A new delthyridoid spiriferid genus from the Dra Valley, Morocco, and its phylogenetic affinities (Brachiopoda, Lower Devonian)
- 52 Downloads
- 1 Citations
Abstract
Multiplicatispirifer, a new delthyridoid brachiopod genus with a gently divergent capillate micro-ornamentation and bifurcating costae on flanks, sulcus and fold, has been identified in the Mdâouer-el-Kbîr Formation of the Dra Valley (southern Anti-Atlas, Morocco). Its type speciesMultiplicatispirifer foumzguidensis n. gen., n. sp., at present the only species of this genus, is described. This form has hitherto been determined asFimbrispirifer orStruveina by various authors. In this work, the new genus is compared with the multiplicate and capillate spiriferid genera with bifurcating costae which also have costae in the sulcus and on the fold:Costispirifer andPerryspirifer from North America,Elymospirifer from South China,Borealispirifer from Asia andMultispirifer from Central Europe. The relation ofMultiplicatispirifer to the fimbriateFimbrispirifer, Struveina andVandercammenina, from which it is distinguished by its capillate micro-ornamentation, is discussed. The new genus is hitherto only known from the Moroccan southern Anti-Atlas. Its colsest relatives, taxa ofCostispirifer, occur in the Eastern American Realm (Gaspé, Canada and the eastern United States) and the Nevada Province (western United States).Multiplicatispirifer occurrences are restricted to the Lower/Upper Emsian boundary interval (upper Lower Devonian). The reasons for a possible migration corridor between North America and North Africa before Emsian time are discussed.
Keywords
Spiriferida Brachiopoda Phylogeny Paläobiogeography Morocco Lower DevonianKurzfassung
In der Mdâouer-el-Kbîr-Formation in der Dra-Ebene (südlicher Anti-Atlas, Marokko) wurde eine neue delthyridoide Brachiopoden-Gattung identifiziert.Multiplicatispirifer ist durch eine leicht divergierende kapillate Mikrostruktur und bifurkierende Rippen auf den Flanken, im Sinus und auf dem Sattel charakterisiert. Die TypusartMultiplicatispirifer foumzguidensis n. gen., n. sp., zur Zeit die einzige bekannte Art dieser Gattung, wird beschrieben. Bisher wurde diese Form von unterschiedlichen Bearbeitern zuFimbrispirifer oderStruveina gestellt. In der vorliegenden Arbeit wird die neue Gattung mit multiplikaten und kapillaten Spiriferen verglichen, die bifurkierende Rippen wie auch einen berippten Sinus und Sattel aufweisen:Costispirifer undPerryspirifer aus Nord-Amerika,Elymospirifer aus Süd-China,Borealispirifer aus Asien undMultispirifer aus Mittel-Europa. Die Verwandtschaft vonMultiplicatispirifer zu den fimbriaten GattungenFimbrispirifer, Struveina undVandercammenina, von denen sie sich durch ihre kapillate Mikrostruktur unterscheidet, wird diskutiert. Die neue Gattung ist bisher nur aus dem marokkanischen Anti-Atlas bekannt. Ihre nächsten Verwandten, Taxa vonCostispirifer, treten im ostamerikanischen Faunenreich (Gaspé, Kanada, und Ostküste der Vereinigten Staaten) sowie in der Faunenprovinz Nevada (westliche USA) auf. Das Vorkommen vonMultiplicatispirifer ist auf den Grenzbereich Unter-/Ober-Emsium beschränkt (oberes Unter-Devon). Die Gründe für einen möglichen Migrations-Korridor zwischen Nord-Amerika und Nord-Afrika zur Zeit des Emsiums werden diskutiert.
Schlüsselwörter
Spiriferida Brachiopoda Phylogenie Paläobiogeographie Marokko Unter-DevonPreview
Unable to display preview. Download preview PDF.
References
- Allan, R.S. 1947. A revision of the Brachiopoda of the Lower Devonian of Reefton, New Zealand. — Journal of Paleontology21 (5): 436–452.Google Scholar
- Becker, R.T.;Jansen, U.;Plodowski, G.;Schindler, E.;Aboussalam Z.S. &Weddige, K. 2004. Devonian litho- and biostratigraphy of the Dra Valley area: An overview. — In:El Hassani, A., ed., Devonian neritic-pelagic correlation and events in the Dra Valley (Western Anti-Atlas, Morocco). International Meeting on Stratigraphy, Rabat, March 1–10, 2004. — Documents de l’Institut Scientifique19: 3–18.Google Scholar
- Billings, E. 1874. Paleozoic fossils, Vol. 2, Part 1. — 144 p., Montreal (Geological Survey of Canada).Google Scholar
- Boucot, A.J. 1959. Brachiopods of the Lower Devonian rocks at High-land Mills, New York. — Journal of Paleontology33 (5): 727–769.Google Scholar
- Boucot, A.J. 1960a. A new Lower Devonian stropheodontid brachiopod. — Journal of Paleontology34 (3): 483–485.Google Scholar
- Boucot, A.J. 1960b. Implications of Rhenish Lower Devonian brachiopods from Nova Scotia. — International Geological Congress XXI session, Part XII: 129–137.Google Scholar
- Boucot, A.J. 1975. Evolution and extinction rate controls. — 426 p., Amsterdam, Oxford, New York (Elsevier).Google Scholar
