El-Domyati M, Attia S, Saleh F, Brown D, Birk DE, Gasparro F, Ahmad H, Uitto J. Intrinsic aging vs. photoaging: a comparative histopathological, immunohistochemical, and ultrastructural study of skin. Experi. Dermatol. 11: 398–405 (2002).
Kuhn A, Zahn S, Patsinakidis N, Landmann A, Graef M, Sauerland C, Surber C, Wenzel J. Resistance to water and abrasion of a broad-spectrum sunscreen: a prospective, open-label study. Experi. Dermatol. 25: 151–152 (2016).
Bickers DR, Athar M. Oxidative stress in the pathogenesis of skin disease. J. Investig. Dermatol. 126: 2565–2575 (2006).
Passeron T, Ortonne J. Skin ageing and its prevention. Press. Medica. 32: 1474–1482 (2003).
Heng MC. Signaling pathways targeted by curcumin in acute and chronic injury: burns and photo-damaged skin. Int. J. Dermatol. 52: 531–543 (2013).
Lu J, Guo JH, Tu XL, Zhang C, Zhao M, Zhang QW, Gao FH. Tiron inhibits UVB-Induced AP-1 binding sites transcriptional activation on MMP-1 and MMP-3 promoters by MAPK signaling pathway in human dermal fibroblasts. PloS ONE 11: e0159998 (2016).
Watson RE, Gibbs NK, Griffiths CE, Sherratt MJ. Damage to skin extracellular matrix induced by UV exposure. Antioxid. Redox. Signal. 21: 1063–1077 (2014).
Chen B, Li R, Yan N, Chen G, Qian W, Jiang HL, Ji C, Bi ZG. Astragaloside IV controls collagen reduction in photoaging skin by improving transforming growth factor-β/Smad signaling suppression and inhibiting matrix metalloproteinase-1. Mol. Med. Rep. 11: 3344–3348 (2015).
Quan T, He T, Kang S, Voorhees JJ, Fisher GJ. Solar ultraviolet irradiation reduces collagen in photoaged human skin by blocking transforming growth factor-β type II receptor/Smad signaling. Am. J. Pathol. 165: 741–751 (2004).
Gilmore TD. Introduction to NF-κB: players, pathways, perspectives. Oncogene 25: 6680–6684 (2006).
Perkins ND. Integrating cell-signalling pathways with NF-κB and IKK function. Nat. Rev. Mol. Cell Biol. 8: 49–62 (2007).
Carmeliet P, Jain RK. Molecular mechanisms and clinical applications of angiogenesis. Nature 473: 298–307 (2011).
Oh Y, Lim H-W, Kim K, Lim C-J. Ginsenoside Re improves skin barrier function in HaCaT keratinocytes under normal growth conditions. Biosci. Biotehcnol. Biochem 80: 2165–2167 (2016).
Rinnerthaler M, Duschl J, Steinbacher P, Salzmann M, Bischof J, Schuller M, Wimmer H, Peer T, Bauer JW, Richter K. Age-related changes in the composition of the cornified envelope in human skin. Experi. Dermatol. 22: 329–335 (2013).
Rajan NS, Bhat R. Antioxidant compounds and antioxidant activities in unripe and ripe kundang fruits (Bouea macrophylla Griffith). Fruits 71: 41–47 (2016).
Heck DE, Vetrano AM, Mariano TM, Laskin JD. UVB light stimulates production of reactive oxygen species unexpected role for catalase. J. Biol. Chem. 278: 22432–22436 (2003).
Park JE, Pyun HB, Woo SW, Jeong JH, Hwang JK. The protective effect of Kaempferia parviflora extract on UVB-induced skin photoaging in hairless mice. Photodermatol. Photoimmunol Photomed. 30: 237–245 (2014).
Anggakusuma, Yanti, Hwang JK. Effects of macelignan isolated from Myristica fragrans Houtt. on UVB-induced matrix metalloproteinase-9 and cyclooxygenase-2 in HaCaT cells. J. Dermatol. Sci. 57: 114–122 (2010).
Draelos ZD. The latest cosmeceutical approaches for anti-aging. J. Cosmet. Dermatol. 6: 2–6 (2007).
Kim YJ, Kim HN, Shin MS, Choi BT. Thread embedding acupuncture inhibits ultraviolet B irradiation-induced skin photoaging in hairless mice. Evid. Based Complement. Alternat. Med. 2015 (2015).
