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The genus Callicarpa (Lamiaceae) in the Philippines

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Summary

A taxonomic revision of the genus Callicarpa L. (Lamiaceae) in the Philippines is presented here. Twenty seven taxa are recognised, and thirteen names newly reduced to synonymy. A key to all species is included, as well as distribution maps and a preliminary conservation status for each species. This account forms part of a treatment of Callicarpa for the Flora Malesiana project.

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Notes

  1. These alternate ‘leaves’, since they are subtending cymes, are strictly bracts, not leaves, but are included here to avoid confusion, since they are identical in structure to the leaves, and are likely to be understood as such.

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Acknowledgements

I would like to thank the curators of A, BM, BO, C, E, G, GH, KEP, L, NY, SING and US for allowing access to their collections for study, or for providing specimens on loan or for making digital images of specimens available. Distribution maps were produced with guidance and assistance from Tim Utteridge and Steve Bachman. I am grateful to David Middleton for his detailed comments on the manuscript.

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Authors and Affiliations

  1. Herbarium, Library, Art & Archives, Royal Botanic Gardens, Kew, Richmond, Surrey, TW9 3AB, UK

    Gemma L. C. Bramley

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  1. Gemma L. C. Bramley
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Correspondence to Gemma L. C. Bramley.

Appendix 1: Identification List

Appendix 1: Identification List

Adduru, M. 44 (C. bicolor) [GH, K]; Adduru, M. 30 (C. pedunculata) [A, K]; Adduru, M. 150 (C. erioclona) [A, K, US]; Ahern, G. P. 662 (C. pedunculata) [US]; Ahern, G. P. 662 (C. pedunculata) [US]; Ahern, G. P. 540 (C. erioclona) [US]; Ahern, G. P. 811 (C. pedunculata) [US,]; Ahern, G. P. 318 (C. surigaensis) [US, US]; Ahern's collector For. Bur. 1888 (C. angusta) [K, NY]; Ahern's collector For. Bur. 1484 (C. bicolor) [K, NY]; Ahern's Collector For. Bur. 3300 (C. erioclona) [K, US]; Alcasid PNH1607 (C. micrantha) [A]; Alcasid, G. L. PNH1869 (C. caudata) [A, K]; Amihan, J. PNH33376 (C. pedunculata) [US]; Anonuevo, P. PNH13622 (C. bicolor) [GH, K]; Anonuevo, P. PNH13579 (C. erioclona) [A].

Baines, P. T. For. Bur. 55 (C. pedunculata) [NY]; Banaga, T. B. PNH33393 (C. pedunculata) [US]; Barbon, Garcia, Romero PPI18943 (C. pedunculata) [GH]; Barbon, Alvarez, Fuentes PPI8765 (C. pedunculata) [K]; Barbon, Alvarez, Fuentes PPI8602 (C. ramiflora) [K]; Barbon, Alvarez, Garcia PPI2021 (C. pedunculata) [K]; Barbon, Alvarez, Garcia PPI5534 (C. pedunculata) [K]; Barbon, Alvarez, Garcia PPI2148 (C. platyphylla) [K, PNH]; Barbon, Garcia, Sagcal PPI6032 (C. pedunculata) [K]; Barbon, Garcia, Sagcal PPI5913 (C. surigaensis) [K]; Barbon, Pedersen, HGG, LSF PPI12843 (C. subglandulosa) [K]; Barbon, Romero, Fernando PPI1650 (C. erioclona) [K]; Barbon, Romero, Fernando PPI1834 (C. pedunculata) [K]; Barbon, Romero, Fuentes PPI8014 (C. aff. arborea) [K,]; Barbon, Romero, Fuentes PPI13108 (C. angusta) [K]; Barnes, P. T. For. Bur. 924 (C. caudata) [K, US]; Barnes, P. T. For. Bur. 924 (C. caudata) [NY]; Barnes, P. T. For. Bur. 55 (C. pedunculata) [K, NY, US]; Barrow, J. V. s.n. (C. bicolor) [K, NY, US]; Bartlett, H. H. 14711 (C. bicolor) [NY]; Bartlett, H. H. 13211 (C. caudata) [A]; Bartlett, H. H. 14698a (C. bicolor) [A]; Bermejos, J. Bur. Sci.1534 (C. bicolor) [NY]; Bolster, F. H. 17 (C. pedunculata) [K]; Borden, T. E. For. Bur. 1595 (C. erioclona) [NY, US]; Bourell, M. 2197 (C. bicolor) [A]; Brenan, J. P. M. 14975 (C. erioclona) [K]; Burley, J. S. 33 (C. caudata) [A]; Burley, J. S. 40 (C. caudata) [A].

