Oecologia

, Volume 143, Issue 1, pp 1–10 | Cite as

Effects of fire on properties of forest soils: a review

Concepts, Reviews, and Syntheses

Abstract

Many physical, chemical, mineralogical, and biological soil properties can be affected by forest fires. The effects are chiefly a result of burn severity, which consists of peak temperatures and duration of the fire. Climate, vegetation, and topography of the burnt area control the resilience of the soil system; some fire-induced changes can even be permanent. Low to moderate severity fires, such as most of those prescribed in forest management, promote renovation of the dominant vegetation through elimination of undesired species and transient increase of pH and available nutrients. No irreversible ecosystem change occurs, but the enhancement of hydrophobicity can render the soil less able to soak up water and more prone to erosion. Severe fires, such as wildfires, generally have several negative effects on soil. They cause significant removal of organic matter, deterioration of both structure and porosity, considerable loss of nutrients through volatilisation, ash entrapment in smoke columns, leaching and erosion, and marked alteration of both quantity and specific composition of microbial and soil-dwelling invertebrate communities. However, despite common perceptions, if plants succeed in promptly recolonising the burnt area, the pre-fire level of most properties can be recovered and even enhanced. This work is a review of the up-to-date literature dealing with changes imposed by fires on properties of forest soils. Ecological implications of these changes are described.

