Intertidal and Subtidal Grazers

  • G. M. Branch
  • C. A. Moreno
Part of the Ecological Studies book series (ECOLSTUD, volume 103)


Grazing animals exert an important influence on the nature of intertidal and shallow subtidal biotic communities, because of their influence on algae. Removal of grazers often leads to a dramatic development of seaweed beds. In this chapter we begin by tracing the effects that human exploitation has on populations of grazers and on algae. These effects are usually obvious and easy to predict. There are, however, indirect effects that can arise from exploiting grazers, and these are often neither obvious nor easy to predict. This is particularly true in multispecies fisheries, and when effects flow from the exploitation of one species to other species, which seem only remotely connected with the target species. We conclude our account by making comparisons between the roles of grazers in Chile and South Africa, which emphasize the difficulty of extrapolating from one species to another and from one part of the world to another.


Rocky Shore Marine Reserve Rocky Intertidal Community Muricid Gastropod Chile Central 


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  1. Anderson AJ (1981) A model of prehistoric collecting on the rocky shore. J Archaeol Sci 8: 109–120CrossRefGoogle Scholar
  2. Barkai R, Griffiths CL (1988) An energy budget for the South African abalone Haliotis midae Linnaeus. J Molluscan Stud 54: 43–51CrossRefGoogle Scholar
  3. Berdichewsky B (1964) Arqueologia de la desembocadura del Aconcagua y zonas de la costa central de Chile. Arqueologia de Chile Central y Areas Vecinas. Tercer Congreso International de Arqueologia de Chile 12–15 de Mayo 1964, Vina del Mar Imprenta Los Andes, Santiago, pp 69–107Google Scholar
  4. Bertness MD, Cunningham C (1981) Crab shell-crushing predation and gastropod architectural defense. J Exp Mar Biol Ecol 50: 213–230CrossRefGoogle Scholar
  5. Black R (1977) Population regulation in the intertidal limpet Patelloida alticostata (Angus, 18651. Oecologia 30: 9–22CrossRefGoogle Scholar
  6. Black R (1979) Competition between intertidal limpets: an intrusive niche on a steep resource gradient. J Anim Ecol 48: 401–411CrossRefGoogle Scholar
  7. Bosman AL, Hockey PAR (1986) Seabird guano as a determinant of rocky intertidal community structure. Mar Ecol Prog Ser 32: 247–257CrossRefGoogle Scholar
  8. Bosman AL, Hockey PAR (1988) The influence of primary production rate on the population dynamics of Patella granularis, an intertidal limpet. PSZNI Mar Ecol 9: 181–198CrossRefGoogle Scholar
  9. Bosman AL, Hockey PAR, Siegfried WR (1987) The influence of coastal upwelling on the functional structure of rocky intertidal communities. Oecologia 72: 226–232CrossRefGoogle Scholar
  10. Branch GM (1974a) The ecology of Patella Linnaeus from the Cape Peninsula, South Africa. II. Reproductive cycles. Trans R Soc S Afr 41: 111–159CrossRefGoogle Scholar
  11. Branch GM (1974b) The ecology of Patella Linnaeus from the Cape Peninsula, South Africa. III. Growth rates. Trans R Soc S Afr 41: 161–193CrossRefGoogle Scholar
  12. Branch GM (1975a) Notes on the ecology of Patella concolor and Cellana capensis, and the effects of human consumption on limpet populations. Zool Afr 10: 75–85Google Scholar
  13. Branch GM (1975b) Intraspecific competition in Patella cochlear Born. J Anim Ecol 44: 263–282CrossRefGoogle Scholar
  14. Branch GM (1975c) Mechanisms reducing intraspecific competition in Patella species: migration, differentiation and territorial behaviour. J Anim Ecol 44: 575–600CrossRefGoogle Scholar
  15. Branch GM (1976) Interspecific competition experienced by South African Patella species. J Anim Ecol 45: 507–529CrossRefGoogle Scholar