Canicosa, F. For. Bur. 30020 (C. caudata) [US]; Cardona, F. T. For. Bur. 23866 (C. pentandra) [US]; Castro, A. PNH5681 (C. pentandra) [A]; Castro, A. PNH5764 (C. surigaensis) [K]; Castro, A. PNH6496 (C. platyphylla) [K]; Celestrino, M. PNH7834 (C. pedunculata) [A]; Clark, W. W. For. Bur. 2534 (C. caudata) [K, US]; Clark, W. W. For. Bur. 2534 (C. caudata) [NY]; Clemens, M. S. Bur. Sci. 15676 (C. bicolor) [NY]; Clemens, M. S. 190 (C. longifolia) [US]; Clemens, M. S. 17400 (C. pedunculata) [NY]; Clemens, M. S. 271 (C. pentandra) [US]; Clemens, M. S. Bur. Sci. 15639 (C. apoensis) [NY]; Clemens, M. S. 9185 (C. longipetiolata) [A]; Co, L. 3756 (C. longipetiolata) [A, K, L]; Co, L. 3536 (C. pedunculata) [GH, K]; Co, L. 3464 (C. platyphylla) [A, K, L, US]; Co, L. 3776 (C. subglandulosa) [A, K, L, SING, US]; Conese, E. PNH15801 (C. pedunculata) [GH, K]; Conklin, H. C. PNH17535 (C. pedunculata) [GH]; Conklin, H. C. PNH19016 (C. erioclona) [GH]; Conklin, H., del Rosario PNH42661 (C. pedunculata) [GH, K]; Conklin, H., del Rosario PNH72689 (C. pedunculata) [GH, K]; Copeland, E. B. 123 (C. pedunculata) [K, US]; Cuming 1425 (C. angusta) [K, NY]; Cuming 1266 (C. arborea) [K, NY]; Cuming 1095 (C. caudata) [K]; Cuming 1330 (C. dolichophylla) [NY]; Cuming 911 (C. erioclona) [GH, K, NY]; Cuming 1778 (C. erioclona) [K]; Cuming 1165 (C. micrantha) [K]; Cuming 1460 (C. micrantha) [K, NY]; Cuming 1641 (C. micrantha) [K]; Cuming 526 (C. pedunculata) [K]; Cuming 1330 (C. dolichophylla) [K, K]; Cuming, H. 1283 (C. bicolor) [K]; Cuming 1773 (C. pentandra) [K]; Curran, H. M. For. Bur. 5140 (C. pedunculata) [US]; Curran, H. M. For. Bur. 17181 (C. platyphylla) [US]; Curran, H. M., Merritt, M. L., Zschokke, T. C. For. Bur. 18162 (C. caudata) [US].

Danao, F. Bur. Sci. 12444 (C. pentandra) [US]; Darling, J. W. For. Bur. 16574 (C. caudata) [US]; Deguchi, K., Tsugaru, S., Inoue, Y. 6313 (C. bicolor) [NY]; Deguchi, K., Tsugaru, S., Inoue, Y. 6339 (C. bicolor) [NY]; DeVore, Hoover 175 (C. bicolor) [US]; DeVore, Hoover 41 (C. caudata) [US]; DeVore, Hoover 106 (C. pedunculata) [US].

Ebalo, L. E., Conklin, H. C. 1364 (C. longifolia) [A]; Edaño, G. Bur. Sci. 46162 (C. bicolor) [NY]; Edaño, G. PNH11436 (C. caudata) [A, SING]; Edaño, G. Bur. Sci. 41685 (C. erioclona) [A, K]; Edaño, G. Bur. Sci. 78500 (C. erioclona) [A]; Edaño, G. Bur. Sci. 35285 (C. micrantha) [A, K]; Edaño, G. Bur. Sci. 41674 (C. micrantha) [BM, A, K, K, SING, US]; Edaño, G. Bur. Sci. 41674 (C. micrantha) [A, K, K, US]; Edaño, G. Bur. Sci. 42365 (C. micrantha) [A]; Edaño, G. Bur. Sci. 48817 (C. micrantha) [NY]; Edaño, G. PNH11126 (C. micrantha) [A, SING]; Edaño, G. PNH11263 (C. micrantha) [A]; Edaño, G. Bur. Sci. 48766 (C. pedunculata) [A, NY]; Edaño, G. Bur. Sci. 48817 (C. pedunculata) [NY]; Edaño, G. Bur. Sci. 78469 (C. pedunculata) [A]; Edaño, G. Bur. Sci. 78550 (C. pedunculata) [A]; Edaño, G. PNH7149 (C. pedunculata) [A]; Edaño, G. Bur. Sci. 45975 (C. pentandra) [K]; Edaño, G. Bur. Sci. 45975 (C. pentandra) [NY]; Edaño, G. PNH1315 (C. pentandra) [K]; Edaño, G. PNH9833 (C. pentandra) [A]; Edaño, G. Bur. Sci. 78610 (C. angusta) [A]; Edaño, G. Bur. Sci. 38333 (C. erioclona) [A]; Edaño, G. Bur. Sci. 78234 (C. erioclona) [A]; Edaño, G. Bur. Sci. 78309 (C. erioclona) [A]; Edaño, G. Bur. Sci. 78325 (C. platyphylla) [A]; Edaño, G. Bur. Sci. 78604 (C. platyphylla) [A]; Edaño, G. Bur. Sci. 79528 (C. platyphylla) [A]; Edaño, G. E. PNH19857 (C. pedunculata) [K]; Edaño, G. E. PNH34552 (C. pedunculata) [A]; Edaño, G. E. PNH3535 (C. pedunculata) [A]; Edaño, G. E. PNH37162 (C. pedunculata) [K]; Edaño, G. E. PNH11680 (C. bicolor) [K]; Edaño, G. E. PNH4142 (C. erioclona) [A]; Edaño, G. E. PNH4168 (C. erioclona) [A]; Edaño, G. E. PNH11263 (C. micrantha) [K]; Edaño, G. E., Gutierrez, H. PNH37721 (C. pedunculata) [K]; Elmer, A. D. E. 11491 (C. apoensis) [BM, BO, GH, NY]; Elmer, A. D. E. 8193 (C. arborea) [K, NY]; Elmer, A. D. E. 9125 (C. arborea) [K, NY]; Elmer, A. D. E. 17575 (C. arborea) [BM, K, NY]; Elmer, A. D. E. 22377 (C. arborea) [BM, K, NY, SING]; Elmer, A. D. E. 18086 (C. bicolor) [GH, K, NY, US]; Elmer, A. D. E. 5784 (C. caudata) [K, NY, US]; Elmer, A. D. E. 5870 (C. caudata) [K, NY, US]; Elmer, A. D. E. 8646 (C. caudata) [GH, K, NY, US]; Elmer, A. D. E. 10375 (C. caudata) [GH, K, US]; Elmer, A. D. E. 11333 (C. caudata) [GH, K, NY, US]; Elmer, A. D. E. 6647 (C. erioclona) [K, NY]; Elmer, A. D. E. 7367 (C. erioclona) [GH, K, NY]; Elmer, A. D. E. 11190 (C. erioclona) [GH, K, US]; Elmer, A. D. E. 15082 (C. erioclona) [GH, K, NY, US]; Elmer, A. D. E. 15124 (C. erioclona) [NY]; Elmer, A. D. E. 17598 (C. erioclona) [GH, K, NY, US]; Elmer, A. D. E. 11851 (C. flavida) [BM, E, K, NY]; Elmer, A. D. E. 13822 (C. flavida) [C, E, K, NY]; Elmer, A. D. E. 13861 (C. flavida) [BM, E, K, NY]; Elmer, A. D. E 13536 (C. longifolia) [K, NY]; Elmer, A. D. E 6266 (C. longipetiolata) [K]; Elmer, A. D. E. 14280 (C. longipetiolata) [NY]; Elmer, A. D. E. 14286 (C. longipetiolata) [K]; Elmer, A. D. E. 12144 (C. micrantha) [BM, GH, US]; Elmer, A. D. E. 14513 (C. paloensis) [BM, K]; Elmer, A. D. E. 16939 (C. paloensis) [BM, GH, K, NY]; Elmer, A. D. E. 5629 (C. pedunculata) [K, NY, US]; Elmer, A. D. E. 7366 (C. pedunculata) [GH, K, NY]; Elmer, A. D. E. 8151 (C. pedunculata) [K, NY]; Elmer, A. D. E. 10985 (C. pedunculata) [GH, NY, US]; Elmer, A. D. E. 13441 (C. pedunculata) [GH, K, NY, US]; Elmer, A. D. E. 14352 (C. pedunculata) [GH, NY, US]; Elmer, A. D. E. 17611 (C. pedunculata) [K, NY]; Elmer, A. D. E. 22211 (C. pedunculata) [GH, K]; Elmer, A. D. E. 7368 (C. pentandra) [K, NY]; Elmer, A. D. E. 13351 (C. pentandra) [NY]; Elmer, A. D. E. 9184 (C. subalbida) [K]; Elmer, A. D. E. 8064 (C. erioclona) [K]; Elmer, A. D. E. 15124 (C. erioclona) [BM, K, NY, US]; Elmer, A. D. E. 7370 (C. paloensis) [NY]; Elmer, A. D. E. 10856 (C. pentandra) [NY]; Elmer, A. D. E. 11102 (C. pentandra) [NY]; Elmer, A. D. E. 6266 (C. longipetiolata) [NY]; Escritor, L. Bur. Sci. 21104 (C. pedunculata) [US]; Escritor, L. Bur. Sci. 21155 (C. pedunculata) [K].