Keywords

Fire Forest ecosystems Forest soils Soil ecology Soil properties 

References

  1. Adams MA, Attiwill PM (1984) Role of Acacia spp. in nutrient balance and cycling in regenerating Eucalyptus regnans F. Muell. Forests. I. Temporal changes in biomass and nutrient content. Aust J Bot 32:205–215Google Scholar
  2. Adams PW, Boyle JR (1980) Effects of fire on soil nutrients in clearcut and whole-tree harvest sites in Central Michigan. Soil Sci Soc Am J 44:847–850Google Scholar
  3. Alcaniz JM, Granada E, Comellas L (1994) Simulating the effects of burning on soil organic matter in a forest soil studied by pyrolysis gas chromatography. In: Senesi N, Miano TM (eds) Humic substances in the global environment and implications on human health. Elsevier, Amsterdam, pp 205–212Google Scholar
  4. Almendros G., Martín F, González-Vila FJ (1988) Effects of fire on humic and lipid fractions in a Dystric Xerochrept in Spain. Geoderma 42:115–127CrossRefGoogle Scholar
  5. Almendros G, Gonzalez-Vila FJ, Martin F (1990) Fire-induced transformation of soil organic matter from an oak forest: an experimental approach to the effects of fire on humic substances. Soil Sci 149:158–168Google Scholar
  6. Almendros G, Gonzalez-Vila FJ, Martin F, Frund R, Ludemann HD (1992) Solid state NMR studies of fire-induced changes in the structure of humic substances. Sci Total Environ 117–118:63–74CrossRefGoogle Scholar
  7. Arocena JM, Opio C (2003) Prescribed fire-induced changes in properties of sub-boreal forest soils. Geoderma 113:1–16CrossRefGoogle Scholar
  8. Bååth E, Frostegård A, Pennanen T, Fritze H (1995) Microbial community structure and pH response in relation to soil organic matter quality in wood-ash fertilized, clear-cut or burned coniferous forest soils. Soil Biol Biochem 27:229–240CrossRefGoogle Scholar
  9. Baar J, Horton TR, Kretzer AM, Bruns TD (1999) Mycorrhizal colonization of Pinus muricata from resistant propagules after a stand-replacing wildfire. New Phytol 143:409–418CrossRefGoogle Scholar
  10. Badìa D, Martí C (2003) Plant ash and heat intensity effects on chemical and physical properties of two contrasting soils. Arid Land Res Manage 17:23–41CrossRefGoogle Scholar
  11. Baldock JA, Smernik RJ (2002) Chemical composition and bioavailability of thermally altered Pinus resinosa (Red pine) wood. Org Geochem 33:1093–1109CrossRefGoogle Scholar
  12. Bhadauria T, Ramakrishnan PS, Srivastava KN (2000) Diversity and distribution of endemic and exotic earthworms in natural and regenerating ecosystems in the central Himalayas, India. Soil Biol Biochem 32:2045–2054CrossRefGoogle Scholar
  13. Bissett J, Parkinson D (1980) Long-term effects of fire on the composition and activity of the soil microflora of a subalpine, coniferous forest. Can J Bot 58:1704–1721Google Scholar
  14. Boerner REJ, Brinkman JA (2003) Fire frequency and soil enzyme activity in southern Ohio oak–hickory forests. Appl Soil Ecol 23:137–146CrossRefGoogle Scholar
  15. Boix Fayos C (1997) The roles of texture and structure in the water retention capacity of burnt Mediterranean soils with varying rainfall. Catena 31:219–236CrossRefGoogle Scholar
  16. Boyer WD, Miller JH (1994) Effect of burning and brush treatments on nutrient and soil physical properties in young longleaf pine stands. For Ecol Manage 70:311–318Google Scholar
  17. Cade-Menun BJ, Berch SM, Preston CM, Lavkulich LM (2000) Phosphorus forms and related soil chemistry of Podzolic soils on northern Vancouver Island. II. The effects of clear-cutting and burning. Can J For Res 30:1726–1741CrossRefGoogle Scholar
  18. Caldararo N (2002) Human ecological intervention and the role of forest fires in human ecology. Sci Total Environ 292:141–165CrossRefGoogle Scholar
  19. Cammeraat LH, Imeson AC (1999) The evolution and significance of soil-vegetation patterns following land abandonment and fire in Spain. The significance of soil, water and landscape processes in banded vegetation patterning. Catena 37:107–127CrossRefGoogle Scholar
  20. Campbell GS, Jungbauer JD Jr, Bidlake WR, Hungerford RD (1994) Predicting the effect of temperature on soil thermal conductivity. Soil Sci 158:307–313Google Scholar
  21. Campbell GS, Jungbauer JD Jr, Bristow KL, Hungerford RD (1995) Soil temperature and water content beneath a surface fire. Soil Sci 159:363–374Google Scholar
  22. Choromanska U, DeLuca TH (2001) Prescribed fire alters the impact of wildfire on soil biochemical properties in a ponderosa pine forest. Soil Sci Soc Am J 65:232–238Google Scholar
