Algae, calcitarchs and the Late Ordovician Baltic limestone facies of the Baltic Basin
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The Late Ordovician succession of the Baltic Basin contains a characteristic fine-grained limestone, which is rich in calcareous green algae. This limestone occurs in surface outcrops and drill-cores in an extensive belt reaching from Sweden across the Baltic Sea to the Baltic countries. This limestone, which is known in the literature under several different lithological names, is described and interpreted, and the term “Baltic limestone facies” is suggested. The microfacies, from selected outcrops from the Åland Islands, Finland and Estonia, consists of calcareous green algae as the main skeletal component in a bioclastic mudstone-packstone lithology with a pure micritic matrix. Three types of calcitarch, which range in diameter from c. 100–180 μm, are common. Basinward, the youngest sections of the facies belt contain coral-stromatoporoid patch reefs and Palaeoporella-algal mounds. The Baltic limestone facies can be interpreted as representing the shallow part of an open-marine low-latitude carbonate platform.
KeywordsLithographic limestone Shallow-water carbonates Dasycladacales Calcitarch Ordovician
The Ordovician succession of the Baltic region is characterized by a drastic change from siliciclastic sediments and cold-water carbonates in the Early Ordovician to warm-water carbonates, with reefs and mud mounds, with little siliciclastic input, in the Late Ordovician. A unique and enigmatic lithofacies within the Late Ordovician warm-water carbonates consists of a dense, hard, stylolitic, fine-grained limestone, with a specific conchoidal fracture and very even texture. Commonly the limestone has been referred to as “calcilutitic”, “aphanitic” or “lithographic” (e.g. Jaanusson and Martna 1948; Spjeldnæs and Nitecki 1994; Hints et al. 2005). It is best described as a fine-grained bioclastic carbonate mudstone—packstone. Its most conspicuous and commonly most abundant skeletal elements are fragments of benthic green algae (e.g. Wiman 1893; Stolley 1896, 1897; Martna 1955; Hucke and Voigt 1967; Kozlowski and Kazmierczak 1968; Vingisaar 1971; Põlma 1972, 1982; Kõrts et al. 1990; Spjeldnæs and Nitecki 1994; Hints et al. 2005).
Herein, the term “Baltic limestone facies” is used to cover all these names. Baltic limestone lithologies occur stratigraphically in an interval that ranges from the Keila Regional Stage (late Sandbian Stage) towards the higher Pirgu Stage (late Katian Stage or lowermost Hirnantian) (Fig. 2). They are stratigraphically most persistent in a wide belt, which reaches from Bothnian Bay and the Åland Islands in the west to northern Gotland in the south and the Saaremaa Island and central Estonia in the east (Fig. 1). The greatest extension of this facies towards the south was reached during the time of Nabala and Rakvere regional stages. During this time the Slandrom Formation extended as far west as to the Siljan area in central Sweden (e.g. Calner et al. 2010) and the Saunja Formation as far south as central Lithuania (Männil 1966).
One particular feature of the Baltic limestone facies is its proximity to mud mounds and reefs, which are especially widespread and thick during the Pirgu Stage (Kröger et al. 2016b) but are also reported from the Rakvere Stage, as in the Puhmu drill core from central Estonia (Kaljo et al. 2017). Notably, green algae such as Palaeoporella, Vermiporella and dasycladaceans are abundant in these structures and in some places they are the dominant skeletal element (Jux 1966; Kröger et al. 2016a, b).
Here, a microfacies analysis of limestone samples from selected localities from the Baltic limestone facies belt is presented. The aim of this study is to understand better the formation processes that led to the widespread deposition of pure limestones in the region. The potential role of algae in producing the Baltic limestone facies is investigated. Spherical calcareous microfossils, which have been mentioned but not previously figured (Spjeldnæs and Nitecki 1994: p. 270), are abundant in some samples. These fossils are determined as calcitarchs and interpreted as algal spores. Calcitarchs are spherical calcareous microfossils (< 550 μm) of unknown and varied biological affinities, previously often subsumed under the more diffuse and less well defined term “calcisphere” (Versteegh et al. 2009). They have been implicated in regional-scale micrite production in later Palaeozoic marine basins, and here we assess the possibility that they were important carbonate sources in the Ordovician Baltic Basin.