Fairchild, D. 157 (C. pentandra) [A]; Fénix, E. Bur. Sci. 15802 (C. caudata) [K, US]; Fénix, E. Bur. Sci. 3622 (C. denticulata) [K]; Fénix, E. Bur. Sci. 28048 (C. micrantha) [GH, K]; Fénix, E. 148 (C. pedunculata) [A]; Fénix, E. Bur. Sci. 26058 (C. pedunculata) [A, BM, US]; Fénix, E. 245 (C. bicolor) [A]; Fénix, E. Bur. Sci. 29884 (C. bicolor) [US,]; Fontanoza, J. For. Bur. 31131 (C. pentandra) [NY]; Fortes, C. O. PNH9604 (C. pedunculata) [A]; Fox, R. B. PNH4725 (C. pedunculata) [A]; Fox, R. B. PNH13401 (C. bicolor) [A]; Foxworthy, F. W. Bur. Sci.12235 (C. pedunculata) [US]; Foxworthy, F. W. Bur. Sci. 660 (C. angusta) [NY]; Foxworthy, F. W. Bur. Sci. 719 (C. angusta) [NY, US]; Foxworthy, F. W. 4 (C. erioclona) [A]; Foxworthy, F. W. 13 (C. erioclona) [US]; Foxworthy, F. W. 18 (C. erioclona) [US]; Fuentes, R., Fernando, L. PPI37156 (C. erioclona) [K]; Fuentes, R., Argent, G. C. G., Mendum, M., Wilkie, P. 99285 (C. caudata) [A, E]; Fuentes, de la Rosa PPI38813 (C. pedunculata) [K].

Gaerlan, F. J. M. PPI9897 (C. pedunculata) [K]; Gaerlan, F. J. M., Sagcal, E. C. PPI10072 (C. nigrescens) [K]; Gaerlan, Fernando, EST. PPI19158 (C. flavida) [K]; Gaerlan, Fernando, Majaducon PPI19995 (C. pentandra) [K]; Gaerlan, F. M. PPI3009 (C. platyphylla) [K]; Gaerlan, Fuentes, Romero PPI26277 (C. micrantha) [GH]; Gaerlan, Romero, Sagcal PPI13404 (C. pentandra) [K]; Gaerlan, Sagcal, Fernando PPI4776 (C. bicolor) [K]; Gaerlan, Sagcal, Fernando PPI4792 (C. erioclona) [K]; Gaerlan, Sagcal, Fernando PPI4572 (C. flavida) [K]; Gaerlan, Sagcal, Fernando PPI4873 (C. pentandra) [K]; Gaerlan, Sagcal, Romero PPI11012 (C. aff. basilanensis) [K]; Gaerlan, Sagcal, Romero PPI11052 (C. bicolor) [K]; Galutera, R. PNH33358 (C. erioclona) [US]; Garcia, H. G., EMR, ECS, REF, PPI15319 (C. pedunculata) [K]; Garcia, Fuentes, Romero PPI18417 (C. pedunculata) [K]; Gates, F. C. 6455 (C. caudata) [NY]; Gates, F. C. 6599 (C. erioclona) [NY]; Gates, F. C. 6286 (C. pedunculata) [NY]; Glassman, S. F. 579 (C. pedunculata) [GH]; Godofredo, Alcasid PNH1591 (C. micrantha) [GH]; Gregory, V. B. 33 (C. pedunculata) [K]; Guerrero, F. For. Bur. 30368 (C. pentandra) [NY]; Gutierrerz, H. R. PNH78040 (C. pedunculata) [GH]; Gutierrez, H. G. PNH78070 (C. pachyclada) [K]; Gutierrez, H. G. PNH117320 (C. pentandra) [K].

