Journal of Ichthyology

, Volume 47, Issue 1, pp 1–20 | Cite as

Materials towards the revision of the genus Triplophysa rendahl, 1933 (Cobitoidea: Balitoridae: Nemacheilinae): A revision of nominal taxa of Herzenstein (1888) described within the species “Nemachilus” stoliczkae and “N.” dorsonotatus, with the description of the new species T. scapanognatha sp. nova

  • A. M. Prokofiev


This study reviews the nominal taxa within the species “Nemachilus” stoliczkae and “N.” dorsonotatus, described by Herzenstein (1888). N. dorsonotatus, N. d. retropinnis, and N. plagiognathus are reviewed within the species Triplophysa stoliczkae; it is suggested that this species is represented by at least 5 subspecies. N. stoliczkae brevicauda, N. s. leptosoma (synonym, N. s. productus), and N. s. crassicauda are considered as valid species, the last 2 are related to the group including the species T. tenuis, T. choprai, T. tenuicauda, and T. yasinensis rather than with T. stoliczkae. It was shown that the “stoliczkae” species group (sensu Prokofiev, 2001) should be divided into three species groups: “stoliczkae” (s. str.), “tenuis,” and “robusta”; their diagnoses and the species composition are presented. A new species from the “stoliczkae” group is described (T. scapanognatha sp. nova), related to T. tanggulaensis (Zhu, 1982).


