Abstract
The genus Catomus Allard and the allied genera of the fauna of Kazakhstan, Middle Asia, the Caucasus, and China are revised. The subgenus Stenomacidius Seidlitz, 1896 is synonymized with the genus Cylindronotus Faldermann, 1837. A new genus, Eustenomacidius gen. n., with the type species Helops luridus Ménétriés, 1848 is described. The following new combinations are proposed: Cylindronotus (s. str.) acutangulus (Seidlitz, 1896), Odocnemis anatolicus (Kaszab, 1961), Eustenomacidius hirtipennis (Seidlitz, 1896), Eustenomacidius turcmenicus (Medvedev, 1964), Eustenomacidius mongolicus (Kaszab, 1968), and Eustenomacidius wagnae (Ren, 1999). The following new subgenera are described: Montanocatomus, Sinocatomus (genus Catomus), and Caucasohelops (genus Eustenomacidius). The name Catomodontus Koch, 1935 is considered invalid. A new genus Xanthohelops closely related to Eustenomacidius is described from the Kara Kum Desert (Turkmenistan). The following new species and subspecies are described: Catomus (s. str.) noctivagus, C. (s. str.) indubitatus, C. (Montanocatomus) fabiani, C. (Sinocatomus) solitarius, Eustenomacidius (Caucasohelops) svetlanae with a new subspecies E. (Caucasohelops) svetlanae araxi subsp. n., Xanthohelops karakumicus Nabozhenko et Medvedev, spp. n. The following new synonyms are established: Cylindronotus Faldermann, 1837 = Stenomacidius Seidlitz, 1896; Eustenomacidius luridus (Ménétriés, 1848) = Stenomax laevicollis Kraatz, 1882, = Stenomax lucidicollis Kraatz, 1882, = Catomus (Stenomacidius) provocator Reitter, 1922; Catomus (Montanocatomus) reinigi (Schuster, 1931) = Catomus (Stenomacidius) alaensis G. Medvedev, 1970; Catomus fragilis (Ménétriés, 1848) = Catomus subniger Reitter, 1901; Catomus karakalensis G. Medvedev, 1964 = Catomus dolini G. Medvedev, 2004. Lectotypes are designated for the following species: Cylindronotus acutangulus (Seidlitz, 1896); Eustenomacidius hirtipennis (Seidlitz, 1896); Catomus fragilis (Ménétriés, 1848); Catomus reinigi (Schuster, 1931); Catomus provocator Reitter, 1922; Catomus subniger Reitter, 1901; Catomus niger (Kraatz, 1882) and Catomus antoniae Reitter, 1890. Morphological adaptations of species to particular environmental conditions, relationships of the new taxa and their positions in the tribe Helopini are considered. Keys to genera of the tribe Helopini of the Caucasus, Middle Asia, and Kazakhstan, and to subgenera and species of Catomus and Eustenomacidius of the Caucasus, Middle Asia, Kazakhstan, and China are given.
Similar content being viewed by others
References
Abdurakhmanov, G.M. and Medvedev, G.S., The Catalogue of Darkling Beetles of the Caucasus (DGPU, Makhachkala, 1994) [in Russian].
Abdurakhmanov, G.M. and Abdulmuslimova, K.M., “Composition, Morphological and Ecological Structure, and Zoogeographical Characteristics of the Tenebrionid Beetle Fauna of the Caucasus,” Russian Entomol. J. 11(1), 41–48 (2002).
Allard, E., “Revision des Helopides vrais de Lacordaire,” Abeille 14, 1–80 (1876).
Allard, E., “Revision des Helopides vrais,” Mitt. Schweiz. Entomol. Ges., 13–268 (1877).
Antoine, M., “Notes d’entomologie Marocaine. XLIV. Matériaux pour l’étude des Helopinae du Maroc (Col. Tenebrionides),” Bull. Soc. Sci. Nat. Maroc. 27, 123–162 (1947).
Bogachev, A.V., “A List of Species of the Beetle Families Tenebrionidae and Scarabaeidae (Collected in 1933 in the Nakhichevan ASSR),” Trudy Inst. Zool. Azerb. Otd. Akad. Nauk SSSR 8, 135–154 (1938).
Bogachev, A.V., “New Species of Tenebrionid Beetles (Tenebrionidae) from the Southern Part of Middle Asia,” Trudy Inst. Zool. i Parazitol. Akad. Nauk TadzhSSR 24, 95–102 (1963).
Bogachev, A.V., Doctoral Dissertation in Biology (Dushanbe, 1965).
Español, F., “Los Probaticus de España (Col. Tenebrionidae),” Eos 32, 83–123 (1956).