  16. Branch GM (1981) The biology of limpets: physical factors, energy flow and ecological interactions. Oceanogr Mar Biol Annu Rev 19: 235–380Google Scholar
  17. Branch GM (1985) Limpets: their role in littoral and sublittoral community dynamics. In: Moore PG, Seed RA (eds) The ecology of rocky shores. Hodder and Stoughton Educational, London, pp 97–116Google Scholar
  18. Branch GM (1988) Activity rhythms of the pulmonate limpet Siphonaria thersites: interdependence between behaviour, morphology and biotic relationships. In: Vannini M, Chelazzi G (eds) Behavioural adaptations to intertidal life. Plenum Press, New York, pp 27–44Google Scholar
  19. Branch GM, Cherry MI (1985) Activity rhythms of the pulmonate limpet Siphonaria capensis Q & G as an adaptation to osmotic stress, predation and wave action. J Exp Mar Biol Ecol 87: 153–168CrossRefGoogle Scholar
  20. Branch GM, Barkai A, Hockey PAR, Hutchings L (1987) Biological interactions: causes or effects of variability in the Benguela ecosystem? S Afr J Mar Sci 5: 425–445CrossRefGoogle Scholar
  21. Branch GM, Harris JM, Parkins C, Bustamante RH, Eekhout S (1991) Algal “gardening” by grazers: a comparison of the ecological effects of territorial fish and limpets. In: John D, Hawkins SJ, Price J (eds) Plant-animal interactions in the marine benthos. Systematics Association Special Volume, vol XX, Clarendon Press, Oxford, pp 405–425Google Scholar
  22. Bretos M, Jiron C (1980) Trematodes in Chilean fissurellid molluscs. Veliger 22: 293Google Scholar
  23. Bretos M, Tesorieri I, Alvarez L (1983) The biology of Fissurella maxima Sowerby (Molluscs: Archaeogastropoda) in northern Chile. 2. Notes on its reproduction. Biol Bull (Woods Hole) 165: 559–568Google Scholar
  24. Brownell WN, Stevely JM (1981) The biology, fisheries and management of the queen conch, Strombus gigas. Mar Fish Rev 43: 1–12Google Scholar
  25. Bustamante RH, Castilla JC (1987) The shellfishery in Chile: an analysis of 26 years of landings (1960–1985). Biol Pesq, 16: 79–97Google Scholar
  26. Cancino J, Santelices B (1984) Importancia ecologica de los discos adhesivos de Lessonia nigrescens Bory ( Phaeophyta) en Chile central. Rev Chi’ Hist Nat 57: 23–33Google Scholar
  27. Capon RJ, Faulkner DJ (1984) Metabolites of the pulmonate Siphonaria lessoni. J Org Chem 49: 2506–2508CrossRefGoogle Scholar
  28. Castilla JC (1986) Sigue existiendo la necesidad de establecer parques y reservas maritimas en Chile? Ambiente Desarollo 11: 53–63Google Scholar
  29. Castilla, JC, Durân LR (1985) Human exclusion from the rocky intertidal zone of central Chile: the effects on Concholepas choncholepas ( Gastropoda ). Oikos 45: 391–399Google Scholar
  30. Castilla JC, Moreno CA (1982) Sea Urchins and Macrocystis pyrifera: experimental test of their ecological relations in southern Chile. In: Lawrence JM (ed) Echinoderms. Proc Int Conf, Tampa Bay. Balkema, Rotterdam, pp 257–263Google Scholar
  31. Castilla JC, Paine RT (1987) Predation and community organisation on eastern Pacific, temperate zone, rocky intertidal shores. Rev Chil Hist Nat 60: 131–151Google Scholar
  32. Catterall CP, Poiner (1987) The potential impact of human gathering on shellfish populations, with reference to some NE Australian intertidal flats. Oikos 50: 114–122Google Scholar
  33. Choat JH (1977) The influence of sessile organisms on the population biology of three species of acmaeid limpets. J Exp Mar Biol Ecol 26: 1–26CrossRefGoogle Scholar
  34. Connor MS (1975) Niche apportionment among the chitons Cyanoplax hartwegi and Mopalia muscosa and the limpets Collisella limatula and Collisella pelta under the brown alga Pelvetia fastigiata. Veliger 18 (Suppl): 9–17Google Scholar
  35. Conteras S, Castilla JC (1987) Feeding behaviour and morphological adaptations in two sympatric sea urchin species in central Chile. Mar Ecol Prog Ser 38: 217–224CrossRefGoogle Scholar