Haenke, T. 579 (C. erioclona) [NY]; Hallier, H. 4024a (C. caudata) [A, K]; Hallier, H. 4027a (C. erioclona) [GH, K]; Hinolan, P. For. Bur. 24043 (C. pentandra) [NY]; Holman, R. M. 187 (C. bicolor) [GH]; Holman, R. M. 40 (C. pedunculata) [A, US]; Hutchinson, W. J. For. Bur. 3974 (C. basilanensis) [K]; Hutchinson, W. J. Forestry Bureau 6124 (C. basilanensis) [NY].

Ingle, N. R. 287 (C. caudata) [K].

Jacobs, M. 7186 (C. caudata) [A, K]; Jacobs, M. 7864 (C. pedunculata) [A, K]; Jacobs, M. 7722 (C. angusta) [A]; Jacobs, M. 7122 (C. subglandulosa) [K]; Jacobs, M. 7482 (C. subglandulosa) [K]; Jose Vera Santos 5810 (C. longipetiolata) [US]; Jose Vera Santos 5239 (C. pedunculata) [US].

Kanehira, R. 2518 (C. pentandra) [NY]; Kanehira, R. 2517 (C. erioclona) [NY]; Kondo, Y., Edaño, G. PNH36778 (C. bicolor) [K]; Kruckeberg, A. R., Brown, H. S. s.n. (C. pedunculata) [NY].

Lam, H. J. 3372 (C. bicolor) [NY]; Leiberg, J. B. 6018 (C. bicolor) [K]; Leiberg, J. B. Bur. Govt. Labs 6000 (C. micrantha) [K]; Lete, R. 252 (C. pedunculata) [A, K, US]; Liborio Ela Ebalo 792 (C. basilanensis) [NY]; Liborio Ela Ebalo 1142 (C. longifolia) [A, K]; Liborio Ela Ebalo 1080 (C. pentandra) [K]; Liborio Ela Ebalo 1131 (C. pentandra) [K]; Liborio Ela Ebalo 1189 (C. pentandra) [K]; Lizardo, L. For. Bur. 29935 (C. pedunculata) [NY]; Llalnos 184 (C. erioclona) [GH]; Loher, A. 4439 (C. caudata) [K]; Loher, A. 4440 (C. caudata) [K, US]; Loher, A. 4438 (C. micrantha) [K]; Loher, A. 4445 (C. micrantha) [K]; Loher, A. 4441 (C. pedunculata) [US]; Loher, A. 12312 (C. angusta) [A]; Loher, A. 4442 (C. bicolor) [K]; Loher, A. 4443 (C. bicolor) [K, US]; Loher, A. 4444 (C. bicolor) [K, US]; Loher, A. 4447 (C. erioclona) [K]; Loher, A. 4448 (C. erioclona) [K]; Loher, A. 4449 (C. erioclona) [K, US]; Loher, A. 6681 (C. erioclona) [K]; Loher, A. 7287 (C. erioclona) [K]; Lopez, G. Bur. Sci. 41342 (C. bicolor) [A, K]; Lynn Zwickey, A. 14 (C. bicolor) [A, NY].

Madulid PNH117871 (C. pentandra) [K]; Madulid, D. A. PPI3151 (C. caudata) [K]; Madulid, D. A. PPI11531 (C. bicolor) [K]; Madulid, Gaerlan PPI23846 (C. pedunculata) [GH, K]; Maliwanag, E. 104 (C. erioclona) [GH]; Maliwanag, E. 191 (C. pedunculata) [A]; Mangubat, L. Bur. Sci. 937 (C. pedunculata) [K, NY, US]; Mangubat, L. Bur. Sci. 937 (C. pedunculata) [NY]; Manuel, F. For. Bur. 23489 (C. caudata) [US]; McGregor, R. C. 1241 (C. bicolor) [NY, US]; McGregor, R. C. 113 (C. pedunculata) [K, NY, US]; McGregor, R. C. Bur. Sci. 22919 (C. pedunculata) [K, US]; McGregor, R. C. Bur. Sci. 1239 (C. pedunculata) [NY, US]; McGregor, R. C. Bur. Sci. 19792 (C. pedunculata) [US]; McGregor, R. C. Bur. Sci. 22019 (C. erioclona) [K]; Mcgregor, R. C. Bur. Sci. 22910 (C. erioclona) [US]; Mearns, E. A., Hutchinson, W. J. For. Bur. 4713 (C. aff. basilanensis) [NY]; Mearns, E. A., Hutchinson, W. J. For. Bur. 4755 (C. caudata) [NY]; Mendoza, D. R. PNH18424 (C. paloensis) [K]; Mendoza, D. R. PNH18195 (C. pedunculata) [K, US]; Mendoza, D. R. PNH18198 (C. pedunculata) [K]; Mendoza, D. R. PNH18303 (C. pentandra) [K]; Mendoza, D. R. PNH12216 (C. erioclona) [A]; Mendoza, D. R. PNH97782 (C. erioclona) [K]; Mendoza, D., Convocar, P. PNH10598 (C. aff. pentandra) [K,]; Mendum, M., Argent, G. C. G., Wilkie, P., Fuentes, R. 99260 (C. pedunculata) [A]; Merrill, E. D. 441 (C. bicolor) [A, K, NY, US]; Merrill, E. D. 2715 (C. bicolor) [A, K, NY, US]; Merrill, E. D. 3442 (C. bicolor) [K, NY, US]; Merrill, E. D. 1727 (C. caudata) [A]; Merrill, E. D. 4613 (C. caudata) [K, US]; Merrill, E. D. 4613 (C. caudata) [NY]; Merrill, E. D. 8115 (C. caudata) [K]; Merrill, E. D. 11689 (C. caudata) [A]; Merrill, E. D. 1667 (C. erioclona) [K, US]; Merrill, E. D. 145 (C. pedunculata) [US]; Merrill, E. D. 892 (C. pedunculata) [A, K, NY, US]; Merrill, E. D. 1214 (C. pedunculata) [K, NY, US]; Merrill, E. D. 1214 (C. pedunculata) [NY]; Merrill, E. D. 1718 (C. pedunculata) [A]; Merrill, E. D. 2522 (C. pedunculata) [A, K, NY, US]; Merrill, E. D. 2688 (C. pedunculata) [K, NY, US]; Merrill, E. D. 3375 (C. pedunculata) [K, NY]; Merrill, E. D. 3375 (C. pedunculata) [NY, US]; Merrill, E. D. 207 (C. bicolor) [US]; Merrill, E. D. 2245 (C. erioclona) [US]; Merrill, E. D. 2536 (C. erioclona) [A, K, US]; Merrill, E. D. 8057 (C. longifolia) [K]; Merrill, E. D. Species Blancoanae 637 (C. pedunculata) [A, K, NY, US]; Merrill, M. L. For. Bur. 8788 (C. angusta) [NY]; Merritt, M. L. For. Bur. 4078 (C. erioclona) [K]; Merritt, M. L. For. Bur. 4077 (C. pedunculata) [K]; Meyer, R. For. Bur. 2520 (C. pedunculata) [K, NY, US].

