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Cultivation of Gracilaria verrucosa (Gracilariales, Rhodophyta) Strain G-16 for agar


The agarophyte, Gracilaria verrucosa Strain G-16, has been grown in sustained cultivation over a period of five years. During this period, a number of experiments were conducted to examine the productivity, agar yield and agar gel strength of this strain. Productivities range from 3–31 g dry wt m−2 d−1 and are generally highest in the summer when annual water temperatures and daily irradiances are highest. In the summer months agar yield from Strain G-16 appears to be lowest whereas the gel strength of the agars was highest (> 750 g cm−2).

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  • Abbott, I. A., 1980. Some field and laboratory studies on colloid producing red algae in central California. Aquat. Bot. 8: 255–266.

    Google Scholar 

  • Asare, S. O., 1980. Seasonal changes in sulfate and 3,6 anhydrogalactose content of phycocolloids from two red algae. Bot. mar. 23: 595–598.

    Google Scholar 

  • Bird, K. T., 1987. Cost analyses of energy from marine biomass. In K. T. Bird & P. Benson (eds), Seaweed cultivation for renewable resources. Elsevier, Amsterdam: 327–350.

    Google Scholar 

  • Bird, K. T., 1988. Agar production and quality from Gracilaria sp. Strain G-16: effects of environmental factors. Bot. mar. 31: 33–39.

    Google Scholar 

  • Bird, K. T., M. D. Hanisak & J. Ryther, 1981. Chemical quality and production of agars extracted from Gracilaria tikvahiae grown in different nitrogen enrichment conditions. Bot. mar. 24: 441–444.

    Google Scholar 

  • Craigie, J. S., Z. C. Wen & J. P. van der Meer, 1984. Interspecific, intraspecific and nutritionally determined variations in the composition of agars from Gracilaria spp. Bot. mar. 27: 55–61.

    Google Scholar 

  • Daugherty, B. K. & K. T. Bird, 1988. Salinity and temperature effects on agar production from Gracilaria verrucosa Strain G-16. Aquaculture 75: 105–113.

    Google Scholar 

  • Dawes, C. J., 1987. The biology of commercially important tropical marine algae. In K. T. Bird & P. Benson (eds), Seaweed cultivation for renewable resources. Elsevier, Amsterdam: 155–190.

    Google Scholar 

  • Edelstein, T., C. J. Bird & J. McLachlan,1976. Studies on Gracilaria spp. Experiments on inocula incubated under greenhouse conditions. J. exp. mar. Biol. Ecol. 30: 249–259.

    Google Scholar 

  • Friedlander, M. & N. Zelikovitch, 1984. Growth rates, phycocolloid yield and quality of the red seaweeds, Gracilaria sp., Pterocladia capillacea, Hypnea musciformis, and Hypnea cornuta, in field studies in Israel. Aquaculture 40: 57–66.

    Google Scholar 

  • Guerin, J. M. & K. T. Bird, 1987. Effects of aeration period on the productivity and agar quality of Gracilaria tikvahiac. Aquaculture 64: 105–110.

    Google Scholar 

  • Hanisak, M. D., 1987. Cultivation of Gracilaria and other macroalgae in Florida for energy production. In K. T. Bird & P. Benson (eds), Seaweed cultivation for renewable resources. Elsevier, Amsterdam: 191–218.

    Google Scholar 

  • Hoyle, M. D., 1978. Agar studies in two Gracilaria species (G. bursa-pastoris (Gmelin) Silva and G. coronopifolia J.Ag.) from Hawaii. II. Seasonal aspects. Bot. mar. 21: 347–352.

    Google Scholar 

  • Ryther, J. H., N. Corwin, T. A. DeBusk & L. D. Williams, 1981. Nitrogen uptake and storage by the red alga, Gracilaria tikvahiae McLachlan. Aquaculture 26: 107–115.

    Google Scholar 

  • Strumski, M. A., K. T. Bird & J. H. Ryther, 1985. Agar production in Gracilaria tikvahiae (Rhodophyta). International Council for the Exploration of the Sea: CM 1985 (F:44 ) Mariculture Committee, 12 pp.

  • Wang, C.-Y. & S.-S. Yang, 1980. Seasonal variation of the quality of Gracilaria cultivation in Taiwan. Proc. natl. Sci. Council R.O.C. 4: 78–86.

    Google Scholar 

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Bird, K.T., Ryther, J.H. Cultivation of Gracilaria verrucosa (Gracilariales, Rhodophyta) Strain G-16 for agar. Hydrobiologia 204, 347–351 (1990).

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Key words

  • agar
  • algae
  • aquaculture
  • Gracilaria
  • seasonality