  36. Crawford RJM, Shannon LV, Pollock DE (1987) The Benguela ecosystem. Part IV. The major fish and invertebrate resources. Oceanogr Mar Biol Annu Rev 25: 353–505Google Scholar
  37. Creese RG (1980) An analysis of distribution and abundance of populations of the high-shore limpet, Notoacmea petterdi (Tenison-Woods). Oecologia 45: 252–260CrossRefGoogle Scholar
  38. Creese RG, Underwood AJ (1982) Analysis of inter-and intra-specific competition amongst intertidal limpets with different methods of feeding. Oecologia 53: 337–346CrossRefGoogle Scholar
  39. Dayton PK (1971) Competition, disturbance and community organisation: the provision and subsequent utilization of space in a rocky intertidal community. Ecol Monogr 41: 351–389CrossRefGoogle Scholar
  40. Dayton PK (1985a) Ecology of kelp communities. Annu Rev Ecol Syst 16: 215–245CrossRefGoogle Scholar
  41. Dayton PK (1985b) The structure and regulation of some South American kelp communities. Ecol Monogr 55: 447–468CrossRefGoogle Scholar
  42. Dethier MN (1981) Heteromorphic algal life histories: the seasonal pattern and response to herbivory of the brown crust Ralfsia californica. Oecologia 49: 333–339CrossRefGoogle Scholar
  43. Dethier MN, Duggins DO (1984) An “indirect commensalism” between marine herbivores and the importance of competitive hierarchies. Am Nat 124: 205–219CrossRefGoogle Scholar
  44. De Villiers G (1976) Observations on the potential exploitation of limpets along the western Cape coast. Fish Bull S Afr 8: 13–16Google Scholar
  45. Durân LR, Castilla JC, Oliva D (1987) Intensity of human predation on rocky shores at Los Cruces in central Chile. Environ Cons 14: 143–149CrossRefGoogle Scholar
  46. Eekhout S, Raubenheimer C, Branch GM, Bosman A, Bergh MO (1992) A holistic approach to the exploitation of virgin intertidal stocks: limpets as a case history. S Afr J Mar Sci 12: 1017–1029CrossRefGoogle Scholar
  47. Fletcher WJ (1984) Intraspecific variation in the population dynamics and growth of the limpet, Cellana tramoserica. Oecologia 63: 110–121CrossRefGoogle Scholar
  48. Geller JB (1991) Gastropod grazing and algal colonisation on a rocky shore in northern California: the importance of body size of grazers. J Exp Mar Biol Ecol 150: 1–17CrossRefGoogle Scholar
  49. George-Nascimento M, Quiroga G (1983) Una nueva especie de trematodo Proctoeces humboldti n. sp. (Digenea: Fellodistomidae), parasito de las lapas Fissurella spp. Brugiere, 1789 (Mollusca: Archeogastropoda ). Parasitol Día (Chile) 7: 100–103Google Scholar
  50. Godoy C, Moreno CA (1989) Indirect effects of human exclusion from the rocky intertidal in southern Chile: a case of cross-linkage between herbivores. Oikos 54: 101–106CrossRefGoogle Scholar
  51. Haven SB (1973) Competition for food between the intertidal gastropods Acmaea scabra and Acmaea digitalis. Ecology 54: 143–151CrossRefGoogle Scholar
  52. Hawkins SJ, Hartnoll RG (1983) Grazing of intertidal algae by marine invertebrates. Oceanogr Mar Biol Annu Rev 21: 195–282Google Scholar
  53. Hockey PAR (1987) The influence of coastal utilization by man on the presumed extinction of the Canarian black oystercatcher Haematopus meadewaldoi. Biol Consery 39: 49–62CrossRefGoogle Scholar
  54. Hockey PAR, Bosman A (1986) Man as an intertidal predator in Transkei: disturbance, com-munity convergence and management of a natural food resource. Oikos 46: 3–14CrossRefGoogle Scholar
  55. Hockey PAR, Branch GM (1984) Oystercatchers and limpets: impact and implications. A pre-liminary assessment. Ardua 72: 199–206Google Scholar
  56. Hockey PAR, Bosman AL, Siegfried WR (1988) Patterns and correlates of shellfish exploita-tion by coastal people in Transkei: an enigma of protein production. J Appl Ecol 25: 353–363CrossRefGoogle Scholar