  23. Choromanska U, DeLuca TH (2002) Microbial activity and nitrogen mineralization in forest mineral soils following heating: evaluation of post-fire effects. Soil Biol Biochem 34:263–271CrossRefGoogle Scholar
  24. Collett NG, Neumann FG, Tolhurst KG (1993) Effects of two short rotation prescribed fires in spring on surface-active arthropods and earthworms in dry sclerophyll eucalypt forest of west-central Victoria. Aust For 56:49–60Google Scholar
  25. Covington WW, Sackett SS (1992) Soil mineral nitrogen changes following prescribed burning in ponderosa pine. For Ecol Manage 54:175–191Google Scholar
  26. Covington WW, DeBano LF, Huntsberger TG (1991) Soil nitrogen changes associated with slash pile burning in pinyon-juniper woodlands. For Sci 37:347–355Google Scholar
  27. Crockford RH, Willett IR (2001) Application of mineral magnetism to describe profile development of toposequences of a sedimentary soil in south-eastern Australia. Aust J Soil Res 39:927–949CrossRefGoogle Scholar
  28. Czimczik CI, Preston CM, Schmidt MWI, Werner RA, Schulze E-D (2002) Effects of charring on mass, organic carbon, and stable carbon isotope composition of wood. Org Geochem 33:1207–1223CrossRefGoogle Scholar
  29. DeBano LF (2000) The role of fire and soil heating on water repellence in wildland environments: a review. J Hydrol 231:195–206CrossRefGoogle Scholar
  30. DeBano LF, Neary DG, Ffolliott PF (1998) Fire effects on ecosystems. Wiley, New YorkGoogle Scholar
  31. Doerr SH, Shakesby RA, Walsh RPD (1998) Spatial variability of soil hydrophobicity in fire-prone eucalyptus and pine forests, Portugal. Soil Sci 163:313–324CrossRefGoogle Scholar
  32. Doerr SH, Shakesby RA, Walsh RPD (2000) Soil water repellence: its causes, characteristics and hydro-geomorphological significance. Earth-Sci Rev 51:33–65CrossRefGoogle Scholar
  33. Durgin PB, Vogelsang PJ (1984) Dispersion of kaolinite by water extracts of Douglas-fir ash. Can J Soil Sci 64:439–443Google Scholar
  34. Everett RL, Java-Sharpe BJ, Scherer GR, Wilt FM, Ottmar RD (1995) Co-occurrence of hydrophobicity and allelopathy in sand pits under burned slash. Soil Sci Soc Am J 59:1176–1183Google Scholar
  35. Fernández I, Cabaneiro A, Carballas T (1997) Organic matter changes immediately after a wildfire in an Atlantic forest soil and comparison with laboratory soil heating. Soil Biol Biochem 29:1–11CrossRefGoogle Scholar
  36. Fernández I, Cabaneiro A, Carballas T (1999) Carbon mineralization dynamics in soils after wildfires in two Galician forests. Soil Biol Biochem 31:1853–1865CrossRefGoogle Scholar
  37. Fisher RF, Binkley D (2000) Ecology and management of forest soils, 3rd edn. Wiley, New YorkGoogle Scholar
  38. Fonturbel MT, Vega JA, Bara S, Bernardez I (1995) Influence of prescribed burning of pine stands in NW Spain on soil microorganisms. Eur J Soil Biol 31:13–20Google Scholar
  39. Franklin SB, Robertson PA, Fralish JS (1997) Small-scale fire temperature patterns in upland Quercus communities. J Appl Ecol 34:613–630Google Scholar
  40. Fritze H, Pennanen T, Pietikainen J (1993) Recovery of soil microbial biomass and activity from prescribed burning. Can J For Res 23:1286–1290Google Scholar
  41. Gillon D, Gomendy V, Houssard C, Marechal J, Valette JC (1995) Combustion and nutrient losses during laboratory burns. Int J Wildland Fire 5:1–12Google Scholar
  42. Giovannini G, Lucchesi S (1997) Modifications induced in soil physico-chemical parameters by experimental fires at different intensities. Soil Sci 162:479–486CrossRefGoogle Scholar
  43. Giovannini G, Lucchesi S, Giachetti M (1988) Effects of heating on some physical and chemical parameters related to soil aggregation and erodibility. Soil Sci 146:255–261Google Scholar
  44. Goh K, Phillips MJ (1991) Effects of clearfell logging and clearfell logging and burning of a Nothofagus forest on soil nutrient dynamics in South Island, New Zealand—changes in forest floor organic matter and nutrient status. N Z J Bot 29:367–384Google Scholar
  45. Gonzalez Parra J, Cala Rivero V, Iglesias Lopez T (1996) Forms of Mn in soils affected by a forest fire. Sci Total Environ 181:231–236CrossRefGoogle Scholar
  46. González-Pérez JA, González-Vila FJ, Almendros G, Knicker H (2004) The effect of fire on soil organic matter—a review. Environ Intl 30:855–870CrossRefGoogle Scholar
