inhabiting a sandy beach at Bermuda, Mar. Biol. 26, 235–248.Google Scholar
  459. Wigley RL and Burns BR (1971) Distribution and biology of mysids (Crustacea, Mysidacea) from the Atlantic coast of the United States in the NMFS Woods Hole collection, Fish. Bull. 69, 717–746.Google Scholar
  460. Wilce RT and Quinlan AV (1983) Fouling of the sandy beaches of Nahant Bay (Massachusetts, USA) by an abnormal free-living form of the macroalga Pilayella littoralis (Phaeophyta) II. Population characteristics. In McLachlan A and Erasmus T, eds., Sandy Beaches as Ecosystems, The Hague, Junk.Google Scholar
  461. Wilce RT, Schneider ZW, Quinlan AV and Van den Bosch K (1982) The life history and morphology of free-living Pilayella littoralis (L.) Kjellm. (Ectocarpaceae, Ectocarpales) in Nahant Bay, Massachusetts, Phycologia 21, 336–354.Google Scholar
  462. Williamson DI (1951) On the mating and breeding of some semiterrestrial amphipods. Dept. Dove Marine Lab. 1949, 49–62.Google Scholar
  463. Wilson WH (1981) Sediment-mediated interactions in a densely populated infaunal assemblage: The effects of the polychaete Abarenicola pacifica. J. mar. Res. 39, 735–748.Google Scholar
  464. Withers RG (1977) Soft-shore macrobentos along the South West Coast of Wales, Estuar. cstl. mar. Sci. 5, 467–484.CrossRefGoogle Scholar
  465. Wolcott TG (1978) Ecological randle of ghost crabs Ocypode quadrata (Fabr.) on an ocean beach: scavengers or predators. J. exp. mar. Biol. Ecol. 31, 67–82.CrossRefGoogle Scholar
  466. Wood DH (1968) An ecological study of a sandy beach near Auckland, New Zealand, Trans. Roy. Soc. N.Z. Zool. 10, 89–115.Google Scholar
  467. Woodin SA (1981) Disturbance and community structure in a shallow water sand flat, Ecology 62, 1052–1066.CrossRefGoogle Scholar
  468. Wooldridge T (1981) Zonation and distribution of the beach mysid, Gastrosaccus psammodytes J. Zool. Lond. 193, 183–189.CrossRefGoogle Scholar
  469. Wooldridge T (1983) The ecology of beach and surf zone mysids in the Eastern Cape, South Africa. In McLachlan A and Erasmus T, eds. Sandy beaches as Ecosystems, The Hague, Junk.Google Scholar
  470. Wooldridge T, Dye Ali and McLachlan A (1981) The ecology of sandy beaches in Transkei, S. Afr. J. Zool. 16, 210–218.Google Scholar
  471. Wormald AP and Stirling HP (1979) A preliminary investigation of nutrient enrichment in experimental sand columns and its effect on tropical intertidal bacteria and meiofauna, Lst. cstl. mar. Sci. 8, 441–453.Google Scholar
  472. Wright LD and Short AD (1982) High-energy near shore and surf zone morphodynamics. JOA (Halifax) Abstracts p 86.Google Scholar
  473. Youngbluth MJ (1979) The variety and abundance of zooplankton in the coastal waters cf Puerto Rico, Northeast Gulf Sci. 3, 15–26.Google Scholar

Copyright information

© Springer Science+Business Media Dordrecht 1983

Authors and Affiliations

  • Anton McLachlan
    • 1
  1. 1.Zoology DepartmentUniversity of Port ElizabethPort ElizabethSouth Africa

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