Español, F. and Viñolas, A., “Revision de los Catomus Ibericos (Col., Tenebrionidae),” Misc. Zool. 10, 181–184 (1986).
Gebien, H., “Katalog der Tenebrioniden,” Mitt. Münch. Entomol. Ges. 33(1), 399–430 (1943).
Gridelli, E., “Materiali per lo studio della fauna eritrea raccolti nel 1901–1903 dal dott. Alfredo Andreini,” Mem. Soc. Entomol. Ital. 18(5), 219–258 (1940 (1939–1940)).
Heyden, L. von and Kraatz, G., “Käfer um Samarkand, gesammelt von Haberhauer,” Deutsche Entomol. Zeitschr. 26(2), 297–338 (1882).
Heyden, L. von and Kraatz, G., “Beiträge zur Coleopteren-Fauna von Turkestan namentlich des Alai-Gebirges,” Deutsche Entomol. Zeitschr. 30, 189–190 (1886).
Iablokoff-Khnzorian, S.M., “Bemerkungen über einige Reitter’sche Typen aus dem Ungarischen Naturwissenschaftlichen Museum (Coleoptera),” Rov. Közlem. (Folia Entomol. Hung.) 17(20), 293–315 (1964).
International Code of Zoological Nomenclature, 4th ed. (St. Petersburg, 2000) [in Russian].
Kaszab, Z., “Zwei neue Tenebrioniden (Coleoptera) aus Arabien und Kleinasien,” Entom. Abh. 26(20), 169–175 (1961).
Kaszab, Z., “Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei,” Acta Zool. Acad. Sci. Hung. 14(3–4), 339–397 (1968).
Kaszab, Z., “Insects of Saudi Arabia. Coleoptera, Fam. Tenebrionidae (Part 2),” Fauna Saudi Arabia 4, 124–243 (1982).
Koch, C., “Wissenschaftliche Ergebnisse de entomologischen Expedition seiner Durchlaucht des Fuersten A. Della Tore e Tasso nach Aegypten und auf die Halbinsel Sinai: VII. Tenebrionidae (Coleoptera),” Bull. Soc. Roy. Entomol. Egypte 19, 2–111 (1935).
Kulzer, H., “Verzeichnis des Typenmaterials der Tenebrionidensammlung des Museums G. Frey, 25. Beitrag zur Kenntnis der Tenebrioniden,” Entomol. Arb. Mus. Frey 14, 375–434 (1963).
Löbl, I. and Merkl, O., “On the Type Species of Several Tenebrionid Genera and Subgenera (Coleoptera, Tenebrionidae),” Acta Zool. Acad. Sci. Hung. 49(3), 243–253 (2003).
Medvedev, G.S., “New Forms of Tenebrionid Beetles (Coleoptera, Tenebrionidae) from Middle Asia,” Entomol. Obozr. 43(3), 651–661 (1964).
Medvedev, G.S., “New Species of Tenebrionid Beetles (Coleoptera, Tenebrionidae) from the Arid Areas of the USSR and Afghanistan,” Entomol. Obozr. 49(2), 393–408 (1970).
Medvedev, G.S., “New species of Tenebrionid Beetles of the Tribe Helopini (Coleoptera, Tenebrionidae) from Middle Asia,” Trudy Zool. Inst. Akad. Nauk SSSR 71, 50–52 (1978).
Medvedev, G.S., “Tenebrionid Beetles of the Genus Zophohelops Rtt. and the Closely Related Genera (Coleoptera, Tenebrionidae) from Middle Asia and Kazakhstan,” Trudy Zool. Inst. Akad. Nauk SSSR 164, 95–129 (1987).
Medvedev, G.S., “A Key to Tenebrionid Beetles of Mongolia,” Trudy Zool. Inst. Akad. Nauk SSSR 220, 253 (1990).
Medvedev, G.S., “New Tenebrionid Species (Coleoptera, Tenebrionidae) from the Palaearctic Deserts,” Entomol. Obozr. 83(3), 553–580 (2004) [Entomol. Rev. 84 (5) 558–580 (2004)].
Medvedev, G.S. and Kaszab, Z., “Ergebnisse Mongolisch-Sovetischen Biologischen Expeditionen in der Mongolischen Volksrepublik (Tenebrionidae, excl. Tentyriini (Coleoptera),” Rov. Közlem. (Folia Entomol. Hung.) 26(1), 79–111 (1973).
Medvedev, G.S. and Lobanov, A.L., “A Faunistic List of Tenebrionid Beetles (Coleoptera, Tenebrionidae) of the Mongolian People’s Republic, with Coordinates of the Localities,” in Insects of Mongolia (Leningrad, 1990), No. 11, pp. 139–204 [in Russian].
