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How not to measure inclusive fitness

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Author notes

  1. Alan Grafen: Marian Dawkins, Mark Ridley and Richard Dawkins are thanked for their comments on an earlier draft of the manuscript.

Authors and Affiliations

  1. Animal Behaviour Research Group, Department of Zoology, Oxford, OX1 3PS

    Alan Grafen

  2. a Research Lecturer of Christ Church College, Oxford

    Alan Grafen

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  1. Alan Grafen
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Grafen, A. How not to measure inclusive fitness. Nature 298, 425–426 (1982). https://doi.org/10.1038/298425a0

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