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The Genus Ringicula Deshayes, 1848 (Ringiculidae, Gastropoda) in the Miocene of the Eastern Paratethys

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Abstract

This paper summarizes data on the distribution of the genus Ringicula in the Miocene sediments of the Eastern Paratethys. Two invasions of the genus from the adjacent waters of the Central Paratethys and the Eastern Mediterranean have been identified: in the Tarkhanian and in the Konkian. The species R. costata penetrated into the Eastern Paratethys in the Tarkhanian and produced several distinct forms of different systematic ranks. Tarkhanian specimens of R. costata belong to the subspecies paratethica subsp. nov., which gave rise the subspecies R. costata chokrakensis subsp. nov., inhabiting shallow areas of the sea with high energy depositional settings. Deeper areas with siltstone-clay deposition were inhabited by the species R. subglobosa sp. nov. By the end of the early Chokrakian, Ringicula species had disappeared due to the freshening of the basin. During the Konkian time, two species of the genus (R. ventricosa and R. laevigata) arrived in the basin. They became extinct without significant changes by the beginning of the Sarmatian.

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Notes

  1. The authors indicated only a single syntype of R. paulucciae from the collections of the museum (Muséum National d’Histoire Naturelle, Paris). However, for R. tournoueri Morlet, 1878 discussed below, only one syntype is also indicated, although the museum website mentions three shells (https://science.mnhn.fr/all/search). A similar situation is known for R. quadriplicata Morlet, 1883: the paper points out one syntype, and two are recorded in the museummuseum site. Moreover, both records of shells are accompanied by photographs, and one of them corresponds to the shell illustrated in the original paper.

  2. Ringicula with four folds is well illustrated in the type material of the species R. quadriplicata Morlet, 1878 (https://science.mnhn.fr/institution/mnhn/collection/f/item/a58584). It is visible that due to the appearance of a furrow below the well-developed parietal fold, the “fourth fold” begins to separate. Probably, in some shells this phenomenon is better developed. The photographs clearly show how the ridge-like thickening of the callus obscures the parietal fold, if observed directly in the apertural view.

  3. This formation is attributed to the lower Badenian, which correlates approximately with the middle part of the Langhian (Harzhauser et al., 2003).

  4. As shown later (Ferrero Mortara et al., 1981), the fauna recorded by F. Sacco for the Colli torinesi and dated as Helvetian, which he placed between Langhian and Tortonian, in fact comes from the interval extending from Aquitanian to Serravallian. A more accurate dating of this material is usually impossible.

  5. Classification is given in Guzhov (2021).

  6. Specimens identified by Iljina as R. costata from Geokoba (2 specimens) and Bryk (1 specimen) are not sufficiently preserved for identification. The Bryk specimen may also be a poorly preserved specimen of R. subglobosa. The R. costata shell from the Konkian of Mount Dubrovaya is probably a rounded spire of R. ventricosa.

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ACKNOWLEDGMENTS

The author is grateful to Jaap Vandervoort and Andre Jansen (The Netherlands) for their help with the literature.

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Guzhov, A.V. The Genus Ringicula Deshayes, 1848 (Ringiculidae, Gastropoda) in the Miocene of the Eastern Paratethys. Paleontol. J. 56, 1114–1141 (2022). https://doi.org/10.1134/S0031030122100033

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