, Volume 5, Issue 4, pp 325–333 | Cite as

Untersuchungen über die Jahresrhythmik von Laufkäfern (Coleoptera, Carabidae) in der Cyrenaika (Libyen, Nordafrika)

  • Wilfried Paarmann

Research on the annual periodicity of carabid beetles in the Cyrenaika (North Africa)


In a 30 km deep coastal area of the Cyrenaika (Libya) a total of 28 species of carabids was collected on regular excursions in 1967 and 1968. In midsummer almost no carabids were caught. They appeared at the beginning of autumn and were caught in winter regularly in equal numbers. During March, April and May the number of catches increased considerably. Using ground traps no more than two species (Carterus silvestrii, Pristonychus picicornis) were caught and that only from December to May. From the pattern of their occurrence we can conclude that in the Cyrenaika carabid beetles propagate mainly during the winter.

By examining the gonads, catching larvae and breeding, it was possible to prove that the speciesBroscus laevigatus, Harpalus litigiosus ssp.cyrenaicus andPristonychus picicornis propagate in winter. Examination of the gonads also indicates a winter propagation of the speciesPterostichus barbarus andLaemostenus complanatus ssp.cyrenaicus. It is not certain, however, thatCarterus silvestrii propagates during the winter.

The speciesCalathus fuscipes andCalathus mollis from the temperate northern regions, which are known as autumn breeders, were caught in Libya too, which indicates a relation between the autumn breeders of Middle and Northern Europe and the ‘winter breeders’ of the South Mediterranean countries. The presence ofPogonus chalceus in Libya strongly suggests that, in certain biotopes, spring breeders may develop besides the dominant autumn breeders.


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  1. Bodenheimer, F. S.: Studies on the ecology of Palestinean Coleoptera. II. Seasonal and diurnal appearance and activity. Bull. Soc. Royale Entom. d'Egyte18, 211–241 (1934).Google Scholar
  2. Larsson, S. G.: Entwicklungstypen und Entwicklungszeiten der dänischen Carabiden. Entom. Meddelels.20, 275–560 (1939).Google Scholar
  3. Paarmann, W.: Vergleichende Untersuchungen über die Bindung zweier Carabidenarten (P. angustatus Dft. undP. oblongopunctatus F.) an ihre verschiedenen Lebensräume. Z. wiss. Zool.174, 83–176 (1966).Google Scholar
  4. Thiele, H. U.: Experimentelle Untersuchungen über die Ursachen der Biotopbindung bei Carabiden. Z. Morph. Ökol. Tiere53, 387–452 (1964).Google Scholar
  5. Thiele, H. U.: Formen der Diapausesteuerung bei Carabiden. Verh. Dtsch. Zool. Ges. (1967) S. 358–364 (1968).Google Scholar
  6. —: The control of larval hibernation and of adult aestivation in the carabid beetlesNebria brevicollis F. andPatrobus atrorufus Stroem. Oecologia (Berl.)2, 347–361 (1969).Google Scholar
  7. —, Krehan, I.: Unterschiedliche Steuerung der Imaginaldiapause bei den beiden Geschlechtern des LaufkäfersAgonum assimile Payk. Naturwissenschaften54, 122–123 (1967).Google Scholar
  8. ——: Experimentelle Untersuchungen zur Larvaldiapause des CarabidenPterostichus vulgaris. Ent. exp. and appl.12, 67–73 (1969).Google Scholar

Copyright information

© Springer-Verlag 1970

Authors and Affiliations

  • Wilfried Paarmann
    • 1
  1. 1.Institut für Angewandte Zoologie der Universität WürzburgWürzburgDeutschland

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