Material and sampling
All samples are from Late Ordovician limestones of the peri-cratonic Baltic Basin, of northeastern Europe. The basin is situated between the Scandinavian shield in the present north and the Sarmatian shield in the present south (Sliaupa et al. 2006) and records a thick early Palaeozoic succession. The Ordovician sediments of the basin have been spatially subdivided by faunal and lithological differences into a number of zones, termed “confacies belts” (Männil 1966; Jaanusson 1976). The confacies belts roughly correspond to the spatial lithofacies distribution along a proximal—distal gradient (e.g. Männil 1966; Jaanusson 1982; Raukas and Teedumäe 1997; Harris et al. 2004; Kröger et al. 2016b). Samples examined here are from drill cores, surface outcrops and erratic boulders of the Åland Islands, Finland and from western and southern Estonia (Fig. 1).
A number of samples were analysed as polished surfaces, in thin-sections under optical light, and under scanning electron microscopy (SEM). For SEM analysis the samples where polished and slightly etched following the methods described in Munnecke et al. (2000).
Tranvik-Lumparn borehole nr. 6, Åland Islands, Finland
The drilling of this core took place during a mainly off-shore drilling campaign of the Partek company (now Nordkalk Oy Ab) in 1964, which produced c. 25 drill-cores from the Lumparn bay area. Core no. 6 was drilled in northern Lumparn Bay, ~ 250 m south of the tip of Tranviknäs (60°10′20.24″N, 20°9′55.55″E) and has been previously mentioned or briefly described (Merrill 1980; Tynni 1982). The core covers a thickness of 34.5 m, but the stratigraphy of the section cannot be resolved in detail because of a lack data.
The topmost beds of borehole no. 6 are composed of a nodular, reddish fine-grained limestone with a Baltic limestone lithology, rich in Vermiporella with ~ 1 mm diameter branched thalli. The succession contains several strongly brecciated horizons, which are almost mylonitic with cm-sized angular intraclasts. The reddish nodular Baltic limestone lithology ranges down to the base of the preserved part of the core at 33 m, where it is rich in recrystallized microgastropods and Vermiporella sp. Generally, the section represents a metre-scale to tens of metre-scale alternation between reddish, argillaceous and lime-rich sections with calcareous algal, gastropod mudstone—wackestone lithologies. Merrill (1980), who described conodonts from older sediments in the Lumparn drill-cores, found none in the drill-core no. 6 samples. Tynni (1982) described the long ranging (upper Haljala Regional Stage, Sandbian–Juuru Regional Stage, Rhuddenian) Tasmanites aff. verrucosus from 21.2 m as the only determinable acritarch and Leiosphaeridia from the argillaceous interval 21.8–19.9 m. In Estonia Leiosphaeridia is not known from Ordovician beds younger than the Keila Regional Stage (uppermost Sandbian) (which is traditionally used as a stratigraphically significant pattern, e.g. Hints et al. 2016), which gives an upper Sandbian (Jõhvi-Keila Stage) age constraint for the 21.2–19.9 m interval. An upper age constraint for borehole no. 6 can be given from the nearby Tranvik-Lumparn borehole no. 7 and the Tranvik Udde outcrop, which produced Amorphognathus superbus zone, Rakvere-Nabala Regional Stage conodonts (Merrill 1980).