  47. Grogan P, Bruns TD, Chapin FS III (2000) Fire effects on ecosystem nitrogen cycling in a Californian bishop pine forest. Oecologia 122:537–544CrossRefGoogle Scholar
  48. Guinto DF, Saffigna PG, Xu ZH, House APN, Perera MCS (1999) Soil nitrogen mineralisation and organic matter composition revealed by 13C NMR spectroscopy under repeated prescribed burning in eucalypt forests of south-east Queensland. Aust J Soil Res 37:123–135Google Scholar
  49. Hartford RA, Frandsen WH (1992) When it’s hot, it’s hot... or maybe it’s not! (Surface flaming may not portend extensive soil heating). Int J Wildland Fire 2:139–144Google Scholar
  50. Haumaier L, Zech W (1995) Black carbon—possible source of highly aromatic components of soil humic acids. Org Geochem 23:191–196CrossRefGoogle Scholar
  51. Henderson GS, Golding DL (1983) The effect of slash burning on the water repellence of forest soils at Vancouver, British Columbia. Can J For Res 13:353–355Google Scholar
  52. Hernandez T, Garcia C, Reinhardt I (1997) Short-term effect of wildfire on the chemical, biochemical and microbiological properties of Mediterranean pine forest soils. Biol Fertil Soils 25:109–116CrossRefGoogle Scholar
  53. Horne DJ, McIntosh JC (2000) Hydrophobic compounds in sands in New Zealand-extraction, characterisation and proposed mechanisms for repellence expression. J Hydrol 231:35–46CrossRefGoogle Scholar
  54. Huffman EL, MacDonald LH, Stednick JD (2001) Strength and persistence of fire-induced soil hydrophobicity under ponderosa and lodgepole pine, Colorado Front Range. Hydrol Process 15:2877–2892CrossRefGoogle Scholar
  55. Imeson AC, Verstraten JM, van Mulligen EJ, Sevink J (1992) The effects of fire and water repellence on infiltration and runoff under Mediterranean type forest. Catena 19:345–361CrossRefGoogle Scholar
  56. Johnson DW, Curtis PS (2001) Effects of forest management on soil C and N storage: meta analysis. For Ecol Manage 140:227–238CrossRefGoogle Scholar
  57. Ketterings QM, Bigham JM (2000) Soil color as an indicator of slash-and-burn fire severity and soil fertility in Sumatra, Indonesia. Soil Sci Soc Am J 64:1826–1833Google Scholar
  58. Ketterings QM, Bigham JM, Laperche V (2000) Changes in soil mineralogy and texture caused by slash-and-burn fires in Sumatra, Indonesia. Soil Sci Soc Am J 64:1108–1117Google Scholar
  59. Khanna PK, Raison RJ (1986) Effect of fire intensity on solution chemistry of surface soil under a Eucalyptus pauciflora forest. Aust J Soil Res 24:423–434Google Scholar
  60. Khanna PK, Raison RJ, Falkiner RA (1994) Chemical properties of ash derived from Eucalyptus litter and its effects on forest soils. For Ecol Manage 66:107–125Google Scholar
  61. Kim EJ, Oh JE, Chang YS (2003) Effects of forest fire on the level and distribution of PCDD/Fs and PAHs in soil. Sci Total Environ 311:177–189CrossRefGoogle Scholar
  62. Klopatek CC, DeBano LF, Klopatek JM (1988) Effects of simulated fire on vesicular–arbuscular mycorrhizae in pinyon–juniper woodland soil. Plant Soil 109:245–249Google Scholar
  63. Knicker H, Almendros G, González-Vila FJ, Martin F, Lüdemann H-D (1996) 13 C- and 15 N-NMR spectroscopic examination of the transformation of organic nitrogen in plant biomass during thermal treatment. Soil Biol Biochem 28:1053–1060CrossRefGoogle Scholar
  64. Kutiel P, Shaviv A (1992) Effects of soil type, plant composition and leaching on soil nutrients following a simulated forest fire. For Ecol Manage 53:329–343Google Scholar
  65. Letey J (2001) Causes and consequences of fire-induced soil water repellence. Hydrol Process 15:2867–2875CrossRefGoogle Scholar
  66. Ludwig B, Khanna PK, Raison RJ, Jacobsen KL (1998) Modelling cation composition of soil extracts under ashbeds following an intense slashfire in a eucalypt forest. For Ecol Manage 103:9–20Google Scholar
  67. Macadam AM (1987) Effects of broadcast slash burning on fuels and soil chemical properties in the sub-boreal spruce zone of central British Columbia. Can J For Res 17:1577–1584Google Scholar
  68. Marcos E, Tarrega R, Luis-Calabuig E (2000) Comparative analysis of runoff and sediment yield with a rainfall simulator after experimental fire. Arid Soil Res Rehab 14:293–307CrossRefGoogle Scholar
  69. Martin DA, Moody JA (2001) Comparison of soil infiltration rates in burned and unburned mountainous watersheds. Hydrol Process 15:2893–2903CrossRefGoogle Scholar
