Abstract
Studies in landscape ecology are concerned with determining the causes, consequences, and functional importance of spatial heterogeneity. Success in accomplishing these ambitious goals requires meaningful, robust methods for quantifying spatial pattern. Landscape ecologists use numerous metrics, and readily available spatial data combined with user-friendly software have made such analyses routine. However, the goal of landscape ecology is not simply to measure landscape pattern, and spatial pattern analysis is but one tool used to unravel the complex phenomena and relationships forming landscapes. Nevertheless, quantifying spatial heterogeneity is a key tool of the trade, and this chapter explains how to select, compute, and interpret landscape metrics. The development of new metrics has slowed (thankfully!), but useful new approaches that have emerged in the past decade are included here. While the nuts-and-bolts of individual metrics and software programs will continue to evolve over time, there are general caveats that apply to any analysis of landscape pattern. We begin this chapter by discussing why pattern is quantified and where landscape data come from, then highlight key caveats and cautions that must be considered before landscape pattern is analyzed. We next present commonly used landscape metrics of composition and configuration, introduce surface metrics briefly, and discuss connectivity measures derived from graph theory. Finally, we address the challenges associated with detecting significant differences in metrics and interpreting multiple pattern metrics, then proffer some added practical advice. This chapter largely focuses on pattern analysis based on categorical data, and we cover spatial statistics in the next chapter.
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Further Reading
Further Reading
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Burnicki AC (2012) Impact of error on landscape pattern analyses performed on land-cover change maps. Landsc Ecol 27:713–729
-
Cushman SA, McGarigal K, Neel MC (2008) Parsimony in landscape metrics: strength, universality and consistency. Ecol Indic 8:691–703
-
Eigenbrod F, Hecnar SJ, Fahrig L (2011) Sub-optimal study design has major impacts on landscape-scale inference. Biol Conserv 144:298–305
-
Fletcher RJ, Acevedo MA, Reichert BE, Pias KE, Kitchens WM (2011) Social network models predict movement and connectivity in ecological landscapes. Proc Natl Acad Sci USA 108:19282–19287
-
Gustafson EJ (1998) Quantifying landscape spatial pattern: what is the state of the art? Ecosystems 1:143–156
-
Kupfer JA (2012) Landscape ecology and biogeography: rethinking landscape metrics in a post-FRAGSTATS landscape. Prog Phys Geogr 36:400–420
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McGarigal K (2014) FRAGSTATS Help. Documentation for FRAGSTATS 4.0. University of Massachusetts, Amherst. http://www.umass.edu/landeco/research/FRAGSTATS/documents/FRAGSTATS_documents.html
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Uuemaa E, Antrop M, Roosaare J, Marja R, Mander Ü (2009) Landscape metrics and indices: an overview of their use in landscape research. Living Rev Landsc Res 3:1–28
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Turner, M.G., Gardner, R.H. (2015). Landscape Metrics. In: Landscape Ecology in Theory and Practice. Springer, New York, NY. https://doi.org/10.1007/978-1-4939-2794-4_4
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