Posterior Margin Lateral Line Head Length Baga Caudal Peduncle 
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  1. 1.
    N. Annandale and S. L. Hora, “The Fishes of Seistan,” Rec. Indian Mus. 18Part 1, 151–189 (1920).Google Scholar
  2. 2.
    V. P. Anikin, Description of New Asiatic Fish Varieties (Tomsk. Univ., Tomsk, 1905) [in Russian].Google Scholar
  3. 3.
    P. Bănărescu and T.T. Nalbant, “A Generical Classification of Nemacheilinae with Description of Two New Genera (Teleostei: Cypriniformes: Cobitidae),” Trav. Mus. Hist. Nat. “Gr. Antipa” 35, 42–495 (1995).Google Scholar
  4. 4.
    L. S. Berg, Fishes of Turkestan (St. Petersburg, 1905) [in Russian].Google Scholar
  5. 5.
    L. S. Berg, “Fishes,” Tr. Pamirsk. Eksped., Vol. 8: Zoology (1932), pp. 23–28.Google Scholar
  6. 6.
    L. S. Berg, Freshwater Fishes of the USSR and Adjacent Countries, 3rd ed. (VNIORKh, Leningrad, 1933), Part 2, pp. 545–903 [in Russian].Google Scholar
  7. 7.
    L. S. Berg, Freshwater Fishes of the USSR and Adjacent Countries (Akad. Nauk SSSR, Moscow, 1949), Vol. 2, pp. 469–925 [in Russian].Google Scholar
  8. 8.
    W.-X. Cao and S.-Q. Zhu, “Two New Species of the Genus Triplophysa from Qinghai-Xizang Plateau, China,” Acta Zootaxon. Sin. 13(2), 201–204 (1988).Google Scholar
  9. 9.
    Y. Chen, The Fishes of the Hengduan Mountains Region (Sci. Press, Beijing, 1998).Google Scholar
  10. 10.
    Q. Cheng and B. Zheng, Systematic Synopsis of Chinese Fishes (Sci. Press, Beijing, 1987), Vol. 1.Google Scholar
  11. 11.
    L. Di Caporiacco, “Pesci raccolti durante la Spedizione italiana al Karakoram 1929-VII,” Monit. Zool. Ital. 44(11–12), 339–344 (1933).Google Scholar
  12. 12.
    P. W. Fang, “On Some Nemachilus Fishes of North-Western China and Adjacent Territory in the Berlin Zoological Museum’s Collection, with Description of Two New Species,” Sinensia 6(6), 749–767 (1935).Google Scholar
  13. 13.
    S. M. Gertsenshtein, Scientific Results of Journeys of N.M. Przheval’skii over Central Asia Published by the Academy of Sciences. Department of Zoology, Vol. 3,Part 2: Fishes (St. Petersburg, 1888), Issue 1, 1–90 [in Russian].Google Scholar
  14. 14.
    S. L. Hora, “Notes on Fishes in the Indian Museum. III. On Fishes Belonging to the Family Cobitidae from High Altitudes in Central Asia,” Rec. Indian Mus. 24Part 1, 63–83 (1922).Google Scholar
  15. 15.
    K. C. Jayaram, The Freshwater Fishes of India, Pakistan, Bangladesh, Burma and Sri Lanka. A Handbook (Zool. Surv. India, Calcutta, 1981).Google Scholar
  16. 16.
    K. Kessler, “Ichthyofauna of Turkestan,” Izv. O-va Lyubitelei Estestvozn., Antropol. Etnografii 10, 1–63 (1874).Google Scholar
  17. 17.
    K. Kessler, “The Journey of A.P. Fedchenko to Turkestan. Fishes,” Izv. O-va Lyubitelei Estestvozn., Antropol. Etnografii 11, 1–63 (1874).Google Scholar
  18. 18.
    K. Kessler, “Beiträge zur Ichthyologie von Central-Asien,” Mélanges Biol. Tirés Bull. Acad. Sci. Pétersb. 10, 282–310 (1879).Google Scholar
  19. 19.
    M. A. V. Lakshmanan, “A Note on the Fishes from the Helmund River in Afghanistan, with a Description of a New Loach,” Rec. Indian Mus. 47Part 2, 217–224 (1950).Google Scholar
  20. 20.
    A. G. K. Menon, The Fauna of India and the Adjacent Countries. Pisces. IV. Teleostei-Cobitoidea. Part 1: Homalopteridae (Zool. Surv. India, Calcutta, 1987).Google Scholar
  21. 21.
    V. P. Mitrofanov, “Genus Noemacheilus van Hasselt, 1823—Charr,” in Fishes of Kazakhstan (Nauka KazSSR, Alma-Ata, 1989), pp. 6–63.Google Scholar
  22. 22.
    A. M. Nikol’skii, “Three New Fish Species from Central Asia (Schizothorax kozlovi sp. n., Ptychobarbus kaznakowi sp. n., Nemachilus fedtschenkoae sp. n.),” Ezhegodn. Zool. Muz. Imp. Akad. Nauk 8, 90–94 (1903).Google Scholar
  23. 23.
    G. V. Nikolsky, Fishes of Tajikistan (Akad. Nauk SSSR, Moscow, 1938) [in Russian].Google Scholar
  24. 24.
    A. M. Prokofiev, “Four New Species of the Triplophysa stoliczkai-Complex from China (Pisces: Cypriniformes: Balitoridae),” Zoosystematica Rossica 10(1), 193–207 (2001).Google Scholar
  25. 25.
    A. M. Prokofiev, “Stone Loaches (Balitoridae: Nemacheilinae) from the Uvs-Nuur Lake Basin at the Boundary of Tuva and Mongolia,” J. Ichthyol. 42(Suppl. 1), S45–S59 (2002).Google Scholar