  70. Martínez-Sánchez JJ, Ferrandis P, de las Heras J, Herranz JM (1999) Effect of burnt wood removal on the natural regeneration of Pinus halepensis after fire in a pine forest in Tus valley (SE Spain). For Ecol Manage 123:1–10Google Scholar
  71. Mataix-Solera J, Doerr SH (2004) Hydrophobicity and aggregate stability in calcareous topsoils from fire-affected pine forests in southeastern Spain. Geoderma 118:77–88CrossRefGoogle Scholar
  72. Matlack GR (2001) Factors determining the distribution of soil nematodes in a commercial forest landscape. For Ecol Manage 146:129–143Google Scholar
  73. McSorley R (1993) Short-term effects of fire on the nematode community in a pine forest. Pedobiologia 37:39–48Google Scholar
  74. Mermut AR, Luk SH, Romkens MJM, Poesen JWA (1997) Soil loss by splash and wash during rainfall from two loess soils. Geoderma 75:203–214CrossRefGoogle Scholar
  75. Miltner A, Zech W (1997) Effects of minerals on the transformation of organic matter during simulated fire-induced pyrolysis. Org Geochem 26:175–182CrossRefGoogle Scholar
  76. Monleon VJ, Cromack K Jr (1996) Long-term effects of prescribed underburning on litter decomposition and nutrient release in ponderosa pine stands in central Oregon. For Ecol Manage 81:143–152Google Scholar
  77. Mroz GD, Jurgensen MF, Harvey AE, Larsen MJ (1980) Effects of fire on nitrogen in forest floor horizons. Soil Sci Soc Am J 44:395–400Google Scholar
  78. Naidu CV, Srivasuki KP (1994) Effect of forest fire on soil characteristics in different areas of Seshachalam hills. Ann For 2:166–173Google Scholar
  79. Neumann FG, Tolhurst K (1991) Effects of fuel reduction burning on epigeal arthropods and earthworms in dry sclerophyll eucalypt forest of west-central Victoria. Aust J Ecol 16:315–330Google Scholar
  80. Oswald BP, Davenport D, Neuenschwander LF (1999) Effects of slash pile burning on the physical and chemical soil properties of Vassar soils. J Sustainable For 8:75–86Google Scholar
  81. Perry DA, Rose S, Pilz D, Schoenberger MM (1984) Reduction of natural ferric iron chelators in disturbed forest soils. Soil Sci Soc Am J 48:379–382Google Scholar
  82. Pietikainen J, Fritze H (1995) Clear-cutting and prescribed burning in coniferous forest: comparison of effects on soil fungal and total microbial biomass, respiration activity and nitrification. Soil Biol Biochem 27:101–109CrossRefGoogle Scholar
  83. Ponomarenko EV, Anderson DW (2001) Importance of charred organic matter in black Chernozem soils of Saskatchewan. Can J Soil Sci 81:285–297Google Scholar
  84. Prieto-Fernandez A, Villar MC, Carballas M, Carballas T (1993) Short-term effects of a wildfire on the nitrogen status and its mineralization kinetics in an Atlantic forest soil. Soil Biol Biochem 25:1657–1664CrossRefGoogle Scholar
  85. Prieto-Fernández A, Acea MJ, Carballas T (1998) Soil microbial and extractable C and N after wildfire. Biol Fertil Soils 27:132–142CrossRefGoogle Scholar
  86. Rab MA (1996) Soil physical and hydrological properties following logging and slash burning in the Eucalyptus regnans forest of southeastern Australia. For Ecol Manage 84:159–175Google Scholar
  87. Rabenhorst MC (1988) Determination of organic and carbonate carbon in calcareous soils using dry combustion. Soil Sci Soc Am J 52:965–969Google Scholar
  88. Raison RJ, Khanna PK, Woods PV (1985) Mechanisms of element transfer to the atmosphere during vegetation fires. Can J For Res 15:132–140Google Scholar
  89. Robichaud PR (2000) Fire effects on infiltration rates after prescribed fire in Northern Rocky Mountain forests, USA. J Hydrol 231:220–229CrossRefGoogle Scholar
  90. Romanya J, Khanna PK, Raison RJ (1994) Effects of slash burning on soil phosphorus fractions and sorption and desorption of phosphorus. For Ecol Manage 65:89–103Google Scholar
  91. Schmidt MWI, Noack AG (2000) Black carbon in soils and sediments: analysis, distribution, implications, and current challenges. Global Biogeochem Cycles 14:777–793CrossRefGoogle Scholar
  92. Schmidt MWI, Skjemstad JO, Gehrt E, Kögel-Knabner I (1999) Charred organic carbon in German chernozemic soils. Eur J Soil Sci 50:351–365CrossRefGoogle Scholar
  93. Schwertmann U, Taylor RM (1989) Iron oxides. In: Dixon JB, Weed SB (eds) Minerals in soil environments, 2nd edn. Soil Science Society of America, Madison, Wis., pp 379–438Google Scholar
  94. Scott DF (2000) Soil wettability in forested catchments in South Africa; as measured by different methods and as affected by vegetation cover and soil characteristics. J Hydrol 231–232:87–104CrossRefGoogle Scholar