Shirakata Y. Regulation of epidermal keratinocytes by growth factors. J. Dermatol. Sci. 59: 73–80 (2010).
Kim HK. Garlic supplementation ameliorates UV-induced photoaging in hairless mice by regulating antioxidative activity and MMPs expression. Molecules 21: 70 (2016).
Zhan JYX, Wang XF, Liu YH, Zhang ZB, Wang L, Chen JN, Huang S, Zeng HF, Lai XP. Andrographolide sodium bisulfate prevents UV-induced skin photoaging through inhibiting oxidative stress and inflammation. Mediators Inflamm. 2016: 1–12 (2016).
Pillai S, Oresajo C, Hayward J. Ultraviolet radiation and skin aging: roles of reactive oxygen species, inflammation and protease activation, and strategies for prevention of inflammation-induced matrix degradation—a review. Int. J. Cosmet. Sci. 27: 17–34 (2005).
Tanaka YT, Tanaka K, Kojima H, Hamada T, Masutani T, Tsuboi M, Akao Y. Cynaropicrin from Cynara scolymus L. suppresses photoaging of skin by inhibiting the transcription activity of nuclear factor-kappa B. Bioorg. Med. Chem. Lett. 23: 518–523 (2013).
Amano S, Ogura Y, Akutsu N, Matsunaga Y, Kadoya K, Adachi E, Nishiyama T. Protective effect of matrix metalloproteinase inhibitors against epidermal basement membrane damage: skin equivalents partially mimic photoageing process. Br. J. Dermatol. 153: 37–46 (2005).
Chen T, Hou H, Lu J, Zhang K, Li B. Protective effect of gelatin and gelatin hydrolysate from salmon skin on UV irradiation-induced photoaging of mice skin. J. Ocean U. China 15: 1–8 (2016).
Nagase H, Visse R, Murphy G. Structure and function of matrix metalloproteinases and TIMPs. Cardiovasc. Res. 69: 562–573 (2006).
Page-McCaw A, Ewald AJ, Werb Z. Matrix metalloproteinases and the regulation of tissue remodelling. Nat. Rev. Mol. Cell Biol. 8: 221–233 (2007).
Pittayapruek P, Meephansan J, Prapapan O, Komine M, Ohtsuki M. Role of matrix metalloproteinases in photoaging and photocarcinogenesis. Int. J. Mol. Sci. 17: 868 (2016).
Kim MS, Oh GH, Kim MJ, Hwang JK. Fucosterol inhibits matrix metalloproteinase expression and promotes type-1 procollagen production in UVB-induced HaCaT cells. Photochem. Photobiol. 89: 911–918 (2013).
Candi E, Schmidt R, Melino G. The cornified envelope: a model of cell death in the skin. Nat. Rev. Mol. Cell Biol. 6: 328–340 (2005).
Yamane M, Sugimura K, Kawasaki H, Tatsukawa H, Hitomi K. Analysis on transglutaminase 1 and its substrates using specific substrate peptide in cultured keratinocytes. Biochem. Biophys. Res. Commun. 478: 343–348 (2016).
Kim H, Lim YJ, Park JH, Cho Y. Dietary silk protein, sericin, improves epidermal hydration with increased levels of filaggrins and free amino acids in NC/Nga mice. Br. J. Nutr. 108: 1726–1735 (2012).
Lai CH, Chang SC, Chen YJ, Wang YJ, Lai YJ, Chang HD, Berens EB, Johnson MD, Wang JK, Lin CY. Matriptase and prostasin are expressed in human skin in an inverse trend over the course of differentiation and are targeted to different regions of the plasma membrane. Biol. Open 5: 1380–1387 (2016).
Descargues P, Sil AK, Sano Y, Korchynskyi O, Han G, Owens P, Wang XJ, Karin M. IKKα is a critical coregulator of a Smad4-independent TGFβ-Smad2/3 signaling pathway that controls keratinocyte differentiation. Proc. Natl. Acad. Sci. 105: 2487–2492 (2008).
Wu N, Sulpice E, Obeid P, Benzina S, Kermarrec F, Combe S, Gidrol X. The miR-17 family links p63 protein to MAPK signaling to promote the onset of human keratinocyte differentiation. PloS ONE, 7: e45761 (2012).