  57. Jara HF, Moreno CA (1984) Herbivory and structure in a midlittoral rocky community: a case in southern Chile. Ecology 65: 28–38CrossRefGoogle Scholar
  58. Jones NS (1946) Browsing of Patella. Nature 158: 557CrossRefGoogle Scholar
  59. Kay EA, Magruder W (1977) The biology of Opihi, Department of Planning and Economic Development, Hawaii, 46 ppGoogle Scholar
  60. Kay EA, Corpuz GC, Magruder WH (1982) Opihi. Their biology and culture. Aquaculture Development Program, Department of Land and Natural Resources, Hawaii, 98 ppGoogle Scholar
  61. Leggett WC, Carscadden JE (1978) Latitudinal variation in reproductive characters of American shad (Alosa sapidissima): evidence for population specific life history strategies in fish. J Fish Res Board Can 35: 1469–1478CrossRefGoogle Scholar
  62. Lodge SM (1948) Algal growth in the absence of Patella on an experimentally cleared strip of foreshore, Port St. Mary, Isle of Man. Proc. Lpool Biol Soc 56: 78–83Google Scholar
  63. Lubchenco J (1978) Plant species diversity in a marine intertidal community: importance of herbivore food preferences and algal competitive abilities. Am Nat 112: 23–39CrossRefGoogle Scholar
  64. McClanahan TR (1989) Kenyan coral reef-associated gastropod fauna: a comparison between protected and unprotected reefs. Mar Ecol Prog Ser 53: 11–20CrossRefGoogle Scholar
  65. McClanahan TR, Muthiga NA (1988) Changes in Kenyan coral reef community structure due to exploitation. Hydrobiol 166: 269–276CrossRefGoogle Scholar
  66. McClanahan TR, Muthiga NA (1989) Patterns of predation on a sea urchin, Echinometra mathaei (de Blainville) on Kenyan coral reefs. J Exp Mar Biol Ecol 126: 77–94CrossRefGoogle Scholar
  67. McLachlan A, Lombard HW (1980) Growth and production in exploited and unexploited populations of a rocky shore gastropod, Turbo sarmaticus. Veliger 23: 221–230Google Scholar
  68. McLean JH (1984) Systematics of Fissurella in the Peruvian and Magellanic faunal provinces (Gastropoda: Prosobranchia). Contrib Sci Nat Hist Mus Los Angeles 345: 1–69Google Scholar
  69. Moreno CA (1986) Un resumen de las consecuencias de la exclusion del hombre en la zona intermareal de Mehuin—Chile. Estud Oceanol 5: 59–66Google Scholar
  70. Moreno CA, Jaramillo E (1983) The role of grazers in the zonation of intertidal macroalgae of the Chilean coast. Oikos 41: 73–76CrossRefGoogle Scholar
  71. Moreno CA, Vega R (1988) Valor cientifico de las reservas marinas costeras: un ejemplo de estudio ecologico con poblaciones intermareales de Loxechinus albus ( Molina ). Inf Unesco Cienc Mar 47: 124–134Google Scholar
  72. Moreno CA, Sutherland JP, Jara HJ (1984) Man as a predator in the intertidal zone of southern Chile. Oikos 42: 155–160CrossRefGoogle Scholar
  73. Moreno CA, Lunecke KM, Lepez MI (1986) The response of an intertidal Concholepas coneholepas ( Gastropoda) population to protection from man in southern Chile and the effects on benthic assemblages. Oikos 46: 359–364Google Scholar
  74. Moreno CA, Godoy C, Villouta E, Lepez I (1987) Explotación de recursos bénticos litorales: una alternitva derivada de la protection de areas. In: Arana P (ed) Manejo y Desarrollo Pesquero Escuela de Ciencias del Mar. Universidad Castolica de Valparaiso, pp 51–58Google Scholar
  75. Newell RC, Field JG, Griffiths CL (1982) Energy balance and significance of micro-organisms in a kelp bed community. Mar Ecol Prog Ser 8: 103–113CrossRefGoogle Scholar
  76. Newman GG (1967) Reproduction of the South African abalone Haliotis midae. Invest Rep Div Sea Fish S Afr 64: 1–24Google Scholar
  77. Newman GG (1968) Growth of the South African abalone Haliotis midae. Invest Rep Div Sea Fish S Afr 67: 1–24Google Scholar
