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A new species of Timorichthys (Ophidiiformes: Bythitidae) from the East China Sea


A new bythitid fish, Timorichthys angustus, is described on the basis of a single specimen (52 mm in standard length, SL) collected from the East China Sea, Japan. The genus Timorichthys Nielsen and Schwarzhans 2011 was only known from the holotype of the type species Timorichthys disjunctus Nielsen and Schwarzhans 2011 caught in the Timor Sea. The new species differs from T. disjunctus in several characters, such as the pectoral fin peduncle being much longer than its width (vs. equal in T. disjunctus), precaudal vertebrae 22 (vs. 16), total vertebrae 62 (vs. 52), long rakers on the first gill arch 14 (vs. 6), horizontal eye diameter 3.1 % SL (vs. 1.3 % SL), interorbital width 0.8 % SL (vs. 3.3 % SL), posterior mandibular pore absent (vs. present), anterior infraorbital pores 1 (vs. 3), opercular spine not covered by skin (vs. covered by skin), otolith height to thickness 2.5 (vs. 1.8), and head and body light brown in alcohol (vs. dark brown).

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  • Nielsen JG, Schwarzhans W (2011) A new genus and species of Bythitidae (Teleostei: Ophidiiformes) from northwestern Australia. Beagle Rec Mus Art Galler North Territ 27:143–146

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  • Nielsen JG, Cohen DM, Markle DF, Robins CR (1999) Ophidiiform fishes of the world (order Ophidiiformes). FAO species catalogue, vol 18. FAO, Rome

  • Schwarzhans W, Møller PR, Nielsen JG (2005) Review of the Dinematichthyini (Teleostei: Bythitidae) of the Indo-West Pacific. Part I. Diancistrus and two new genera with 26 new species. Beagle Rec Mus Art Galler North Territ 21:73–163

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We are sincerely grateful to the captain, officers, and crew of R/V Kumamoto-maru (Kumamoto Reiyou High School), M. Fukuwaka, M. Yoda, H. Kurota, R. Yukami, K. Hidaka (Seikai National Fisheries Research Institute), M. Kawazu (Kyusyu University), and K. Minotani (Tokyo University of Marine Science and Technology) for their assistance in sampling. We also thank D. G. Smith (Smithsonian Institution) and K. Hoshino (Seikai National Fisheries Research Institute) for their critical reading of this manuscript, M. A. Krag (Natural History Museum of Denmark) for producing the photographs, and H. Senou (Kanagawa Prefectural Museum of Natural History) and M. McGrouther (Australian Museum) for their advice in this study. This cruise was funded by the Fisheries Agency of Japan.

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Correspondence to Jørgen G. Nielsen.

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This article was registered in the Official Register of Zoological Nomenclature (ZooBank) as 02B45AD9-B2D2-4F10-B9FF-D1A3B71E053C.

This article was published as an Online First article on the online publication date shown on this page. The article should be cited by using the doi number.

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Nielsen, J.G., Okamoto, M. & Schwarzhans, W. A new species of Timorichthys (Ophidiiformes: Bythitidae) from the East China Sea. Ichthyol Res 60, 237–240 (2013).

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  • Ophidiiformes
  • Bythitinae
  • Timorichthys
  • Japan
  • East China Sea
  • New species