Ocampo, M. Bur. Sci. 27980 (C. pedunculata) [A, US]; Otanes, F. Bur. Sci. 17792 (C. micrantha) [A, US]; Otanes, F. Bur. Sci. 17851 (C. micrantha) [US]; Otanes, F. Bur. Sci. 17903 (C. pedunculata) [A, US].

Paniza, I. P. PNH9419 (C. pedunculata) [A]; Paraiso, F. Bur. Sci. 30197 (C. pedunculata) [NY]; Pascua, A. For. Bur. 30280 (C. basilanensis) [NY]; Peralta, A. PNH35478 (C. pedunculata) [K]; Ponce, S. For. Bur. 31291 (C. pentandra) [NY].

Quadras, J. F. 335 (C. erioclona) [NY, US]; Quisumbing, E. PNH2488 (C. pedunculata) [A]; Quisumbing, E., Sulit, M. Bur. Sci. 82430 (C. longipetiolata) [A]; Quisumbing, E., Sulit, M. Bur. Sci. 82319 (C. caudata) [A].

Ramos, M. Bur. Sci. 26976 (C. albidotomentella) [A, K]; Ramos, M. Bur. Sci. 1030 (C. angusta) [NY]; Ramos, M. Bur. Sci. 39455 (C. angusta) [NY, SING]; Ramos, M. Bur. Sci. 27041 (C. caudata) [A, US]; Ramos, M. Bur. Sci. 27388 (C. caudata) [K, US]; Ramos, M. Bur. Sci. 39816 (C. caudata) [A, US]; Ramos, M. Bur. Sci. 43310 (C. caudata) [A]; Ramos, M. Bur. Sci. 5790 (C. caudata) [US]; Ramos, M. Bur. Sci. 76891 (C. caudata) [A]; Ramos, M. Bur. Sci. 1063 (C. dolichophylla) [K, NY]; Ramos, M. Bur. Sci. 2037 (C. dolichophylla) [BM, NY, SING]; Ramos, M. Bur. Sci. 8268 (C. dolichophylla) [K]; Ramos, M. Bur. Sci. 23798 (C. erioclona) [K, US]; Ramos, M. Bur. Sci. 1039 (C. erioclona) [A, US]; Ramos, M. Bur. Sci. 30328 (C. longifolia) [K, NY, US]; Ramos, M. Bur. Sci. 13893 (C. micrantha) [K]; Ramos, M. Bur. Sci. 22310 (C. micrantha) [A, BM, US]; Ramos, M. Bur. Sci. 27662 (C. micrantha) [A, SING, US]; Ramos, M. Bur. Sci. 33146 (C. micrantha) [A, US]; Ramos, M. Bur. Sci. 43103 (C. micrantha) [A, SING]; Ramos, M. Bur. Sci. 43186 (C. micrantha) [A, SING]; Ramos, M. Bur. Sci. 23582 (C. paloensis) [K, NY]; Ramos, M. Bur. Sci. 77033 (C. pedunculata) [A]; Ramos, M. Bur. Sci. 32921 (C. pedunculata) [K]; Ramos, M. Bur. Sci. 17427 (C. pedunculata) [US]; Ramos, M. Bur. Sci. 1863 (C. pedunculata) [A, NY]; Ramos, M. Bur. Sci. 21755 (C. pedunculata) [US]; Ramos, M. Bur. Sci. 27610 (C. pedunculata) [US]; Ramos, M. Bur. Sci. 33019 (C. pedunculata) [NY]; Ramos, M. Bur. Sci. 42808 (C. pedunculata) [A]; Ramos, M. Bur. Sci. 43346 (C. pedunculata) [A]; Ramos, M. Bur. Sci. 80655 (C. pedunculata) [A]; Ramos, M. Bur. Sci. 80668 (C. pedunculata) [A]; Ramos, M. Bur. Sci. 8131 (C. pedunculata) [US]; Ramos, M. Bur. Sci. 1691 (C. pentandra) [NY]; Ramos, M. Bur. Sci. 14710 (C. pentandra) [K]; Ramos, M. Bur. Sci. 43349 (C. pentandra) [A, US]; Ramos, M. Bur. Sci. 41540 (C. ramiflora) [K, SING]; Ramos, M. Bur. Sci. 15278 (C. ramiflora) [K]; Ramos, M. Bur. Sci. 15278 (C. ramiflora) [BM, K]; Ramos, M. Bur. Sci. 30275 (C. ramiflora) [K]; Ramos, M. Bur. Sci. 41540 (C. ramiflora) [NY]; Ramos, M. Bur. Sci. 40778 (C. stenophylla) [A, K]; Ramos, M. Bur. Sci. 5739 (C. stenophylla) [K]; Ramos, M. Bur. Sci. 41009 (C. subalbida) [K]; Ramos, M. 1958 (C. angusta) [A]; Ramos, M. Bur. Sci. 39497 (C. angusta) [A]; Ramos, M. Bur. Sci. 41148 (C. bicolor) [A, US]; Ramos, M. Bur. Sci. 33012 (C. caudata) [NY]; Ramos, M. Bur. Sci. 7724 (C. caudata) [NY]; Ramos, M. 1901 (C. erioclona) [A]; Ramos, M. Bur. Sci. 33179 (C. erioclona) [NY]; Ramos, M. Bur. Sci. 40933 (C. erioclona) [A, K]; Ramos, M. Bur. Sci. 42770 (C. erioclona) [A]; Ramos, M. Bur. Sci. 8176 (C. erioclona) [K, US]; Ramos, M. Bur. Sci. 46442 (C. pedunculata) [NY]; Ramos, M. Bur. Sci. 7724 (C. pedunculata) [NY, US]; Ramos, M. Bur. Sci. 24383 (C. pentandra) [A]; Ramos, M. Bur. Sci. 40767 (C. subintegra) [A, US]; Ramos, M. & Convocar, P. Bur. Sci. 83993 (C. surigaensis) [A]; Ramos, M. & Convocar, P. Bur. Sci. 84082 (C. surigaensis) [A]; Ramos, M. & Edaño, G. Bur. Sci. 26367 (C. angusta) [A, US]; Ramos, M. & Edaño, G. Bur. Sci. 29506 (C. bicolor) [K]; Ramos, M. & Edaño, G. Bur. Sci. 84887 (C. bicolor) [A]; Ramos, M. & Edaño, G. Bur. Sci. 44601 (C. caudata) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 85264 (C. erioclona) [A]; Ramos, M. & Edaño, G. Bur. Sci. 37718 (C. subglandulosa) [K]; Ramos, M. & Edaño, G. Bur. Sci. 40363 (C. subglandulosa) [K]; Ramos, M. & Pascasio, J. Bur. Sci. 35185 (C. flavida) [NY]; Ramos, M. & Pascasio, J. Bur. Sci. 34697 (C. bicolor) [K]; Ramos, M. & Convocar, P. Bur. Sci. 83554 (C. pentandra) [A]; Ramos, M. & Convocar, P. Bur. Sci. 83762 (C. pentandra) [A]; Ramos, M. & Edaño, G. Bur. Sci. 34051 (C. angusta) [NY, SING]; Ramos, M. & Edaño, G. Bur. Sci. 46685 (C. angusta) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 46741 (C. angusta) [NY, SING]; Ramos, M. & Edaño, G. Bur. Sci. 36737 (C. basilanensis) [K, US]; Ramos, M. & Edaño, G. Bur. Sci. 31513 (C. bicolor) [A]; Ramos, M. & Edaño, G. Bur. Sci. 31537 (C. bicolor) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 48634 (C. bicolor) [A, K, NY]; Ramos, M. & Edaño, G. Bur. Sci. 49172 (C. bicolor) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 26312 (C. caudata) [A, US]; Ramos, M. & Edaño, G. Bur. Sci. 37388 (C. caudata) [A, K, US]; Ramos, M. & Edaño, G. Bur. Sci. 37481 (C. caudata) [A, US]; Ramos, M. & Edaño, G. Bur. Sci. 37635 (C. caudata) [A, US]; Ramos, M. & Edaño, G. Bur. Sci. 37679 (C. caudata) [A]; Ramos, M. & Edaño, G. Bur. Sci. 38035 (C. caudata) [A, US]; Ramos, M. & Edaño, G. Bur. Sci. 