Lefua tokaiensis, a new species of nemacheilid loach from central Japan (Teleostei: Nemacheilidae)

  • Taiki ItoEmail author
  • Kazumi Hosoya
  • Jun-Ichi Miyazaki
Full Paper


A new nemacheilid loach, Lefua tokaiensis sp. nov., is described from the small mountain streams in the Tokai region, central Honshu, Japan. Lefua tokaiensis is distinguished from all other species of Lefua by the following combination of characters: eyes located dorsally on the head, presence of a narrow conspicuous longitudinal mark between the base of the outer rostral barbel and the eye, absence of a rhomboid or triangular dark blotches on the middle of the caudal fin base, absence of a black longitudinal stripe on both body sides in mature males, absence of dusky cross bars on the dorsal area of the body, absence of a dusky bar beside the dorsal fin base, typical presence of small dark spots on the dorsal and caudal fins, presence of the obscure dark band in skin of the caudal fin base, caudal fin slightly squared off at the apex, and modally four hypurals.


Endangered species Head water Parallel evolution Tokai region 



We are grateful to M. Sugiura, T. Asaka, T. Morishita, and M. Takemoto for collecting several specimens and data to clarify the total variance of L. tokaiensis, and M. Matsushita for drawings of the holotype of L. tokaiensis. We sincerely thank M. Matsunuma (KUN) for observations of materials under his care.


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Copyright information

© The Ichthyological Society of Japan 2019

Authors and Affiliations

  1. 1.Wetlands International JapanTokyoJapan
  2. 2.Department of Environmental Management, Faculty of AgricultureKindai UniversityNaraJapan
  3. 3.Faculty of EducationUniversity of YamanashiKofuJapan
  4. 4.Tokushima Prefectural MuseumTokushimaJapan

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