Abstract
A new family, Ivapteridae fam. nov. (Insecta; Grylloblattida), is described from the Middle Permian locality of Soyana (Arkhangelsk Region; Kazanian Stage). It is most similar to Sojanoraphidiidae O. Martynova, 1952, differing from it in the subcostal field being traversed in the basal half of the wing by long, curved, and strongly oblique crossveins that form a double row of cells, the base of CuA being free, and CuA1 thin compared to CuA2. The new family is represented by a single species, Ivaptera sharovi, gen. et sp. nov. An overview of the modern system of the order Grylloblattida is included.
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References
D. S. Aristov, “A New Family of Early Permian Grylloblattids (Insecta: Grylloblattida) from the Ural Moutains,” Far Eastern Entomol., No. 85, 1–4 (2000).
D. S. Aristov, “The Fauna of Grylloblattid Insects (Grylloblattida) of the Lower Permian Locality of Tshekarda,” Paleontol. J. 38(Suppl. 2), 80–145 (2004a).
D. S. Aristov, “Grylloblattids of the Family Chaulioditidae (= Tomiidae syn. nov.) (Insecta: Grylloblattida) from the Upper Permian of the Orenburg Region,” Paleontol. J. 38(Suppl. 2), 146–149 (2004b).
D. S. Aristov, “The Fauna of Grylloblattid Insects (Grylloblattida) from the End of the Late Permian to the First Half of the Triassic,” Paleontol. Zh., No. 5, 45–62 (2004) [Paleontol. J. 38 (5), 514–521 (2004c)].
O. Béthoux, “Emptying the Paleozoic Wastebasket for Insects Members of a Carboniferous ‘Protorthopterous Family’ Assigned to Natural Groups,” Alavesia 1, 41–48 (2007).
O. Béthoux, A. Nel, J. Lapeyrie, and G. Gand, “New Data on Paleozoic Grylloblattid Insects (Neoptera),” J. Paleontol. 79(1), 125–138 (2005).
F. M. Carpenter, Treatise on Invertebrate Paleontology, Vol. 3 Superclass Hexapoda Boulder (Geol. Soc. Am. Univ. Kansas Press, Lawrence, 1992).
A. Handlirsch, Die Fossilen Insecten und die Phylogenie der rezenten Formen: Ein Handbuch für Paläontologen und Zoologen (Engelmann, Leipzig, 1906–1908).
V. G. Novokshonov, “Novyi Barmalei from the Upper Permian of the Arkhangelsk Region (Insecta; Gryllobattida: Tillyardembiidae),” Vestn. Perm. Univ., Ser. Biol., No. 3, 202–205 (1997).
V. G. Novokshonov, “New Extinct Insects (Insecta: Grylloblattida, Ordinis incertis) from the Lower Permian of the Middle Ural Mountains,” Paleontol. Zh., No. 5, 42–47 (2000) [Paleontol. J. 34 (5), 511–516 (2000)].
V. G. Novokshonov and D. S. Aristov, “New and Little- Known Permian Insects (Insecta: Grylloblattida; Orthoptera) from the Chekarda Locality, Central Ural Mountains),” Paleontol. Zh., No. 6, 73–77 (2002) [Paleontol. J. 36 (6), 648–652 (2002)].
V. G. Novokshonov, V. V. Ivanov, and D. S. Aristov, “New Insects from the Late Permian of the Ural Mountains,” Paleontol. Zh., No. 2, 39–42 (2002) [Paleontol. J. 36 (2), 157–160 (2002)].
V. G. Novokshonov, V. V. Ivanov, and D. S. Aristov, “New Insects from the Late Permian of the Ural Mountains,” Paleontol. Zh., No. 2, 39–42 (2002) [Paleontol. J. 36 (2), 157–160 (2002)].
V. G. Novokshonov and S. Yu. Storozhenko, “New Genus of the Family Sojanoraphidiidae (Grylloblattida) from Lower Permian of Ural,” Far Eastern Entomol., No. 33, 1–4 (1996).
A. P. Rasnitsyn, “The Origin and Volume of the Class Insecta: Cohort Paoliiformes, Infraclass Gryllones, Superorder Blattidea, Order Eoblattida, Superorder Perlidea, Order Grylloblattida, Order Perlida, Order Gerarida, Superorder Gryllidea,” Tr. Paleontol. Inst. Akad. Nauk SSSR 178 (Historical Development of the Class Insecta), 19–20, 29–30, 134–138, 148–160, 164–166 (1980).
A. P. Rasnitsyn, I. D. Sukacheva, and D. S. Aristov, “Permian Insects of the Vorkuta Group in the Pechora Basin, and Their Stratigraphic Implications,” Paleontol. Zh., No. 4, 63–75 (2005) [Paleontol. J. 39 (4), 404–416 (2005)].
A. V. Sharov, “Paleozoic Insects of the Kuznetsk Basin: The Orders Protoblattodea and Paraplecoptera,” Tr. Paleontol. Inst. Akad. Nauk SSSR 85, 157–234 (1961).
A. V. Sharov, “The Orders Protoblattodea and Paraplecoptera,” in Fundamentals of Paleontology: Arthropods, Tracheates, and Chelicerates (Akad. Nauk SSSR, Moscow, 1962), pp. 116–134 [in Russian].
S. Yu. Storozhenko, “Classification and Phylogeny of the Order Grylloblattida (Insecta),” in Lectures in Memory of Aleksei Ivanovich Kurentsov (Dal’nauka, Vladivostok, 1997a), Vol. 7, pp. 21–56 [in Russian].
S. Yu. Storozhenko, “Classification of the Order Grylloblattida (Insecta), with Description of New Taxa,” Far Eastern Entomol., No. 42, 1–20 (1997b).
S. Yu. Storozhenko, Systematics, Phylogeny, and Evolution of Grylloblattid Insects (Insecta: Grylloblattida) (Dal’nauka, Vladivostok, 1998) [in Russian].
S. Yu. Storozhenko, “Order Grylloblattida Walker, 1914,” in History of Insects (Kluwer, Dordrecht, 2002), pp. 278–281.
S. Yu. Storozhenko and V. G. Novokshonov, “To Knowledge of the Fossil Family Permosialidae (Insecta; Miomoptera),” Far Eastern Entomol., No. 76, 1–5 (1999).
R. J. Tillyard, “Kansas Permian Insects: Part 10. The New Order Protoperlaria: A Study of Typical Genus Lemmatophora Sellards,” Am. J. Sci. 16, 185–220 (1928).
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Original Russian Text © D.S. Aristov, 2009, published in Paleontologicheskii Zhurnal, 2009, No. 2, pp. 59–63.
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Aristov, D.S. A new family of the order Grylloblattida (Insecta) from the Middle Permian of Russia. Paleontol. J. 43, 178–182 (2009). https://doi.org/10.1134/S0031030109020087
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DOI: https://doi.org/10.1134/S0031030109020087