Summary
Timothy hays grown with intensive soil treatments, including trace elements, (without significant differences in yields of hay per acre), were put under bioassay by weanling rabbits, only to have the high temperatures of a summer heat wave demonstrate by deaths of these test animals, regardless of soil treatments, that the protein intake (and what might accompany it) was not equal to the animal's needs. No deaths resulted in the stock rabbits in the same environment, fed on the same grain but with green grass in addition. Also, ten grams of skim-milk powder per rabbit per day prevented the death of the test rabbits as the extended heat wave permitted the demonstration. Similarly red clover hay, a replacement of the timothy, demonstrated that it also provided the protein needs to prevent deaths of the rabbits by the heat. According to the bioassays, the timothy suggests its own classification as of low potential as animal nutrition, even under extensive soil treatments growing it.
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Albrecht, W.A., de Jonge, J. Biosynthesis of amino acids according to soil fertility. Plant Soil 18, 298–308 (1963). https://doi.org/10.1007/BF01347230
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DOI: https://doi.org/10.1007/BF01347230