Helwingiaceae Decne, Ann. Sci. Nat. II, 6: 69 (1836).
  • Q.-Y. XiangEmail author
Part of the The Families and Genera of Vascular Plants book series (FAMILIES GENERA, volume 14)


Dioecious evergreen or deciduous shrubs, rarely small trees up to 8 m. Leaves simple, pinnately veined, alternate, petiolate and stipulate; blade chartaceous, sub-coriaceous or coriaceous; margin glandular-serrate; stipules 2, deciduous, thread-like, divided or not. Flowers small, 3–4-merous, rarely 5-merous, green or purple green; calyx small, limbs rudimentary; petals triangular to ovate, valvate; floral disk fleshy and flat; male flowers 3–20 in an umbel; stamen filaments short; anthers short, with two nearly separate locules; female flowers 1–4 in an umbel born on leaf blades, rarely on petioles; style short, with 3–4(5) distal stigmatic branchlets; ovary inferior, 3–4(5)-locular; ovules 1 per locule, pendant, apotropous, with dorsal raphe. Fruit drupaceous, with 3–4(5)-lobed stone covered by a fleshy layer, ovate or oblong, green when young, turning red and eventually black when mature, 1–4(5) obtusely angular when dry. Embryo straight.


Vessel Element Lateral Vein Axial Parenchyma Stamen Filament Quartet Move 
These keywords were added by machine and not by the authors. This process is experimental and the keywords may be updated as the learning algorithm improves.

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Copyright information

© Springer International Publishing Switzerland 2016

Authors and Affiliations

  1. 1.Department of Plant and Microbial BiologyNorth Carolina State UniversityRaleighUSA

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