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The forest and scrub vegetation in limestone areas of Shikoku, Japan

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Summary

Eleven communities in limestone areas of Shikoku have been described and considered from a phytosociological point of view.

The evergreen broad-leaved forests and scrubs in the warm temperate region are included in the Camellietea japonicae, and deciduous broad-leaved and coniferous forests excluding the Carpinus turczaninovii-Zabelia integrifolia association in the cool temperate region belong to the Fagetea crenatae. These classes are inclusive of climatic climaxes, and the major dominants are not characteristic of limestone areas. A noticeable fact, however, is the occurrence of the Quercus glauca-Nandina domestica association in limestone areas, where the Castanopsis cuspidata forests prevailing in climatic climaxes are replaced by this association.

The limestone vegetation varies considerably with the situation and the physiographic factors, and the most characteristic community typified by the Carpinus turczaninovii-Zabelia integrifolia association appears in xeric sites. This association, having a wide altitudinal range, is not included in any known classes, and the floristic composition is particularized by a large number of endemic and disjunctive elements.

Zusammenfassung

Als Ergebnis der pflanzensoziologischen Studien über Wälder und Sträucher in Kalkgegenden von Shikoku, Süd-Japan, wurden elf Pflanzengesellschaften beschrieben.

Die immergrünen Wälder und Sträucher im warmtemperierten Gebiet werden in die Klasse Camellietea japonicae eingereiht, während im kalttemperierten Gebiet die Sommergrün- und Nadelwälder ausschließlich der Carpinus turczaninovii-Zabelia integrifolia-Assoziation, Klasse Fagetea crenatae, gehören.

Diese Klassen bestehen aus klimatischen Klimaxgesellschaften, und die wichtigeren Dominanten sind keine Kalk-Spezialisten. Es ist aber zu bemerken, daß sich die Quercus glauca-Nandina domestica-Assoziation in Kalkgegenden befindet, wo die in klimatischen Klimaxgesellschaften überwiegenden Castanopsis cuspidata-Wälder durch diese Assoziation ersetzt werden.

Mit dem Standort und den physiographischen Faktoren verändert sich die Kalkvegetation ansehnlich, und die durch die Carpinus turczaninovii-Zabelia integrifolia-Assoziation höchst charakteristisch dargestellte Pflanzengesellschaft erscheint in trockenen Standorten. Diese Assoziation wird in keine der bekannten Klassen eingeschlossen, die floristische Zusammensetzung kennzeichnet sich durch eine große Anzahl endemischer und disjunkter Elemente.

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Yamanaka, T. The forest and scrub vegetation in limestone areas of Shikoku, Japan. Vegetatio Acta Geobot 19, 286–307 (1969). https://doi.org/10.1007/BF00259016

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