Erratic boulders from southwestern Finland
Hoitberg reef, Vormsi Island, Estonia
The Hoitberg reef is a natural surface outcrop in the centre of western Vormsi Island, Estonia, at 59°0′16″N, 23°10′58″E. The outcrop consists of an overgrown 5–10 m high elongate hill with a width of c. 30 m, length of c. 80–100 m, and a maximum expansion in NNW/SSE -direction. The reef is lithologically embedded within an algal-rich fine-grained limestone of the Moe Formation (Hints et al. 2005). Several large (metre-scale) blocks of the reef are exposed mostly on its northern part and provide insight into the composition of the reef core and flanks. The core consists of a coral-boundstone with a non-skeletal matrix content of 56–72%. The most abundant skeletal reef-builder is the heliotid tabulate Eocatenipora, which comprises 18–41% of the rock volume in the sampled areas. Stromatoporoids and bryozoans, mostly dendroid growth forms, were minor skeletal reef-builders with less than 5% of the rock volume. The matrix consists of mudstone—wackestone lithologies, in several places with a fenestral-microbial texture. The skeletal components of the matrix are Vermiporella algae and ostracods, calcitarchs, sponge spicules and bryozoan fragments are abundant.
Kaugatuma 509 borehole, Saarema Island, Estonia
The Kaugatuma 509 drill hole is situated close to the north-east coast of the Sõrve Peninsula at 58°7′25.572″N, 22°11′39.4794″E, on the southern Saaremaa Island, Estonia. The core is ~ 466 m deep and ranges from the Upper Silurian down to the Middle Ordovician. Parts of the Ordovician section have been previously described and figured (Põlma 1973; Oraspõld 1982; Oraspõld and Kala 1982; Nõlvak 1989; Kaljo 2001; Brenchley et al. 2003; Ainsaar et al. 2010). Herein, two samples are described: Kaug-2 from 411.38 m, Rägavere Formation, Rakvere Stage, and Kaug-3, 387.6 m from the upper part of the Saunja Formation, Nabala Stage. The top of the Keila Stage Kahula Formation at Kaugatuma 509 is a distinct hardground at 413.7 m, which marks an abrupt facies change from greenish-grey argillaceous wackestone toward a decimetre-scale intercalation of nodular fine-grained limestone and dark grey argillaceous interlayers of the Tõrremägi Member. The Tõrremägi Member grades toward the top into pure pale pinkish-grey nodular stylolitic, fine-grained limestone with a bioclastic mudstone-wackestone lithology. From 413.1 to 411.1 m a pure fine-grained limestone with bioclastic mudstone-wackestone lithology of the Piilse Member prevails. Its top, at 411.1 m, is a pyrite stained planar hardground with distinct Trypanites burrows. Sample Kaug-2 is from 0.18 m below this hardground. The overlying Paekna Formation of the Nabala Stage is again an alternation of decimetre-scale intercalation of nodular fine-grained limestone and dark green argillaceous interlayers. The top of the Paekna Formation is relatively abrupt, with a hardground forming the base of the Saunja Formation at 407.4 m. The Saunja Formation consists of a massive, pure, pale pinkish-grey bioturbated stylolitic, fine-grained limestone with bioclastic mudstone-wackestone lithology; its top is a karstic, heavily burrowed and eroded surface at 387.1 m. Sample Kaug-3 is from 0.5 m below the top Saunja hardground.
Niibi trench, western Estonia
The outcrop is a shallow trench on both sides of the unpaved road that leads from Niibi hamlet to road 16,122, c. 150–400 m west of Niibi, Lääne county, western Estonia at 59°2′29.60″N, 23°39′28.89″E. The trench exposes the top of a number of small patch reefs, each with a maximum diameter of less than 5 m. Similar reefs also crop out on a fallow land ~ 50 m south of the road and have been described from a now nearly overgrown quarry ~ 180 m to the west of the road. Fossils collected from these and probably from other shallow reef outcrops in the direct vicinity have been described (Kaljo 1961; Nestor 1964; Klaamann 1966; Preobrazhensky and Klaamann 1975).