  95. Scott DF, van Wyk DB (1990) The effects of wildfire on soil wettability and hydrological behaviour of an afforested catchment. J Hydrol 121:239–256CrossRefGoogle Scholar
  96. Serrasolsas I, Khanna PK (1995) Changes in heated and autoclaved forest soils of S.E. Australia. II. Phosphorus and phosphatase activity. Biogeochemistry 29:25–41Google Scholar
  97. Sevink J, Imeson AC, Verstraten JM (1989) Humus form development and hillslope runoff, and the effects of fire and management, under Mediterranean forest in NE-Spain. Catena 16:461–475CrossRefGoogle Scholar
  98. Shakesby RC, Coelho C, Ferreira AD, Terry JP, Walsh RPD (1993) Wildlife impacts on soil erosion and hydrology in wet Mediterranean forest, Portugal. Int J Wildland Fire 3:95–110Google Scholar
  99. Sharpley A (2000) Phosphorous availability. In: Sumner ME (ed) Handbook of soil science. CRC, Boca Raton, Fla., pp D18–D38Google Scholar
  100. Simard DG, Fyles JW, Paré D, Nguyen T (2001) Impacts of clearcut harvesting and wildfire on soil nutrient status in the Quebec boreal forest. Can J Soil Sci 81:229–237Google Scholar
  101. Soto B, Diaz-Fierros F (1993) Interactions between plant ash leachates and soil. Int J Wildland Fire 3:207–216Google Scholar
  102. Tan KH, Hajek BF, Barshad I (1986) Thermal analysis techniques. In: Klute A (ed) Methods of soil analysis. 1. Physical and mineralogical methods. American Society of Agronomy and Soil Science Society of America, Madison, WI, pp 151–183Google Scholar
  103. Thomas AD, Walsh RPD, Shakesby RA (1999) Nutrient losses in eroded sediment after fire in eucalyptus and pine forests in the wet Mediterranean environment of northern Portugal. Catena 36:283–302CrossRefGoogle Scholar
  104. Tomkins IB, Kellas JD, Tolhurst KG, Oswin DA (1991) Effects of fire intensity on soil chemistry in a eucalypt [Eucalyptus sp.] forest. Aust J Soil Res 29:25–47Google Scholar
  105. Torres P, Honrubia M (1997) Changes and effects of a natural fire on ectomycorrhizal inoculum potential of soil in a Pinus halepensis forest. For Ecol Manage 96:189–196Google Scholar
  106. Ulery AL, Graham RC (1993) Forest fire effects on soil color and texture. Soil Sci Soc Am J 57:135–140Google Scholar
  107. Ulery AL, Graham RC, Amrhein C (1993) Wood-ash composition and soil pH following intense burning. Soil Sci 156:358–364Google Scholar
  108. Ulery AL, Graham RC, Bowen LH (1996) Forest fire effects on soil phyllosilicates in California. Soil Sci Soc Am 60:309–315Google Scholar
  109. Vihnanek RE, Ballard TM (1988) Slashburning effects on stocking, growth, and nutrition of young Douglas-fir plantations in salal-dominated ecosystems of eastern Vancouver Island. Can J For Res 18:718–722Google Scholar
  110. Vilariño A, Arines J (1991) Numbers and viability of vesicular-arbuscular fungal propagules in field soil samples after wildfire. Soil Biol Biochem 23:1083–1087CrossRefGoogle Scholar
  111. Walstad JD, Radosevich SR, Sandberg DV (eds) (1990) Natural and prescribed fire in the pacific Northwest forest. Oregon State University Press, CorvallisGoogle Scholar
  112. Wanner M, Xylander WER (2003) Transient fires useful for habitat-management do not affect soil microfauna (testate amoebae)—a study on an active military training area in eastern Germany. Ecol Eng 20:113–119CrossRefGoogle Scholar
  113. Wardle DA, Zackrisson O, Hornberg G, Gallet C (1997) Influence of island area on ecosystem properties. Science 277:1296–1299CrossRefGoogle Scholar
  114. Wardle DA, Zackrisson O, Nilsson MC (1998) The charcoal effect in Boreal forests: mechanisms and ecological consequences. Oecologia 115:419–426CrossRefGoogle Scholar
  115. Weston CJ, Attiwill PM (1996) Clearfelling and burning effects on nitrogen mineralization and leaching in soils of old-age Eucalyptus regnans forests. For Ecol Manage 89:13–24Google Scholar
  116. Weston CJ, Attiwill PM (1990) Effects of fire and harvesting on nitrogen transformations and ionic mobility in soils of Eucalyptus regnans forests of south-eastern Australia. Oecologia 83:20–26CrossRefGoogle Scholar
  117. Zackrisson O, Nilsson MC, Wardle DA (1996) Key ecological function of charcoal from wildfire in the Boreal forest. Oikos 77:10–19Google Scholar

Copyright information

© Springer-Verlag 2005

Authors and Affiliations

  1. 1.Dipartimento di Scienza del Suolo e Nutrizione della PiantaUniversità degli Studi di FirenzeFlorenceItaly

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