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Wellstediaceae

Wellstediaceae (Pilg.) Novák in S. Prát (ed.), Rostlinopis 9: 530 (1943).Boraginaceae subfam. Wellstedioideae Pilg., Bot. Jahrb. Syst. 46: 558 (1912).
  • H. H. HilgerEmail author
  • M. Weigend
Chapter
Part of the The Families and Genera of Vascular Plants book series (FAMILIES GENERA, volume 14)

Abstract

Annual herbs, sometimes with basal rosette of large leaves, or more often subperennial to perennial shrublets; branches and leaves often more or less distichous and congested; stems lignified, strong taproot present; indumentum sericeous, of unicellular, densely scabrid trichomes, usually acroscopically appressed. Leaves alternate, exstipulate, shortly petiolate or subsessile, entire, linear to (ob)ovate-elliptical, narrowed at base, apex acute, acuminate or apiculate, indistinctly veined, sometimes primary vein visible below, rarely venation pinnate with ascending secondary veins. Inflorescences bracteose, of dense scorpioid monochasia or dichasia, rarely reduced to single flowers, morphologically terminal but overtopped by and sometimes partly fused with uppermost lateral shoot; when fused, flowers arranged in alternate rows on this shoot. Flowers erect, tetramerous, actinomorphic, bisexual, hypogynous; calyx divided nearly to base, calyx lobes equal, narrowly triangular to narrowly ovate or linear, pubescent or sericeous, slightly enlarging in fruit; corolla pink, white or yellowish, sympetalous, hypocrateriform with very short tube and half-erect to spreading lobes, tube inside with 4 protrusions often fused into a distinct rim, lobes triangular-ovate, margin entire or denticulate to erose; stamens 4, alternating with petal lobes, filaments equal, shorter or longer than anthers, anthers included or exserted; nectary unknown; ovary bicarpellate, style terminal with two very short stigmatic lobes. Fruit a bivalved capsule. Seeds one, rarely two, asymmetrically ovoid in lateral view, strongly laterally compressed, pubescent often with ring of longer hairs near funicular pole, exendospermous.

Keywords

Pollen Morphology Corolla Tube Secondary Vein Calyx Lobe Primary Vein 
These keywords were added by machine and not by the authors. This process is experimental and the keywords may be updated as the learning algorithm improves.

Selected Bibliography

  1. Balfour, I.B. 1888. Botany of Socotra. Edinburgh: R. Grant, pp. 1–446.Google Scholar
  2. Constance, L., Chuang, T.I. 1982. SEM survey of pollen morphology and classification in Hydrophyllaceae (waterleaf family). Amer. J. Bot. 69: 40–53.CrossRefGoogle Scholar
  3. Friedrich-Holzhammer, M. 1967. 121. Wellstediaceae. In: Merxmüller, H. (ed.) Prodromus einer Flora von Südwestafrika, Lieferung 13. Lehre: Cramer, p. 1Google Scholar
  4. Gürke, M. 1893. Borraginaceae (Asperifoliaceae). In: Engler, A., Prantl, K. (eds.) Die natürlichen Pflanzenfamilien, IV, 3a. Leipzig: Engelmann, pp. 71–131.Google Scholar
  5. Hunt, D.R. 1969. Wellstedia socotrana, W. dinteri var. dinteri, W. dinteri var. gracilior. Hook. Ic. Pl. 37: tab. 3665–3667.Google Scholar
  6. Novák, F.A. 1943. Wellstediaceae. Prát, Rostlinopsis 9: 530.Google Scholar
  7. Pilger, R. 1912. Die Gattung Wellstedia in Südwestafrika. Bot. Jahrb. Syst. 46: 558–561.Google Scholar
  8. Retief, W., van Wyk, A.E. 2008. The genus Wellstedia (Boraginaceae: Wellstedioideae) in southern Africa. Bothalia 38: 57–63.Google Scholar
  9. Retief, E., van Wyk, A.E., Van der Merwe, C.F. 2001.Tapetal orbicules in southern African Boraginaceae. Proc. Micros. Soc. Southern Africa 31: 56.Google Scholar
  10. Thulin, M. 2006. Boraginaceae. In: Thulin, M. (ed.) Flora of Somalia 3. Angiospermae (Ericaceae-Asteraceae). Kew: Royal Botanic Gardens, pp. 31–60.Google Scholar
  11. Thulin, M., Johansson, A.N.B. 1996. Taxonomy and biogeography of the anomalous genus Wellstedia. In: Van der Maesen, L.J.G., van der Burgt, X.M., van Medenbach de Rooy, J.M. (eds.) The Biodiversity of African Plants. Proc. XIV AETFAT Congr. Dordrecht: Kluwer, pp. 73–86.Google Scholar
  12. Weigend, M., Luebert, F., Selvi, F., Brokamp, G., Hilger, H.H. 2013. Multiple origins for Hounds tongues (Cynoglossum L.) and Navel seeds (Omphalodes Mill.) – the phylogeny of the borage family (Boraginaceae s.str.). Mol. Phylogenet. Evol. 68: 604–618.CrossRefPubMedGoogle Scholar
  13. Weigend, M., Luebert, F., Gottschling, M., Couvreur, T.L.P., Hilger, H.H., Miller, J. 2014. From capsules to nutlets—phylogenetic relationships in the Boraginales. Cladistics 30(5): 508–518.CrossRefGoogle Scholar

Copyright information

© Springer International Publishing Switzerland 2016

Authors and Affiliations

  1. 1.Systematische Botanik und Pflanzengeographie, Institut für BiologieFreie Universität BerlinBerlinGermany
  2. 2.Nees-Institut für Biodiversität der PflanzenRheinische Friedrich-Wilhelms-UniversitätBonnGermany

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