Journal of Mammalian Evolution

, Volume 23, Issue 4, pp 319–335 | Cite as

New Remains of the Multituberculate Mammal Barbatodon from the Upper Cretaceous of the Hațeg Basin (Romania)

  • Alexandru SolomonEmail author
  • Vlad Codrea
  • Márton Venczel
  • Mihai Dumbravă
  • Thierry Smith
Original Paper


The so-called “Hațeg Island” (Transylvania, Romania) is well known for its rich and peculiar Late Cretaceous vertebrate assemblages, including dwarf dinosaurs, first reported at the end of the 19th century. Besides dinosaurs, other important members of the “Hațeg Island” terrestrial ecosystems were the multituberculate mammals. Late Cretaceous (Maastrichtian) multituberculates are extremely rare in Europe, being known exclusively from the “Hațeg Island.” Two genera are reported from this area: Kogaionon, with the single species K. ungureanui, and Barbatodon, with two species: B. transylvanicus and B. oardaensis. Both genera belong to the European endemic family Kogaionidae. We report herein new remains of B. transylvanicus from the Maastrichtian locality of Pui (Hunedoara District) in the Hațeg Basin, including the most complete dentaries found to date with complete cheek teeth rows. Intraspecific variability of this species in dentary shape, tooth morphology, and size difference is highlighted.


Multituberculata Kogaionidae Intraspecific variability Maastrichtian Romania 



This research was supported by grant PN-II-PCE-2011-3-0381 of the National Council of Scientific Research - CNCS (A.S., V.C., M.V.), Projects MO/36/001-004 of the Belgian Science Policy Office (T.S.), the financial support of the Sectorial Operational Programme for Human Resources Development 2007–2013, co-financed by the European Social Fund, under the projects POSDRU/159/1.5/S/133391-“Doctoral and postdoctoral excellence programs for training highly qualified human resources for research in the fields of Life Sciences, Environment and Earth” (A.S.) and POSDRU/159/1.5/S/132400 with the title “Young successful researchers-professional development in an international and interdisciplinary environment” (M.D.), Sepkoski grant (A.S). The authors thank all people involved over a decade in the field missions in the “Hațeg Island,” too numerous to be mentioned here. Also, we thank the anonymous reviewers, as well as the Editor-in-chief John Wible, for their useful comments that greatly helped to improve the manuscript.

Supplementary material

10914_2016_9322_MOESM1_ESM.docx (17 kb)
Table S1 (DOCX 16.5 kb)
10914_2016_9322_MOESM2_ESM.docx (18 kb)
Table S2 (DOCX 17 kb)


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Copyright information

© Springer Science+Business Media New York 2016

Authors and Affiliations

  • Alexandru Solomon
    • 1
    Email author
  • Vlad Codrea
    • 1
  • Márton Venczel
    • 2
  • Mihai Dumbravă
    • 1
  • Thierry Smith
    • 3
  1. 1.Department of Geology, Faculty of Biology-GeologyBabeş-Bolyai UniversityCluj-NapocaRomania
  2. 2.Department of Natural HistoryŢării Crișurilor MuseumOradeaRomania
  3. 3.Directorate Earth and History of Life, Royal Belgian Institute of Natural SciencesBrusselsBelgium

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