The reefs occur within the Moe Formation, Pirgu Stage, late Katian. Some of these patch reefs are covered and surrounded by a coarse, partly argillaceous greenish echinoderm packstone—echinoderm grainstone. The reefs themselves are composed of a matrix-rich coral-boundstone. The matrix has a skeletal mudstone—wackestone lithology, in several places with a fenestral-microbial texture. The dominant skeletal element of the boundstone is the heliotid coral Eocatenipora (see Klaamann 1966) and locally large colonies of Sarcinula. Stromatoporoid colonies are common (see Nestor 1964). The most abundant skeletal elements in the matrix are dasycladal green algae, echinoderm ossicles and fragments of bryozoan skeletons.
Valga 10 borehole, southern Estonia
The Valga 10 drill-hole is located in southern Estonia, near the Latvian border at 57°48′14.4″N, 26°04′39″E on the NE outskirts of the town of Valga. The section was described in detail in a monographic treatment (Põldvere 2001); it is 424.4 m deep and penetrates Quaternary, Devonian, Silurian and Ordovician sediments. Since 2001 individual sections of the core have been described and discussed in numerous publications from which a few are relevant herein because they give additional age constraints and lithological information (Ainsaar and Meidla 2001; Nõlvak and Bauert 2006; Ainsaar et al. 2010). Sample Vg-1 is from 379.65 m, from 0.25 above the base of the Rägavere Formation, Rakvere Stage. The Rägavere Formation, ~ 6.7 m thick at Valga 10, is unconformity bounded and consists of greenish to brownish, wavy thin-bedded fine-grained limestone with a skeletal mudstone lithology, and with finely dispersed pyrite and dolomite mainly in its upper parts. The limestone is cracked and contains predominantly sub-centimetre to centimetre-thick sparite-filled veins which contain geopetal argillaceous crystal dolomitic silt (see also Ainsaar and Meidla 2001) which we interpret as palaeokarst following Calner et al. (2010).
Võhma borehole, central Estonia
The Võhma drill-hole is located south of the parish Võhma, Viljandi county, central Estonia, at 58°36′44.136″N, 25°33′34.056″E. The drill-core section is 438 m deep with a stratigraphic range from the Proterozoic crystalline basement up to the lower Silurian Adavere Stage. Parts of the section have been described in publications; most relevant are Oraspõld (1982) and Kröger et al. (2016a, b), which cover the parts of the Pirgu Stage. The Pirgu Stage is exceptional at the Võhma drill-core, because it exposes ~ 52 m of a Palaeoporella mud mound.
Here, seven samples, Vm01–Vm07, are analysed ranging stratigraphically from the Rakvere Stage to the upper part of the mound. The samples Vm02 and Vm06 are from 220.55 m and 218.78 m, Rakvere Stage, which ranges in the Võhma drill-core from 221.75 to 214.50 m (Fig. 3b, c). The position of the lower boundary of the Rakvere Stage is not fully resolved because the lithology grades from a predominantly argillaceous decimetre-scale intercalation of greenish-grey nodular fine-grained limestone and dark-grey argillaceous interlayers toward massive pinkish fine-grained limestone in a ~ 0.5 m interval. Here, the notebook no. 6 of Põlma (http://geokogud.info/doi/10.15152/GEO.126) is followed where the boundary is positioned at the base of the lowermost pinkish nodular fine-grained limestone bed. The upper boundary of the Rägavere Formation is a distinct, pyrite-stained hardground, which marks a sharp lithological boundary towards the more argillaceous greenish limestone of the Paekna Formation, Nabala Stage. The limestone of the Rägavere Formation is cracked at some levels and contains predominantly sub-centimetre to centimetre-thick sparite-filled veins containing partly gravitationally filled argillaceous crystal dolomitic silt which is interpreted as a palaeokarst following Calner et al. (2010). Samples from the algal-mud mound, Pirgu Stage, are from depths 144.20 m (Vm04), 155.37 m (Vm03), 160.15 (Vm07); 187.29 m (Vm01) and comprise the two key lithologies of the mound. Samples Vm01 and Vm03 are from fenestral, stromatactis-rich bioclastic wackestone—packstone lithologies and samples Vm04 and Vm07 represent originally algal-rich parts of the mound.
