, Volume 91, Issue 3, pp 353–373 | Cite as

The oldest bryozoans of Baltoscandia from the lowermost Floian (Ordovician) of north-western Russia: two new rare, small and simple species of Revalotrypidae

  • Petr V. FedorovEmail author
  • Anna V. Koromyslova
  • Silviu O. Martha
Research Paper


We describe two new species of esthonioporate bryozoans, Revalotrypa inopinata sp. nov. and R. yugaensis sp. nov., from the Lower Ordovician, south-east of Lake Ladoga, north-western Russia. The colonies of the two species are very small and were extracted from limestone nodules found in the lower part of glauconitic sandstone in the Joa Member (lowermost Floian). Revalotrypa inopinata sp. nov. and R. yugaensis sp. nov. are the oldest known bryozoans from Baltoscandia. Study of the two species includes scanning electron microscopy (SEM) imaging and X-ray microtomography (micro-CT). The palaeogeographic distribution of the oldest known bryozoans from the Lower Ordovician and the advantages and disadvantages of micro-CT for the study of Palaeozoic bryozoans are discussed. The authorship of Revalotrypidae, which has been inconsistently allocated in the literature, is discussed.


Bryozoa Esthonioporata Taxonomy Ordovician Floian Micro-CT 


Wir beschreiben zwei neue Arten esthonioporater Bryozoen (Revalotrypa inopinata sp. nov. und R. yugaensis sp. nov.) aus dem Unter-Ordovizium südöstlich des Ladoga-Sees im Nordwesten Russlands. Die Kolonien der beiden Bryozoenarten sind sehr klein und wurden aus Kalksteinknollen extrahiert, die im unteren Bereich von Glaukonitsanden der Joa-Einheit (unterstes Floium) gefunden worden sind. R. inopinata sp. nov. und R. yugaensis sp. nov. sind die ältesten bekannten Bryozoen Balto-Skandinaviens. Die beiden Arten wurde mit dem Rasterelektronenmikroskop (REM) und der Röntgen-Mikro-Computertomographie (Micro-CT) untersucht. Die paläogeographische Verbreitung der ältesten bekannten Bryozoen aus dem Unter-Ordovizium sowie die Vor- und Nachteile von Micro-CT für die Untersuchung paläozoischer Bryozoen wird diskutiert. Die in der Literatur bisher uneinheitlich gehandhabte Autorenschaft der Revalotrypidae wird ebenso diskutiert.


Bryozoa Esthonioporata Taxonomie Ordovizium Floium Micro-CT 



Alexandr Alekseev (Lomonosov Moscow State University, Moscow), Andrej Ernst (Uni Hamburg), Dennis P. Gordon (National Institute of Water and Atmospheric Research, Wellington), Joachim Scholz (Senckenberg Forschungsinstitut und Naturmuseum, Frankfurt) and Paul D. Taylor (Natural History Museum, London) are thanked for advice regarding the authorship of Revalotrypidae. Andrej Ernst, Paul D. Taylor and the editor-in-chief Mike Reich (Bayerische Staatssammlung für Paläontologie und Geologie, München) are also thanked for helpful reviews and comments on the manuscript. The third author (S.O.M.) gratefully acknowledges financial support of the Deutsche Forschungsgemeinschaft (Project SCHO 581/12–1 “Enhancing documentation and digitalization of the Bryozoa collection donated by Professor Voigt (1905–2004) to the Senckenberg Research Institute, Frankfurt am Main, Germany”).


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Copyright information

© Paläontologische Gesellschaft 2017

Authors and Affiliations

  • Petr V. Fedorov
    • 1
    Email author
  • Anna V. Koromyslova
    • 2
  • Silviu O. Martha
    • 3
  1. 1.Department of Sedimentary Geology, Institute of Earth SciencesSt. Petersburg State UniversitySt. PetersburgRussia
  2. 2.Borissiak Paleontological Institute of the Russian Academy of SciencesMoscowRussia
  3. 3.Senckenberg Forschungsinstitute und Naturmuseen, Sektion Marine Evertebraten III (Bryozoologie)FrankfurtGermany

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