38850 (C. caudata) [A, K]; Ramos, M. & Edaño, G. Bur. Sci. 40505 (C. caudata) [A, US]; Ramos, M. & Edaño, G. Bur. Sci. 44895 (C. caudata) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 48504 (C. caudata) [A, K]; Ramos, M. & Edaño, G. Bur. Sci. 48504 (C. caudata) [K]; Ramos, M. & Edaño, G. Bur. Sci. 49295 (C. caudata) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 46779 (C. dolichophylla) [BM, NY]; Ramos, M. & Edaño, G. Bur. Sci. 26417 (C. erioclona) [A, US]; Ramos, M. & Edaño, G. Bur. Sci. 75120 (C. erioclona) [NY, US]; Ramos, M. & Edaño, G. Bur. Sci. 44064 (C. longifolia) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 44326 (C. longifolia) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 37563 (C. magnifolia) [K]; Ramos, M. & Edaño, G. Bur. Sci. 26233 (C. micrantha) [K]; Ramos, M. & Edaño, G. Bur. Sci. 26233 (C. micrantha) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 29137 (C. micrantha) [US]; Ramos, M. & Edaño, G. Bur. Sci. 29137 (C. micrantha) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 45614 (C. micrantha) [A]; Ramos, M. & Edaño, G. Bur. Sci. 49011 (C. micrantha) [A, SING, US]; Ramos, M. & Edaño, G. Bur. Sci. 49011 (C. micrantha) [BM, NY]; Ramos, M. & Edaño, G. Bur. Sci. 44297 (C. nigrescens) [K, NY]; Ramos, M. & Edaño, G. Bur. Sci. 45640 (C. pachyclada) [BM, K, NY]; Ramos, M. & Edaño, G. Bur. Sci. 28513 (C. pedunculata) [A, K]; Ramos, M. & Edaño, G. Bur. Sci. 29051 (C. pedunculata) [A, US]; Ramos, M. & Edaño, G. Bur. Sci. 29116 (C. pedunculata) [A]; Ramos, M. & Edaño, G. Bur. Sci. 29306 (C. pedunculata) [A,US]; Ramos, M. & Edaño, G. Bur. Sci. 33905 (C. pedunculata) [A, K, US]; Ramos, M. & Edaño, G. Bur. Sci. 44573 (C. pedunculata) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 44745 (C. pedunculata) [A, NY]; Ramos, M. & Edaño, G. Bur. Sci. 45067 (C. pedunculata) [A, NY]; Ramos, M. & Edaño, G. Bur. Sci. 45203 (C. pedunculata) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 47223 (C. pedunculata) [A, K, NY, US]; Ramos, M. & Edaño, G. Bur. Sci. 75187 (C. pedunculata) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 75187 (C. pedunculata) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 75283 (C. pedunculata) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 75712 (C. pedunculata) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 85196 (C. pedunculata) [A]; Ramos, M. & Edaño, G. Bur. Sci. 49510 (C. pentandra) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 31179 (C. pentandra) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 38748 (C. pentandra) [K]; Ramos, M. & Edaño, G. Bur. Sci. 44362 (C. pentandra) [A]; Ramos, M. & Edaño, G. Bur. Sci. 46928 (C. plumosa) [A, B, BM, K, NY, US]; Ramos, M. & Edaño, G. Bur. Sci. 75343 (C. ramiflora) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 28969 (C. subalbida) [BM, K]; Ramos, M. & Edaño, G. Bur. Sci. 45741 (C. subalbida) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 29707 (C. subintegra) [A, BM, K]; Ramos, M. & Edaño, G. Bur. Sci. 26465 (C. subintegra) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 26465 (C. subintegra) [K]; Ramos, M. & Edaño, G. Bur. Sci. 26619 (C. subintegra) [K]; Ramos, M. & Edaño, G. Bur. Sci. 26619 (C. subintegra) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 33838 (C. subintegra) [NY]; Ramos, M. & Edaño, G. Bur. Sci. 48530 (C. subintegra) [NY]; Ramos, M. & Pagasio, J. Bur. Sci. 34775 (C. caudata) [NY, SING]; Ramos, M. & Pascasio, J. Bur. Sci. 35123 (C. fasciculiflora) [K]; Ramos, M. & Pascasio, J. Bur. Sci. 35239 (C. pedunculata) [US,]; Ramos, M. & Pascasio, J. Bur. Sci. 34538 (C. pentandra) [K]; Ramos, M. & Pascasio, J. Bur. Sci. 34545 (C. surigaensis) [K, US, US]; Reillo, J. Bur. Sci. 15450 (C. basilanensis) [BM, K]; Reillo, J. Bur. Sci. 15450 (C. basilanensis) [K]; Reillo, J. Bur. Sci. 19265 (C. bicolor) [NY, US]; Reynoso, E. J. PNH87816 (C. erioclona) [K]; Reynoso, Alvarez & Fernando PPI6575 (C. pedunculata) [K]; Reynoso, Fuentes & Rivers PPI14203 (C. bicolor) [K]; Reynoso, Fuentes & Rivers PPI14120 (C. micrantha) [K]; Reynoso & Majaducon PPI21912 (C. pentandra) [K]; Reynoso & Majaducon PPI24171 (C. pentandra) [GH]; Reynoso & Majaducon PPI24996 (C. subglandulosa) [K]; Reynoso, Romero & Fuentes PPI1492 (C. erioclona) [K]; Reynoso, Sagcal & Fernando PPI17983 (C. bicolor) [GH]; Reynoso, Sagcal & Fernando PPI21095 (C. bicolor) [K]; Reynoso, Sagcal & Fernando PPI21443 (C. pedunculata) [GH, K]; Reynoso, Sagcal & Fernando PPI17892 (C. pentandra) [K]; Reynoso, Sagcal & Garcia PPI7322 (C. erioclona) [K]; Reynoso, Sagcal & Garcia PPI7109 (C. pedunculata) [K]; Ridsdale, C. E. 1051 (C. pentandra) [K]; Robinson, C. B. Bur. Sci. 11831 (C. basilanensis) [K]; Robinson, C. B. Bur. Sci. 11802 (C. caudata) [K, US]; Robinson, C. B. & Merrill, M. L. Bur. Sci. 6124 (C. bicolor) [NY]; Robinson, C. B. Bur. Sci. 6861 (C. platyphylla) [K]; Rogerson, C. T. 1016 (C. bicolor) [US]; Romero, E. M. & Chavez, R. V. PPI29251 (C. pedunculata) [K]; Romero & Fuentes PP137498 (C. bicolor) [K]; Romero & Fuentes PPI37557 (C. micrantha) [K]; Romero, Garcia & Majaducon PPI15345 (C. pedunculata) [GH, K]; Romero, Garcia & Majaducon PPI15432 (C. platyphylla) [A].