Description of the calcitarchs
Type-II calcitarchs. Small spheres with thin, simple wall. Diameter ~ 130 μm (median, 131 μm; 1. quantile, 103 μm; 3. quantile, 174 μm) (Fig. 4ad, ae, ba). Wall thickness ~ 0.05–0.15 of sphere diameter. Occur in most samples of the Baltic limestone facies and in the mounds and reefs.
Description of the microfacies
The only non-matrix component in the mudstone lithology consists of round microspar dots (diameter < 50 μm) with diffuse margins and type-I calcitarchs (Figs. 3a, 5e). The skeletal wacke-packstone lithology contains abundant ostracod shell hush and fragments of Moniliporella sp. thalli (Fig. 5d). The thalli are partly strongly disintegrated and marginally micritized. Shell fragments of brachiopods, trilobites and echinoderms occur. The packstone lithology contains predominantly skeletal fragments of Moniliporella sp. and strongly disintegrated and marginally micritized fragments of Vermiporella sp. Few type-I and type-II calcitarchs occur.
Rägavere Formation, Rakvere Regional Stage, early Katian Stage. The samples from the Rägavere Formation range in lithology from mudstone to skeletal packstone. The matrix is similar to the Baltic limestone samples. The skeletal components of the mudstone (sample Vg-1, Fig. 5f) are rare type-I calcitarchs, disintegrated shell fragments of ostracods and monaxial and triaxial sponge spicules. In the wacke-packstone (sample Vm02, Figs. 3b, 5b), dasyclad algae (Moniliporella sp.) are predominant; disintegrated ostracod shell fragments and microgastropods are abundant and a few echinoderm ossicles occur. Type-I and type-III calcitarchs are rare. The sample Vm06 (Figs. 3c, 5a) has a skeletal packstone lithology with c. 50% fragments of green algal skeletons. Fragments of thalli of Vermiporella sp. are predominant, Moniliporella sp. are second abundant. The partly strongly disintegrated fragments are marginally micritized to different degrees. Ostracod shell hash, microgastropods and echinoderm ossicles occur. Type-I and type-II calcitarchs, and brachiopod shell fragments are rare.
Saunja Formation, Nabala Regional Stage, early Katian Stage. Two samples from the Saunja Formation were analysed (samples Kaug-2, Kaug-3) with mudstone–packstone lithologies. The matrix is similar to the Rägavere Formation samples. The mudstone contains only a few echinoderm ossicles and type-II calcitarchs. The skeletal components of the wacke–packstone lithologies contain predominantly fragments of Moniliporella sp. and strongly disintegrated Vermiporella sp. Shell fragments of brachiopods and bryozoans occur; microgastropods and type-II calcitarchs are rare.
Coral-stromatoporoid reefs, Pirgu regional stage, late Katian. The samples from the Hoitberg reef are boundstone to packstone lithologies with the tabulate Eocatenipora vormsiense Klaamann, as the main skeletal component and abundant fragments of bryozoans and ostracod shells as secondary skeletal components (Fig. 8e–f). The matrix of the boundstone consists of a mudstone-wackestone lithology with a clotted fabric and partly with irregular fenestrae that are filled with sparite and a gravitational micritic infill. Type-II and type-III calcitarchs occur. Notably patches and individual thalli of questionable Apidium sp. occur in this matrix. The intraclastic, fenestral packstone lithology is rich in fragments of dasycladaceans, predominantly Vermiporella sp. Type-II and type-III calcitarchs occur. Similar skeletal components and calcitarchs occur also in the matrix of the boundstone and packstone lithologies of the Niibi reefs.
