Swiss Journal of Palaeontology

, Volume 138, Issue 1, pp 87–97 | Cite as

Regurgitated ammonoid remains from the latest Devonian of Morocco

  • Christian KlugEmail author
  • Lothar H. Vallon
Regular Research Article


Accumulations of ammonoid shell fragments have been recovered from the Hangenberg Black Shale (latest Devonian) of the southern Maïder (eastern Anti-Atlas, Morocco). They are here interpreted as regurgitalites and ascribed tentatively to gnathostomes as possible tracemakers. The recognition of fossil regurgitations is reviewed and a checklist provided.


Ammonoidea Mass extinctions Hangenberg event Chondichthyes Food web Digestichnia 



We greatly appreciate the support by the Swiss National Science Foundation (Project Numbers 200020_132870, 200020_149120, 200021_156105). We thank the Ministère de l’Energie, des Mines, de l’Eau et de l’Environnement (Direction du Développement Minier, Division du Patrimoine, Rabat, Morocco) for working and sample export permits. We acknowledge the thorough reviews of Kenneth De Baets (Erlangen) and an anonymous reviewer that helped to improve the current paper.


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© Akademie der Naturwissenschaften Schweiz (SCNAT) 2018

Authors and Affiliations

  1. 1.Paläontologisches Institut und MuseumZurichSwitzerland
  2. 2.Geomuseum Faxe (Østsjællands Museum)FaxeDenmark

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