The occurrence of sex-linked yellow is described and designated as tortoiseshell (symbol To). The phenotype produced by the heterozygous female (To+) is a mosaic of that of ToTo and ++. Genetic inactivation of one X chromosome per cell is thus indicated. The To gene interacts phenotypically with genes Ba, Ds and Wh.
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Robinson, R. Genetic studies of the Syrian hamster. VIII. The sex-linked gene Tortoiseshell. Genetica 43, 239–243 (1972). https://doi.org/10.1007/BF00123631
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DOI: https://doi.org/10.1007/BF00123631