The micritic matrix of the Baltic limestone facies
One of the distinctive features of the Baltic limestone facies is its fine-grained, homogenous micritic matrix. In macroscopic view, patches commonly occur with different colours between pink and yellow, probably tracing bioturbation patterns. These patches are caused by differences in organic matter and skeletal content, or the distribution of finely dispersed pyrite and locally by the occurrence of dolomite, all probably related to minor differences in original porosity. In thin-section, centimetre-sized burrows are commonly preserved with perfect circular cross-sections in a horizontal or subhorizontal direction, which indicate little or no diagenetic compaction of the limestone and hence early lithification. Early lithification is also indicated by swiss-cheese karst features crossing down from the top Rägavere and top Saunja formation surfaces, respectively, and from the uncompressed fossils themselves. Under high magnification the micritic matrix commonly shows flake-like, angular patches or lumps with sizes of less than 50 μm consisting of μm-sized clusters of micrite crystals.
The relatively high abundance of non-disintegrated, marginally micritized dasycladacean thalli among the skeletal fragments in the limestones is evidence for a shallow-water environment with a depositional depth of less than 25 m within the shallow euphotic zone (Granier 2012). This is an environment in which other calcareous green algae could have flourished with less rigid skeletons, such as Acetabularia. Recent calcareous green algae are known to produce 750 g CaCO3/m2 (Acetabularia, Marszalek 1975) to up to 2400 g CaCO3/m2 (Freile et al. 1995). Under SEM several micritic samples exhibit a microporous texture, which is very similar to the one described in Lasemi and Sandberg (1984) and Munnecke et al. (1997), probably representing relics of aragonite needles within a μm-range, and which can be interpreted as representing a texture of an original aragonite-dominated lime-mud (Fig. 6f).
It is, therefore, coherent to assume a green algal, aragonitic origin and rapid recrystallisation of the micritic matrix of the Baltic limestone facies analogous to the diagenetic process known from recent aragonite-dominated tropical platform sediments (Munnecke et al. 1997). Our interpretation is also in accordance with the long-standing opinion that the appearance of a widespread micrite-rich limestone facies is related to the palaeogeographical drift of the Baltica continent toward lower latitudes and warmer climatic conditions (Dronov and Rozhnov 2007; see also Nestor and Einasto 1997).
Comparison and interpretation of Late Ordovician calcitarchs
Few descriptions of Palaeozoic calcitarch occurrences exist. The specimens described and figured herein are unique and differ from previously described forms in terms of size and partly in wall structure. The spores found in situ in the thalli of Late Ordovician Vermiporella sp. are smaller (~ 35 μm, Kozlowski and Kazmierzak 1968). The spherical microfossils from the Silurian Visby and Högklint formations are also generally smaller (< 100 μm, Munnecke and Servais 2008; Servais et al. 2009). The simple-walled spherical microfossils described from the Devonian Choteč and Třebotov formations are similar in general appearance and structure to our type-II calcitarchs but are also smaller (80–120 μm, Berkyova and Munnecke 2010). Late Palaeozoic radiospheres (Kaźmierczak and Kremer 2005) differ in having external spines.
The calcitarchs described herein, show no signs of organic walls in thin-section or under SEM observation and, therefore, cannot be interpreted as calcified acritarchs. However, all three calcitarch groups distinguished herein are close to the size range known from recent calcified spherical non-motile asexual spores (aplanospores) of chlorophyceans, such as Acetabularia (140–185 μm, Marszalek 1975). Notably, all these Ordovician calcitarchs co-occur with green algae and have their mass occurrences, where green algae are abundant. This co-occurrence is not a taphonomic effect of preferred preservation within specifically fine-grained micritic matrix, because calcitarchs occur also in peloidal and intraclastic packstones of the algal-rich reef facies (see above).