  78. Oliva D, Castilla JC (1986) The effects of human exclusion on the population structure of keyhole limpets Fissurella crassa and F. limbata on the coast of central Chile. PSZNI Mar Ecol 7: 201–217CrossRefGoogle Scholar
  79. Ortega S (1985) Competitive interactions among tropical intertidal limpets. J Exp Mar Biol Ecol 90: 11–25CrossRefGoogle Scholar
  80. Ortega S (1987) The effect of human predation on the size distribution of Siphonaria gigas (Mollusca: Pulmonata) on the Pacific coast of Costa Rica. Veliger 29: 251–255Google Scholar
  81. Osorio C, Diaz H, Rios M (1986) Prevalencia de Proctoeces humboldti George-Nascimento Quiroga 1983 (Digenea: Fellodistomidae) en la poblacion de Fissurella maxima Sowerby 1835 (Mollusca, Gastropoda) de Los Vilos, Chile. Rev Bio Mar (Valparaiso) 22: 157–168Google Scholar
  82. Paine RT (1980) Food webs: linkage, interaction strength and community infrastructure. J Anim Ecol 9: 667–686Google Scholar
  83. Paine RT, Vadas RL (1969) The effects of grazing by sea urchins, Strongylocentrotus spp., on benthic algal populations. Limnol Oceanogr 14: 710–719CrossRefGoogle Scholar
  84. Paine RT, Castilla JC, Cancino J (1985) Perturbation and recovery patterns of starfish-dominated intertidal assemblages in Chile, New Zealand, and Washington State. Am Nat 125: 679–691Google Scholar
  85. Palmer AR (1979) Fish predation and the evolution of gastropod shell structure: experimental and geographic evidence. Evolution 33: 697–713CrossRefGoogle Scholar
  86. Palmer AR (1985) Adaptive value of shell variation in Thais lamellosa: effect of thick shells on vulnerability to and preference by crabs. Veliger 27: 349–356Google Scholar
  87. Parkington J (1976) Coastal settlement between the mouth of the Berg and Olifants Rivers, Cape Province. S Afr Archaeol Bull 31: 127–140CrossRefGoogle Scholar
  88. Parra B (1986) Analisis regional de la diversidad, abundancia y zonacion de las algas intermareales en un gradiente de intervencion antropica en al coasta de Valdivia Chile. MSc Thesis, Instituto de Botanica, Universidad Austral de ChileGoogle Scholar
  89. Poiner R, Catterall CP (1988) The effects of traditional gathering on populations of the marine gastropod Strombus luhuanus Linne 1758, in southern Papua New Guinea. Oecologia 76: 191–199CrossRefGoogle Scholar
  90. Prince JD, Sellers TL, Ford WB, Talbot SR (1987) Experimental evidence for limited dispersal of haliotid larvae (genus Haliotis; Mollusca: Gastropoda). J Exp Mar Biol Ecol 106: 243–263Google Scholar
  91. Prince JD, Sellers TL, Ford WB, Talbot SR (1988) Confirmation of a relationship between the localised abundance of breeding stock and recruitment for Haliotis rubra Leach ( Mollusca: Gastropoda). J Exp Mar Biol Ecol 122: 91–104Google Scholar
  92. Robson G (1986) A new species of South African limpet, Patella aphanes (Mollusca: Gastropoda: Prosobranchia), with a discussion of P. obtecta Krauss 1843. Durban Mus Novit 13: 305–324Google Scholar
  93. Santelices B (1987) South American marine herbivory. Rev Chil Hist Nat 60: 153–158Google Scholar
  94. Santelices B, Correa J (1985) Differential survival of macroalgae to digestion by intertidal herbivore molluscs. J Exp Mar Biol Ecol 88: 183–191CrossRefGoogle Scholar
  95. Santelices B, Ojeda FP (1984) Recruitment, growth and survival of Lessonia nigrescens ( Phaeo-phyta) at various tide levels in exposed habitats of central Chile. Mar Ecol Prog Ser 19: 73–82Google Scholar
  96. Santelices B, Ugarte R (1987) Algal life-history strategies and resistance to digestion. Mar Ecol Prog Ser 35: 267–275CrossRefGoogle Scholar
  97. Santelices B, Castilla JC, Cancino J, Schmiede P (1980) Comparative ecology of Lessonia ni-grescens and Durvillaea antarctica ( Phaeophyta) in central Chile. Mar Biol 59: 119–132Google Scholar
  98. Santelices B, Montalva S, Oliger P (1981) Competitive algal community organisation in ex-posed intertidal habitats from central Chile. Mar Ecol Prog Ser 6: 267–276CrossRefGoogle Scholar
