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Long-term growth and yield effects of respacing natural regeneration of Sitka spruce in Britain

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Abstract

Stands of Sitka spruce (Picea sitchensis (Bong.) Carr.) originating from natural regeneration can be extremely dense with high mortality, poor growth form and low volume production. Although costly, respacing (pre-commercial thinning) can reduce resource competition for the remaining trees, altering the final crop produced. Three experiments were established to examine the effect of respacing 1–2 m tall trees to different target densities. The impact on stand growth and yield was measured 11–17 years after respacing, and the longer-term impact was modelled to year 50. Unrespaced control treatments self-thinned at a similar and constant rate at two sites. At the third, extremely high initial density resulted in a higher rate of self-thinning and lower density after 11 years. Wider respacing treatments resulted in larger mean tree diameter, but there was no significant effect of respacing on stand volume 11–17 years after respacing; greater diameter growth did not compensate for low tree number. Results indicate a volume penalty associated with delaying respacing until trees were 4 m tall, but this treatment was unreplicated. Modelled stand volume in year 50 was higher for the 2.6 m × 2.6 m respacing treatment than for the 1.8 m × 1.8 m, 2.1 m × 2.1 m or 3.3 m × 3.3 m treatments. However, open-grown conditions may reduce timber quality compared to closer spacing treatments. The results are presented using a stand density management diagram for Sitka spruce growing in Canada and support recent suggestions that British stands have a shallower size–density relationship than Canadian stands.

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References

  • Adam GA (1993) Mechanised respacing with tractor mounted horizontal shaft flails. Technical Development Branch Report 20/93. Technical Development, Ae, Dumfries

  • Barbour RJ, Bailey RE, Cook JE (1992) Evaluation of relative density, diameter growth and stem form in a red spruce (Picea rubens) stand 15 years after precommercial thinning. Can J For Res 22:229–238

    Article  Google Scholar 

  • Brazier JD, Mobbs ID (1993) The influence of planting distance on the structural wood yields of unthinned Sitka spruce. Forestry 66:332–353

    Article  Google Scholar 

  • Cameron AD (2002) Importance of early selective thinning in the development of long-term stand stability and improved log quality: a review. Forestry 75:25–35

    Article  Google Scholar 

  • Clarke GC (1992) The natural regeneration of spruce. Scott For 46:107–129

    Google Scholar 

  • Claveau Y, Messier C, Comeau PG, Coates KD (2002) Growth and crown morphological responses of boreal conifer seedlings and saplings with contrasting shade tolerance to a gradient of light and height. Can J For Res 32:458–468

    Article  Google Scholar 

  • Comeau PG, White M, Kerr G, Hale SE (2010) Maximum size–density relationships for Sitka spruce and coastal Douglas-fir in Britain and Canada. Forestry 83(5):461–468

    Article  Google Scholar 

  • Deans JD, Milne R (1999) Effects of respacing on young Sitka spruce crops. Forestry 72(1):47–57

    Article  Google Scholar 

  • Drew TJ, Flewelling JW (1979) Stand density management: an alternative approach and its application to Douglas fir plantations. For Sci 25:518–532

    Google Scholar 

  • Edwards PN, Christie JM (1981) Yield models for forest management. Forestry Commission booklet 48. HMSO, London

  • Forestry Commission (1992) Motor manual respacing of naturally regenerated Sitka spruce in Wales. Technical Development Branch Report 3/92. Technical Development, Machynlleth, Powys

  • Gardiner BA, Stacey GR, Belcher RE, Wood CJ (1997) Field and wind tunnel assessments of the effects of respacing on tree stability. Forestry 70:233–252

    Article  Google Scholar 

  • Gardiner B, Suarez J, Achim A, Hale S, Nicoll B (2006) ForestGALES—a PC-based wind risk model for British forests. User’s guide. Version 2.1. Forestry Commission, Edinburgh, p 54

    Google Scholar 

  • Gardiner B, Leban J-M, Auty D, Simpson H (2011) Models for predicting wood density of British-grown Sitka spruce. Forestry 84(2):119–132