The calcitarchs, described herein, occur together with calcified green algae as the most conspicuous skeletal component of the Baltic limestone facies and its associated reefs. It appears likely that the calcitarchs described herein are aplanospores of various green algae, probably of taxa that are not preserved as articulated fossils, like the fast disintegrating Acetabularia. This interpretation is in accordance with the idea that the characteristic fine-grained and homogenous micritic matrix of the Baltic limestone facies originated from aragonitic algal mud.
Comparison of the Baltic limestone facies with similar Palaeozoic carbonates elsewhere
Probably the most similar facies to the Baltic limestone facies can be found in the algal-rich platform carbonates of the Ordos-Basin (northwest China) within the early Katian Beiguoshan Formation (Zheng et al. 2018). Zheng et al. (2018) described a situation with coral-stromatoporoid patch reefs, a platform margin-reef belt and micritic carbonates in a back-reef shallow-water environment, which is rich in dasycladaceans, other green algae, gastropods and crinoids.
Algal-rich massive limestone deposits, algal mounds and reefs are also described from the Katian Lianglitage Formation of the western Tarim Basin (northwest China) and are interpreted as tropical shallow-water carbonates (Zhang et al. 2014; Shen and Neuweiler 2016). Notably the Ordos Basin and the Tarim Basin carbonates contain Vermiporella sp. as the dominant algal constituent, similar to the Baltic limestone facies. Calcitarchs are not described from these northwest China carbonate platforms.
They are described from the micritic limestone of the marl-limestone alternations of the Upper Visby and Högklint formations (latest Llandovery to early Wenlock, Munnecke et al. 1999; Servais et al. 2009). These occurrences are also from a shallow tropical carbonate environment with small patch reefs but differ in lacking the abundant green algal component and in being generally more argillaceous than the massive and pure Baltic limestone facies. The calcitarch bearing Devonian Choteč and Třebotov formations represent different depositional environments ranging from relatively deep to extremely shallow, and do not contain reefs, mounds or green algae.
In conclusion, the Baltic limestone facies appears to be related to time-equivalent tropical shallow platform carbonates, probably reflecting primarily similar climatic conditions such as high temperatures and extremely low terrigenous sediment input.
Along the eastern and northeastern margin of the Baltic Basin a wide facies-belt developed during the Late Ordovician, characterised by an extremely pure limestone with a fine homogenous micritic matrix. The limestone is rich in dasycladacean algae and contains abundant calcitarchs. This algal-rich micritic limestone which comprises several regional chronostratigraphic zones and lithostratigraphic units is combined under the new term “Baltic limestone facies”. Three types of calcitarchs and a number of genera of green algae (Apidium sp., Dasyporella sp., Mastipora sp., Moniliporella, Vermiporella sp.) can be distinguished within the samples analysed in our study. The dasycladaceans indicate an original deposition of the limestone within the shallow euphotic and euryhaline zone under low-latitude climatic conditions. Basinward the limestone contains coral-stromatoporoid patch reefs and Palaeoporella-algal mounds. The Baltic limestone facies can be interpreted as representing the shallower part of an open-marine carbonate platform.
Open access funding provided by University of Helsinki including Helsinki University Central Hospital. BK is grateful to Susan Scholze and Duncan Matthews (Helsinki) for support with data compilation and preparation of samples. We are grateful to Kenneth Fjäder and Gerhard Hakkarainen (Nordkalk Oy, Parainen) for access to Åland drill cores. Ursula Toom (Tallinn), Mare Isakar (Tartu) and Gennady Baranov (Tallinn) where wonderful field work companions in Estonia, and Dina Schultze and Radoslaw Michallik (Helsinki) provided technical assistance and useful discussions. Many thanks also to Olle Hints (Tallinn) for general support during field work in Estonia. BK and AP were funded by the Academy of Finland. This is a contribution to the IGCP Project 653 ‘The Onset of the Great Ordovician Biodiversification Event’.
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