Medvedev, G.S. and Nepesova, M.G., A Key to Tenebrionid Beetles of Turkmenistan (Ylym, Ashkhabad, 1985) [in Russian].
Ménétriés, E., “Descriptions des Insectes recueillis par feu M. Lehmann,” Mém. Acad. Sci. St.-Petersbourg. Sér 6. Sci. Math. Phys. Nat. 8, part 2 (= Sci. Nat. 6), livr. 1/2, pp. 17–66 (1848 [II]); 8, part 2 (= Sci. nat. T. 6), livr. 4, pp. 217–328 (1849 [XI]).
Mordkovich, V.G., Zoological Diagnostics of Soils of the Forest-steppe and Steppe Zones of Siberia (Nauka, Novosibirsk, 1977) [in Russian].
Nabozhenko, M.V., “Taxonomic Notes on the Genus Zophohelops Reitter, 1901 with Description of New Species from Tadzhikistan and New Genus Pseudoprobaticus gen. n. (Coleoptera, Tenebrionidae),” Ann. Zool. 51(4), 113–117 (2001).
Nabozhenko, M.V., Candidate’s Dissertation in Biology (St. Petersburg, 2002a).
Nabozhenko, M.V., “Taxonomic Significance of the Female Genital Tubes in the Tribe Helopini (Coleoptera, Tenebrionidae),” in Proceedings of XII Congress of the Russian Entomological Society, St. Petersburg, August 19–24, 2002 (St. Petersburg, 2002b).
Nabozhenko, M.V., “Interstructural Correlations in the Evolution of Tenebrionid Beetles of the tribe Helopini (Coleoptera, Tenebrionidae),” Caucas. Entomol. Bull. 1(1), 37–48 (2005).
Nepesova, M.G., Tenebrionid Beetles of Turkmenistan (Biology and Ecology) (Ylym, Ashkhabad, 1980) [in Russian].
Pirnazarov, B., Candidate’s Dissertation in Biology (Ashkhabad, 1973).
Reinig, W.F., “Entomologische Ergebnisse der Deutsch-Russischen Alai-Pamir-Expedition 1928 (II): 5. Coleoptera II. Tenebrionidae,” Mitt. Zool. Mus. Berlin 16(6), 865–912 (1931).
Reitter, E., “Neue Coleopteren aus Europa, den angrenzenden Ländern, und Sibirien, mit Bemerkungen über bekannte Arten,” Deutsche Entomol. Zeitschr. X, 165–176 (1890).
Reitter, E., “Verschiedenes über die Coleopteren der Tenebrioniden-Abtheilung Helopinae,” Deutsche Entomol. Zeitschr., No. 2, 209–224 (1901).
Reitter, E., “Bestimmungs-Tabellen der europaischen Coleopteren. H. 92. Tenebrionidae. 16. Teil: Unterfamilie Helopina, I,” Wien. Entomol. Ztg. 39, 1–44 (1922).
Ren, G. and Youzhi, Yu., The Darkling Beetles from Deserts and Semideserts of China (Coleoptera, Tenebrionidae) (Hebei Univ. Publishing House, Hebei, 1999).
Seidlitz, G., Naturgeschichte der Insecten Deutschlands. Coleoptera. Tenebrionidae, Helopinae (Berlin, 1896), Vol. 5.
Skopin, N.G., “Contributions to the Knowledge of the Tenebrionid Beetle (Coleoptera, Tenebrionidae) Fauna of Mangyshlak and Northwestern Ustyurt,” Trudy Nauch.-Issled. Inst. Zashchity Rastenii (Alma-Ata) 8, 277–297 (1964).
Tadzhibaev, M., “Tenebrionid Beetles (Coleoptera, Tenebrionidae) from the Low-Mountain Areas of Southern Tajikistan,” Entomol. Obozr. 51(2), 274–281 (1972).
Vauloger, M., “Contribution au catalogue des coléoptères du nord de l’Afrique. Helopini,” Ann. Soc. Entomol. France 68, 669–722 (1899).
Author information
Authors and Affiliations
Additional information
Original Russian Text © M.V. Nabozhenko, 2006, published in Entomologicheskoe Obozrenie, 2006, Vol. 85, No. 4, pp. 798–857.
Rights and permissions
About this article
Cite this article
Nabozhenko, M.V. A revision of the genus Catomus Allard, 1876 and the allied genera (Coleoptera, Tenebrionidae) from the Caucasus, Middle Asia, and China. Entmol. Rev. 86, 1024–1072 (2006). https://doi.org/10.1134/S0013873806090065
Received:
Issue Date:
DOI: https://doi.org/10.1134/S0013873806090065