    Article  Google Scholar 

  • Harding J (1993) Motor manual respacing by clearing saw. Technical Development Branch Report 17/93. Technical Development, Ae, Dumfries

  • Harding J (1996) Mechanical respacing by Bruunett/Berti flail. Technical Development Branch Report 6/94. Technical Development, Ae, Dumfries

  • Kennedy F (2002) The identification of soils for forest management. Forestry Commission field guide. Forestry Commission, Edinburgh, p 56

    Google Scholar 

  • Kobe RK, Coates RD (1997) Models of sapling mortality as a function of growth to characterise interspecific variation in shade tolerance of eight tree species of north western British Columbia. Can J For Res 27:227–236

    Article  Google Scholar 

  • Lieffers V, Messier C, Stadt KJ, Gendron F, Comeau PG (1999) Predicting and managing light in the understory of boreal forests. Can J For Res 29:796–811

    Article  Google Scholar 

  • Macdonald E, Hubert J (2002) A review of the effects of silviculture on timber quality of Sitka spruce. Forestry 75:107–138

    Article  Google Scholar 

  • Manley A, Richardson J (1995) Silviculture and economic benefits of producing wood energy from conventional forestry systems and measures to mitigate negative impacts. Biomass Bioenergy 9:89–105

    Article  Google Scholar 

  • Mason WL (2008) Natural regeneration of Sitka spruce in the Forest of Ae: development over 25 years. Scott For 62:2–8

    Google Scholar 

  • Mason WL (2010) Respacing naturally regenerating Sitka spruce and other conifers. Forestry Commission practice note 16. Forestry Commission, Edinburgh

  • Matthews RW, Mackie ED (2006) Forest mensuration: a handbook for practitioners. Forestry Commission, Edinburgh

    Google Scholar 

  • McNeill JD, Thompson DA (1982) Natural regeneration of Sitka spruce in the Forest of Ae. Scott For 36:269–282

    Google Scholar 

  • Messier C, Doucet R, Ruel J-C, Claveau Y, Kelly C, Lechowicz MJ (1999) Functional ecology of advance regeneration in relation to light in boreal forests. Can J For Res 29:812–823

    Article  Google Scholar 

  • Monserud RA, Ledermann T, Sterba H (2005) Are self-thinning constraints needed in a tree-specific mortality model? For Sci 50(6):848–858

    Google Scholar 

  • Moore J, Achim A, Lyon A, Mochan S, Gardiner B (2009) Effects of early re-spacing on the physical and mechanical properties of Sitka spruce structural timber. For Ecol Manag 258:1174–1180

    Article  Google Scholar 

  • Nelson DG (1991) Management of Sitka spruce natural regeneration. Forestry Commission research information note 204. Forestry Commission, Edinburgh

  • Nixon CJ, Worrell R (1999) Natural regeneration of conifers in Britain. Forestry Commission bulletin 120. HMSO, London

  • O’ Hara KL (1998) Silviculture for structural diversity: a new look at multi-aged stands. J For 96:4–10

    Google Scholar 

  • Ogawa K (2005) Time-trajectory of mean phytomass and density during a course of self-thinning in a sugi (Cryptomeria japonica D. Don) plantation. For Ecol Manag 214:104–110

    Article  Google Scholar 

  • Payne RW (ed) (2005) The guide to GenStat release 8.1. Part 2: statistics. Lawes Agricultural Trust (Rothamsted Experimental Station), Oxford

    Google Scholar 

  • Peet KR, Christensen NL (1987) Competition and tree death. Bioscience 37(8):586–595

    Article  Google Scholar 

  • Peterson EB, Peterson NM, Weetman GF, Martin PJ (1997) Ecology and management of Sitka spruce, emphasising its natural range in British Columbia. UBC Press, Vancouver

    Google Scholar 

  • Pothier D (2002) Twenty-year results of precommercial thinning in a balsam fir stand. For Ecol Manag 168:177–186

    Article  Google Scholar 

  • Pretzsch H (2002) A unified law of spatial allometry for woody and herbaceous plants. Plant Biol 4:159–166

    Article  Google Scholar 

  • Reineke LH (1933) Perfecting a stand-density index for even-aged forests. J Agric Res 46:627–638

