Landscape Ecology

, Volume 28, Issue 8, pp 1465–1478 | Cite as

Characterizing wild ass pathways using a non-invasive approach: applying least-cost path modelling to guide field surveys and a model selection analysis

Research article


Movement of animals is a key process affecting population dynamics. Information on factors that affect pathway use is essential for identifying and protecting pathways, and important for maintaining connectivity among populations. We present an innovative, non-invasive, approach for predicting pathways of reintroduced Asiatic wild ass (Equus hemionus) in Israel, which is based on understanding the effects of landscape factors on pathways use. The approach includes: Predicting pathways, by employing a least cost pathway (LCP) GIS models based on several landscape factors, so as to efficiently direct a field survey and explore the wild ass’s general preferences of pathway types; Collecting empirical data by surveying the dung density of wild ass along each of the predicted pathways and using the data as an index of pathway use; Evaluating the predicted pathways against the empirical data collected, to estimate the general pathway preferences of the wild ass; and Developing and evaluating alternative generalized linear models, according to a priori hypotheses based on empirical data so as to quantify the effect of different landscape factors on pathway use. The analyses were conducted for the entire landscape, and then for two distinct landscape types, open landscape and landscape-barriers (mountain ridges), as subsets of the entire landscape. There were clear differences in the mean number of faeces counts between the LCPs, indicating that the wild ass prefers certain pathway types as a function of landscape features. We further found that the factors affecting E. hemionus pathway usage—vegetation; slopes; canyons; and 4-wheel drive routes—varied largely between the two major landscape types studied, demonstrating the importance of studying space use patterns at different landscape terrains. This information can be applicable to landscape planning measures that aim to enhance protection of the species. This approach provides a framework for studying animal space-use patterns of a variety of species, including elusive species, in a heterogeneous landscape.


Akaike’s information criterion (AIC) Conservation Equus hemionus Faeces GIS Landscape barriers Landscape connectivity Least-cost models Reintroduction 

Supplementary material

10980_2013_9915_MOESM1_ESM.doc (886 kb)
Supplementary material 1 (DOC 885 kb)


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Copyright information

© Springer Science+Business Media Dordrecht 2013

Authors and Affiliations

  • Achiad Davidson
    • 1
  • Yohay Carmel
    • 2
  • Shirli Bar-David
    • 1
  1. 1.Mitrani Department of Desert EcologyJacob Blaustein Institutes for Desert Research, Ben-Gurion University of the NegevMidreshet Ben-GurionIsrael
  2. 2.Faculty of Civil and Environmental Engineering, TechnionIsrael Institute of TechnologyHaifaIsrael

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