Sablaya, M. 95 (C. pentandra) [K]; Sandkuhl, H. For. Bur. 20428 (C. longipetiolata) [US]; Santos, J. K. Bur. Sci. 31659 (C. caudata) [A, K]; Santos, J. K. Bur. Sci. 31935 (C. longipetiolata) [K, SING]; Santos, J. K. Bur. Sci. 31935 (C. longipetiolata) [NY]; Santos, J. K. 11 (C. pedunculata) [K]; Santos, J. K. Bur. Sci. 31935 (C. longipetiolata) [A]; Sinclair, J. & Edaño, G. E. 9644 (?SFN) (C. pedunculata) [K,US]; Soejarto, D. A., Madulid, D. A., Fernando, O. & Majaducon, R. 8041 (C. pentandra) [A]; Soejarto, D. D., Madulid, D. A., Fernando, O. & Majaducon, R. 7997 (C. platyphylla) [A]; Soejarto, D. D., Gaerlan, F., Sagcal, E. & Fernando, O. 8321 (C. pentandra) [A]; Soejarto, D. D., Gaerlan, F., Sagcal, E. & Fernando, O. 8448 (C. pentandra) [A]; Steiner, M. L. 418 (C. erioclona) [US]; Stone, B. C. PPI811 (C. caudata) [K]; Stone, B. C. PPI6717 (C. micrantha) [K]; Stone, B. C. PPI6141 (C. pentandra) [K]; Stone, Fuentes & Fernando PPI5456 (C. pedunculata) [K]; Sulit, M. D. PNH15736 (C. caudata) [A]; Sulit, M. D. PNH8516 (C. caudata) [A]; Sulit, M. D. PNH14576 (C. erioclona) [A]; Sulit, M. D. PNH6232 (C. erioclona) [A, K]; Sulit, M. D. PNH14311 (C. pentandra) [A]; Sulit, M. D. PNH21651 (C. pentandra) [K]; Sulit, M. D. PNH2715 (C. pentandra) [K]; Sulit, M. D. PNH6094 (C. pentandra) [A]; Sulit, M. D. PNH6124 (C. pentandra) [A, K]; Sulit, M. D. PNH6344 (C. ramiflora) [K]; Sulit, M. D. PNH6406 (C. ramiflora) [K]; Sulit, M. D. PNH12342 (C. angusta) [A]; Sulit, M. D. PNH12444 (C. angusta) [A]; Sulit, M. D. PNH9744 (C. bicolor) [A]; Sulit, M. D. 6916 (C. erioclona) [A]; Sulit, M. D. PNH11744 (C. erioclona) [A]; Sulit, M. D. PNH14576 (C. erioclona) [K]; Sulit, M. D. PNH7706 (C. longipetiolata) [A, K]; Sulit, M. D. PNH7771 (C. longipetiolata) [A, K]; Sulit, M. D. PNH14405 (C. pachyclada) [K]; Sulit, M. D. & Conklin, H. C. PNH17699 (C. caudata) [A, K]; Sulit, M. D. & Conklin, H. C. PNH16953 (C. erioclona) [A]; Sulit, M. D. & Conklin, H. C. PNH16863 (C. pedunculata) [A]; Sulit, M. D. & Conklin, H. C. PNH17617 (C. pedunculata) [A, K]; Susara, J. B. PNH37323 (C. pedunculata) [K].