  99. Santelices B, Correa J, Avila M (1983) Benthic algal spores surviving digestion by sea urchins. J Exp Mar Biol Ecol 70: 263–269CrossRefGoogle Scholar
  100. Santelices B, Vasquez J, Meneses I (1986) Patrones de distribution y diitas de un gremio de moluscos herbivoros en habitats intermareales expuestos de Chile central. Monogr Biol 4: 147–171Google Scholar
  101. Seeley RH (1986) Intense natural selection caused a rapid morphological transition in a living snail. Proc Natl Acad Sci 83: 6897–6901PubMedCrossRefGoogle Scholar
  102. Siegfried WR, Hockey PAR, Crowe AA (1985) Exploitation and conservation of brown mussel stocks by coastal people of Transkei. Environ Consery 39: 49–62Google Scholar
  103. Simenstad CA, Estes JA, Keyton KW (1978) Aleuts, sea otters, and alternative stable-state communities. Science 200: 403–411PubMedCrossRefGoogle Scholar
  104. Slocum CJ (1980) Differential susceptibility to grazers in two phases of an intertidal alga: advantages of heteromorphic generations. J Exp Mar Biol Ecol 46: 99–110CrossRefGoogle Scholar
  105. Southward AJ, Southward EC (1978) Recolonization of rocky shores in Cornwall after use of toxic dispersants to clean up Torrey Canyon Spill. J Fish Res Board Can 35: 682–706CrossRefGoogle Scholar
  106. Steneck RS (1982) A limpet-coralline alga association: adaptations and defenses between a selective herbivore and its prey. Ecology 63: 507–522CrossRefGoogle Scholar
  107. Steneck RS (1986) The ecology of coralline algal crusts: convergent patterns and adaptive strategies. Annu Rev Ecol Syst 17: 273–303CrossRefGoogle Scholar
  108. Sutherland JP, Ortega S (1986) Competition conditional on recruitment and temporary escape from predators on a tropical rocky shore. J Exp Mar Biol Ecol 95: 155–166CrossRefGoogle Scholar
  109. Swadling P (1976) Changes induced by human exploitation in prehistoric shellfish populations. Mankind Q 10: 156–162Google Scholar
  110. Swadling P (1977a) Central province shellfish resources and their utilisation in the prehistoric past of PNG. Veliger 19: 293–302Google Scholar
  111. Swadling P (1977b) The implications of shellfish exploitation for New Zealand prehistory. Mankind Q 11: 11–18Google Scholar
  112. Thompson GB (1980) Distribution and population dynamics of the limpet Patella vulgata L. in Bantry Bay. J Exp Mar Biol Ecol 45: 173–217Google Scholar
  113. Tutschulte TC, Connell JH (1988) Feeding behaviour and algal food of three species of abalones (Haliotis) in southern California. Mar Ecol Prog Ser 49: 57–64CrossRefGoogle Scholar
  114. Underwood Ai (1978) An experimental evaluation of competition between three species of intertidal prosobranch gastropods. Oecologia 33: 185–202CrossRefGoogle Scholar
  115. Underwood AJ (1980) The effects of grazing by gastropods and physical factors on the vertical limits of distribution of intertidal macroalgae. Oecologia 46: 201–213CrossRefGoogle Scholar
  116. Underwood AJ, Fairweather PG (1988) Supply-side ecology and benthic marine assemblages. TREE 4: 16–20Google Scholar
  117. Underwood AJ, Denley EJ, Moran MJ (1983) Experimental analyses of the structure and dynamics of midshore rocky intertidal communities in New South Wales. Oecologia 56: 202–219CrossRefGoogle Scholar
  118. Yesner DR (1984) Population pressure in coastal environments: an archaeological test. World Archaeol 16 (1): 108–127CrossRefGoogle Scholar
  119. Zamorano JH (1986) Rattus rattus (Rodentia, Muridae) un depredador intermareal poco conocido. Medio Ambiente 8:58–62Google Scholar

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© Springer-Verlag Berlin Heidelberg 1994

Authors and Affiliations

  • G. M. Branch
  • C. A. Moreno

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