• L. A. S. Johnson
  • K. L. Wilson
Part of the The Families and Genera of Vascular Plants book series (FAMILIES GENERA, volume 2)


Dioecious or monoecious trees or shrubs; bark persistent, smooth or furrowed or corky. Branchlets articulate, slender, with several short basal articles and one to many elongated articles; articles with as many ridges (phyllichnia) as there are teeth (reduced leaves), the phyllichnia separated by furrows (shallow and open in Gymnostoma, deep and closed in other genera) containing the stomates. Leaf laminae reduced to teeth in a whorl of 4–20 at apex of each article of assimilatory branchlets. Inflorescences of alternating whorls of tooth-like bracts; within each bract 2 lateral scale-like bracteoles, persistent (deciduous in some male Allocasuarina), and a single flower. Male inflorescence a short to elongated spike. Male flowers: tepals 1 or 2, anteroposterior, hooded, scale-like, deciduous at anthesis; stamen 1; anther 2-locular, basifixed. Female inflorescence a globular or ovoid head. Female flowers: perianth absent; carpels 2, anteroposterior, fused in the ovary and in the proximal part of the style; placentation axile (apparently basal at maturity); ovules 2, rarely 4, orthotropous, epitropous, bitegmic, crassinucellate; style 2-branched, reddish, proximally winged in fruit. Infructescence a ± woody compound “cone”, the 2 floral bracteoles enlarged as valves. Fruit a compressed, terminally winged nut (samara). Seed solitary; cotyledons large; radicle superior; often with more than 1 embryo; endosperm absent at maturity.


Fossil Pollen Female Inflorescence Male Inflorescence Ninetyeast Ridge Fossil History 
These keywords were added by machine and not by the authors. This process is experimental and the keywords may be updated as the learning algorithm improves.


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Copyright information

© Springer-Verlag Berlin Heidelberg 1993

Authors and Affiliations

  • L. A. S. Johnson
  • K. L. Wilson

There are no affiliations available

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