Tarrosa & Almagro For. Bur. 14928 (C. pentandra) [K]; Tesoro, B. T. PNH9603 (C. erioclona) [A].

Usteri, A. s.n. (C. bicolor) [NY].

Vanoverbergh, M. 1376 (C. longipetiolata) [A, NY]; Vidal 3431 (C. arborea) [K]; Vidal 3280 (C. dolichophylla) [K]; Vidal 3426 (C. dolichophylla) [K]; Vidal 1641 (C. micrantha) [GH]; Vidal 3239 (C. micrantha) [K]; Vidal 3428 (C. pedunculata) [GH, K]; Vidal 3430 (C. pentandra) [GH]; Vidal 3430 (C. pentandra) [GH]; Vidal 1643 (C. caudata) [GH, K]; Vidal 1644 (C. caudata) [GH, K]; Vidal 495 (C. angusta) [K]; Vidal 493 (C. bicolor) [K]; Vidal 493c (C. bicolor) [GH]; Vidal 3429 (C. caudata) [K]; Vidal 494 (C. erioclona) [K]; Vidal 843 (C. erioclona) [K]; Vidal 3427 (C. longifolia) [K]; Vidal 3425 (C. pedunculata) [K]; Vidal 3432 (C. pedunculata) [K].

Weber, C. M. 1055 (C. pedunculata) [GH, K, US]; Wenzel, C. A. 2701 (C. pentandra) [K]; Wenzel, C. A. 3389 (C. bicolor) [GH, K, NY]; Wenzel, C. A. 2849 (C. pentandra) [NY]; Wenzel, C. A. 2849 (C. pentandra) [K]; Wenzel, C. A. 1328 (C. bicolor) [GH, US]; Wenzel, C. A. 1479 (C. erioclona) [GH, US]; Wenzel, C. A. 609 (C. pentandra) [GH]; Wenzel, C. A. 690 (C. pentandra) [GH]; Wenzel, C. A. 1235 (C. pentandra) [GH]; Whitford, H. N. 487 (C. erioclona) [NY, US]; Whitford, H. N. 569 (C. bicolor) [US]; Whitford, H. N. For. Bur. 19757 (C. erioclona) [US]; Whitford, H. N. 404 (C. pedunculata) [K, NY, US]; Williams, R. S. 2307 (C. caudata) [A, K, NY, US]; Williams, R. S. 2964 (C. erioclona) [A, K, NY, US]; Williams, R. S. 2116 (C. longifolia) [K, NY]; Williams, R. S. 340 (C. micrantha) [A,US]; Williams, R. S. 115 (C. pedunculata) [K, NY, US]; Williams, R. S. 115 (C. pedunculata) [NY]; Williams, R. S. 1158 (C. pedunculata) [US]; Williams, R. S. 1003 (C. arborea) [K, NY]; Williams, R. S. 1060 (C. caudata) [NY, US]; Williams, R. S. 2577 (C. pedunculata) [NY]; Williams, R. S. 2577 (C. pedunculata) [NY].

Zwickey, A. L. 117 (C. pentandra) [A]; Zwickey, A. L. 287 (C. pentandra) [A]; Zwickey, A. L. 14 (C. bicolor) [A].

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Bramley, G.L.C. The genus Callicarpa (Lamiaceae) in the Philippines. Kew Bull 68, 369–418 (2013). https://doi.org/10.1007/s12225-013-9456-y

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