    Google Scholar 

  • Reynolds B (2004) Continuous cover forestry: possible implications for surface water acidification in the UK uplands. Hydrol Earth Syst Sci 8:306–313

    Article  CAS  Google Scholar 

  • Roberts G, Crane SB (1997) The effects of clear felling established forestry on stream-flow losses from the Hore sub-catchment. Hydrol Earth Syst Sci 1:477–482

    Article  Google Scholar 

  • Rollinson TJ (1988) Respacing Sitka spruce. Forestry 61:1–22

    Article  Google Scholar 

  • Ruha T, Varmola M (1997) Precommercial thinning in naturally regenerated Scots pine stands in northern Finland. Silva Fenn 31(4):401–415

    Google Scholar 

  • Talbot B, Suadicani K (2005) Analysis of two simulated in-field chipping and extraction systems in spruce thinnings. Biosyst Eng 91(3):283–292

    Article  Google Scholar 

  • Ulvcrona KA, Claesson S, Sahlén K, Lundmark T (2007) The effects of timing of pre-commercial thinning and stand density on stem form and branch characteristics of Pinus sylvestris. Forestry 80(3):323–325

    Article  Google Scholar 

  • Varmola M, Salminen H (2004) Timing and intensity of precommercial thinning in Pinus sylvestris stands. Scand J For Res 19(2):142–151

    Article  Google Scholar 

  • von Ow F, Joyce P, Keane M (1996) Factors affecting the establishment of natural regeneration Sitka spruce (Picea sitchensis (Bong.) Carr.) in Ireland. Ir For 53(1):2–18

    Google Scholar 

  • Weiskittel A, Gould P, Temesgen H (2009) Sources of variation in the self-thinning boundary line for three species with varying levels of shade tolerance. For Sci 55(1):84–93

    Google Scholar 

  • Weller DE (1987) A re-evaluation of the—3/2 power rule of self-thinning. Ecol Monogr 57:23–43

    Article  Google Scholar 

  • Williams H, Messier C, Kneeshaw DD (1999) Effects of light availability and sapling size on the growth and crown morphology of understory Douglas-fir and lodgepole pine. Can J For Res 28:222–231

    Article  Google Scholar 

  • Wright EF, Canham CD, Coates KD (2000) Effects of suppression and release on sapling growth for 11 tree species of northern, interior British Columbia. Can J For Res 30:1571–1580

    Article  Google Scholar 

  • Yoda K, Kira T, Ogawa H, Hozumi K (1963) Self thinning in overcrowded pure stands under cultivated and natural conditions. J Biol Osaka City Univ 14:107–129

    Google Scholar 

  • Zeide B (1985) Tolerance and self-tolerance of trees. For Ecol Manag 13:149–166

    Article  Google Scholar 

  • Zeide B (1987) Analysis of the 3/2 power law of self-thinning. For Sci 33(2):517–537

    Google Scholar 

Download references

Acknowledgments

The experiments were designed and established by D. G. Nelson and later maintained and managed by Chris Nixon and Bill Mason. Operations, maintenance and assessments were carried out by Forest Research Technical Services Staff. The M1 model was developed from J. Christie’s yield models by R. Matthews, C. Arcangeli and P. Henshall of Forest Research. The SDMD for planted Sitka spruce was developed by Craig Farnden working under contract to the British Columbia Ministry of Forests and is based on data from the managed stand yield tables produced by TIPSY (www.for.gov.bc.ca/hre/gymodels/SDMD/model.htm). Andy Peace provided statistical advice, and Tom Jenkins provided helpful comments on an earlier draft. The authors are grateful for helpful comments made by two anonymous reviewers.

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The authors declare that they have no conflict of interest.

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Correspondence to Victoria Stokes.

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Communicated by A. Weiskittel.

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Stokes, V., Kerr, G. Long-term growth and yield effects of respacing natural regeneration of Sitka spruce in Britain. Eur J Forest Res 132, 351–362 (2013). https://doi.org/10.1007/s10342-012-0679-5

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  • DOI: https://doi.org/10.1007/s10342-012-0679-5

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