Journal of Plant Research

, Volume 108, Issue 4, pp 417–428 | Cite as

Tsuga shimokawaensis, a new species of permineralized conifer leaves from the Middle Miocene Shimokawa Group, Hokkaido, Japan

  • Midori Matsumoto
  • Takeshi Ohsawa
  • Makoto Nishida
Original Articles


Silicified pinaceous leaves from the late Middle Miocene bed of Shimokawa Town, central Hokkaido, Japan are described asTsuga shimokawaensis sp. nov. Comparisons with leaves of extant species ofTsuga show that the new species is assigned to sect.Tsuga, resembling the extant speciesT. heterophylla in having usually one-cell-layered hypodermis, andT. sieboldii in having mesophyll cells between the resin canal and the hypodermis. From cladistic analyses we infer thatTsuga shimokawaensis is positioned the basal in sect.Tsuga and that sect.Tsuga comprises four species groups.

Key words

Fossil leaf Middle Miocene Pinaceae Tsuga shimokawaensis 


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Copyright information

© The Botanical Society of Japan 1995

Authors and Affiliations

  • Midori Matsumoto
    • 1
  • Takeshi Ohsawa
    • 2
  • Makoto Nishida
    • 3
  1. 1.Department of Earth Sciences, Faculty of ScienceChiba UniversityChibaJapan
  2. 2.Laboratory of Phylogenetic Botany, Faculty of ScienceChiba UniversityChibaJapan
  3. 3.Research Institute of Evolutionary BiologyTokyoJapan

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