  26. 26.
    A. M. Prokofiev, “Review of the Triplophysa bombifrons-Species Group (Balitoridae: Nemacheilinae) with a Description of a New Species from Kam Valley, China,” Vopr. Ikhtiol. 42(4), 468–474 (2002) [J. Ichthyol. 42 (6), 427–433 (2002)].Google Scholar
  27. 27.
    A. M. Prokofiev, “Little Known and New Species of the Genus Triplophysa Rendahl, 1933 (Osteichthyes: Balitoridae: Nemacheilinae) from Northern China and Tibet with Additional Remarks on the “Bombifrons” Species Group,” Estestv. Tekhn. Nauki, No. 1, 53–72 (2003).Google Scholar
  28. 28.
    A. M. Prokofiev, “Revision of the Species-Complex of Triplophysa labiata with Description of a New Species, T. kaznakowi sp. n. (Osteichthyes, Balitoridae, Nemacheilinae),” Senckenberg. Biol. 83(2), 181–208 (2004).Google Scholar
  29. 29.
    H. Rendahl, “Stüdien über Innerasiatische Fische,” Arkiv Zool. 25, Ser. A (11), 1–51 (1933).Google Scholar
  30. 30.
    F. Steindachner, “Ichthyologische Mittheilungen VI. Zur fisch-fauna Kaschmirs und der benachbarten Landerstriche,” Verh. K.-K. Zool.-Bot. Ges. Wien. 16, 761–796 (1866).Google Scholar
  31. 31.
    P. K. Talwar and A. G. Jhingran, Inland Fishes of India and Adjacent Countries (A.A. Balkema, Rotterdam, 1992), Vol. 1.Google Scholar
  32. 32.
    T.-L. Tchang, T.-H. Yueh, and H.-C. Hwang, “Notes on Fishes of the Genus Nemachilus of Southern Tibet, China, with Description of a New Species,” Acta Zool. Sin. 15(4), 624–635 (1963).Google Scholar
  33. 33.
    F. A. Turdakov, “On the Systematics and Distribution of Some Species of the Genus Nemachilus Hasselt (Pisces, Cypriniformes) in Kirgizia and Middle Asia,” Tr. Kirgiz. Ped. Inst. 1946(1948) 1(1), 49–59 (1946).Google Scholar
  34. 34.
    F. A. Turdakov, Fishes of Kirgizia (KirgizFAN, Frunze, 1952) [in Russian].Google Scholar
  35. 35.
    F. A. Turdakov, “Materials on the Ichthyofauna of the Ters (Assa) River and Talas,” Tr. Inst. Zool. Parazitol. Kirgiz. Fil. Akad. Nauk SSSR 1, 113–122 (1954).Google Scholar
  36. 36.
    F. A. Turdakov, Fishes of Kirgizia, 2nd ed. (Akad. Nauk KirgSSR, Frunze, 1963) [in Russian].Google Scholar
  37. 37.
    Y.-F. Wu and Y. Chen, “Notes on Fishes from Golog and Yushu Region of Qinghai Province, China,” Acta Zootaxon. Sin. 4(3), 287–296 (1979).Google Scholar
  38. 38.
    Y.-F. Wu and C.-Z. Wu, “Notes on Fishes from the Sources of the Huanghe River and Cocha Lake, China,” Acta Zootaxon. Sin. 13(1), 195–200 (1988).Google Scholar
  39. 39.
    Y.-F. Wu and C.-Z. Wu, The Fishes of the Qinghai-Xizang Plateau (Sichuan Publ. House of Sci. and Technol., Chengdu, 1992).Google Scholar
  40. 40.
    T.-Q. Xu and C.-G. Zhang, “A New Species of Cobitid Fishes from Tibet, China (Cypriniformes: Cobitidae),” Acta Zootaxon. Sin. 21(3), 377–379 (1996).Google Scholar
  41. 41.
    G.-R. Yang and G.-X. Xie, “New Species of Loaches (Pisces: Cobitidae) from Upper Changjiang River,” Acta Zootaxon. Sin. 8(3), 314–316 (1983).Google Scholar
  42. 42.
    T.-Q. Zhao, “Taxonomic Problems of Some Nemachilus Fishes,” Zool. Res. 5(4), 341–346 (1984).Google Scholar
  43. 43.
    T.-Q. Zhao, “Fish-Fauna and Zoogeographical Division of Hexi-Alashan Region, the Northwest China,” Acta Zool. Sin. 37(2), 153–167 (1991).Google Scholar
  44. 44.
    T. Zhao and X. Wang, “Fishes and Fishery Problems of Hexi Area, Gansu Province, with Descriptions of a New Subspecies and an Unidentified Species of Cobitidae,” J. Lanzhou Univ. Nat. Sci. 24(4), 109–119 (1988).Google Scholar
  45. 45.
    S.-Q. Zhu, “New Species of the Genus Nemachilus (Pisces: Cobitidae) from Quinghai Province, China,” Acta Zootaxon. Sin. 7(2), 223–224 (1982).Google Scholar
  46. 46.
    S.-Q. Zhu, The Loaches of the Subfamily Nemacheilinae in China (Cypriniformes: Cobitidae) (Jiangsu Sci. and Technol. Publ. House, Nanjing, 1989).Google Scholar

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© Pleiades Publishing, Ltd. 2007

Authors and Affiliations

  • A. M. Prokofiev
    • 1
  1. 1.Severtsov Institute of Ecology and Evolution (IPEE)Russian Academy of SciencesMoscowRussia

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