Genetic dissection of fruit quality traits in the octoploid cultivated strawberry highlights the role of homoeo-QTL in their control

Table 2 Significant QTL detected for all the traits based on CIM analysis with LOD > LOD threshold (2.86 for the female map and 2.90 for the male map)

Traits^a	No. QTL^b	Location of QTL^c (nb of year of detection)^d
Development
FD	4	IIb-m (2), IIIa-m (2), IIIc1-m (1), IVa-m (1)
FL	7	IIa-f (1), IIc-m (3), IIIa-m (1), IIId-m (1), IVd2-f (1), Vb-f (1), VIb-m (1)
FD/FL	5	IIIa-m (3)/f (2), IIId-m (2), Va-m (2), Vb-m (1), VIa-m (2)
DW	3	IIIa-f (1), VIa-m (1), VIIb-m (1)
FW	3	Ia-f (1), IIb-m (2), IIIc1-m (1)
RD	6	IIb-m (1), IIId-f (1), IVc-m (1), Vc-m (1), VIb-m (1), VIIa-m (2)
Firmness
FIRM	7	IIb-m (1), IIIa-m (1), IIIa-f (1), IIIb-m (1), IIIc-f (1), IVa-f (1), Vc-m/f (1)
Colour
a	6	Ia-f (1), IIa-f (3), IVd-m (1), Vb-m (1), VIa-m/f (2), VIb-f (1)
b	4	Ia-m (1), IIIa-f (1), VIa-m/f (2), VIb-f (1)
L	4	IIa-f (1), IIIa-m/f (2), IVd-m (1), VIa-m (2)
ANTH	5	Ia-f (1), IIa-f (2), IIIa-f (1), VIa-m/f (2), VIb-f (1)
Sugar-related traits
SSC	3	IIIa-m/f (1), Va-f (1), VIa-m/f (2)
FRU	2	IVb-m (1), Vc-f (1)
GLU	5	Ib-f (1), IIIa-f (1), IIIc-f (1), IVa-f (1), VIa-f (1)
SUCR	2	VIa-m (1), VIIa-f (2)
Acid-related traits
pH	6	Ia-m (1), IIIb-m (1), IIIc-f (1), IVc-f (1), Vb-f (1), VIa-f (1)
TA	5	Ia-m (1), IIc-m (1), IIId-m (1), IVa-f (1), Vb-m/f (2)
CIT	6	Ia-f (1), IIc-m (1), IIIa-f (1), Vb-f (1), VIa-f (1), VIIb-f (1)
MAL	4	Vd-f (1), VIa-m (1), VIb-m (1), VIId-m (1)

^aTraits are the following: development: FD, fruit diameter; FL, fruit length; FD/FL, ratio; DW, dry weight; FW, fruit weight; RD, ripening date of the fruit. Texture: FIRM, fruit firmness. Colour: skin colour parameters, a, b and L; ANTH, total anthocyanin. Sugar-related traits: SSC, soluble-solids content; FRU, fructose; GLU, glucose; SUCR, sucrose. Acid-related traits: pH; TA, titratable acidity; CIT, citrate; MAL, malate
^bNumber of unique QTL. QTL were considered unique when, the same regions overlapped with one LOD support confidence interval
^cName of the linkage group according to Rousseau-Gueutin et al. (2008). The Roman number indicates the homoeologous group number. The letter, a, b c, or d, indicates one of the four homoeologous linkage groups and the final letter, m or f, indicates if the linkage group belongs to the male or the female linkage maps
^dIn brackets, the number of detection years of the individual QTL