- Boucot, A.J.;MacDonald, G.J.F.;Milton, C. &Thompson, J.B. Jr. 1958. Metamorphosed middle Paleozoic fossils from Central Massachusetts, Eastern Vermont, and Western New Hampshire. — Bulletin f the Geological Society of America60: 855–870.CrossRefGoogle Scholar
- Bouyx, E.;Blaise, J.;Brice, D.;Gourvennec, R.;Lardeux, H. &Le Menn, J. 1992. Implications paléogéographiques des affinités nord-gondwanniennes et rhénanes des faunes dévoniennes de la zone de Meguma (Appalaches septentrionales). — Comptes Rendus de l’Academie des Sciences serie 2, Mechanique, Physique, Chimie, Sciences de l’Univers, Sciences de la Terre315 (3): 337–343.Google Scholar
- Carls, P. 1987. Ein Vorschlag zur biostratigraphischen Redefinition der Grenze Gedinnium/Siegenium und benachbarter Unter-Stufen. 1. Teil: Stratigraphische Argumente und Korrelation. — Courier Forschungsinstitut Senckenberg92: 77–121.Google Scholar
- Carls, P.;Meyn, H. &Vespermann, J. 1993. Lebensraum, Entstehung und Nachfahren vonHowellella (Iberhowellella)hollmanni n. sg., n. sp. (Spiriferacea; Lochkovium, Unter-Devon). — Senckenbergiana lethaea73 (2): 227–267.Google Scholar
- Carter, J.L.;Johnson, J.G.;Gourvennec, R. &Hou Hongfei 1994. A revised classification of the spiriferid brachiopods. — Annals of the Carnegie Museum63 (4): 327–374.Google Scholar
- Chlupáč, I. &Kukal, Z. 1988. Possible global events and the stratigraphy of the Barrandian Palaeozoic (Cambrian and Devonian). — Zprávy o geologických výzkumech43: 83–146.Google Scholar
- Clarke, J.M. 1907. Some New Devonian Fossils. — Geological Papers, New York State Museum Bulletin107: 153–291.Google Scholar
- Conrad, T.A. 1839. Descriptions of new species of organic remains. — 3rd Annual Report of the New York State Geological Survey: 57–66.Google Scholar
- Cooper, G.A. 1942. New genera of North American brachiopods. — Journal of Washington Academy of Sciences32 (8): 228–235.Google Scholar
- Drot, J. 1964. Rhynchonelloidea et Spiriferoidea siluro-dévoniens du Maroc pré-saharien. — Notes et Mémoires du Service Géologique du Maroc178: 1–288.Google Scholar
- Gentil, L. 1929. L’Anti-Atlas et le djebel Bani (Exploration de 1923). — Revue Géographique Physique & Géologie Dynamique2 (1): 5–66 + 68–69.Google Scholar
- Gourvennec, R. 1989. Brachiopodes Spiriferida du Dévonien inférieur du Massif Armoricain. Systématique, paléobiologie, évolution, biostratigraphie. — Biostratigraphie du Paléozoïque9: 1–281.Google Scholar
- Hall, J. 1857. Description of Palaeozoic Fossils. — 10th Annual Report of the Regents of the University of the State New York on Conditions of the State Cabinet of Natural History, Part C, Appendix: 41–186.Google Scholar
- Helmbrecht, W. &Wedekind, R. 1923. Versuch einer biostratigraphischen Gliederung der Siegener Schichten auf Grund von Rensselaerien und Spiriferen. — Glückauf41: 949–953.Google Scholar
- Hollard, H. 1960. La découverte de goniatites du genreSellanarcestes dans le Dévonien du Draa (Maroc présaharien) et ses conséquences stratigraphiques. — Notes et Mémoires du Service Géologique du Maroc19: 55–60.Google Scholar
- Hollard, H. 1963. Un tableau stratigraphique du Dévonien du Sud de l’Anti-Atlas. — Notes du Service Géologique du Maroc23: 105–109.Google Scholar
- Hollard, H. 1967. Le Dévonien du Maroc et du Sahara nord-occidental. — In:Oswald, D.H., ed., International Symposium on the Devonian System, Calgary1: 203–244.Google Scholar
- Hollard, H. 1981. Principaux charactères des formations dévoniennes de l’Anti-Atlas. — Notes du Service Géologique du Maroc42: 15–22.Google Scholar
- Hollard, H. &Lafuste, J. 1961.Trachypora limbata (Eaton) 1832 (Coelenterata, Tabulata) dans le Dévonien du Sud Marocain. — Notes et Mémoires du Service Géologique du Maroc20: 71–79.Google Scholar
- Ivanova, E.A. 1972. Osnovnyye zakonomernosti evolyutsee spiriferid (Brachiopoda). — Paleontologicheskii Zhurnal1972 (3): 28–42 [in Russian].Google Scholar
- Jansen, U. 2000. The Early Devonian genusAcrospirifer. A taxonomic odyssey. — The Millenium Brachiopod Congress, Abstracts; London.Google Scholar
- Jansen, U. 2001a. Morphologie, Taxonomie und Phylogenie unter-devonischer Brachiopoden aus der Dra-Ebene (Marokko) und dem Rheinischen Schiefergebirge (Deutschland). — Abhandlungen der senckenbergischen naturforschenden Gesellschaft554: 1–389.Google Scholar
- Jansen, U. 2001b. On the genusAcrospiriferHelmbrecht etWedekind, 1923 (Brachiopoda, Lower Devonian). Contributions to Lower Devonian brachiopods from the Rheinisches Schiefergebirge and adjacent areas, 1. — In:Fryda, J.;Blodgett, R.B. &Mergl, M., eds., Havlíček Volume. — Journal of the Czech Geological Society46 (3–4): 131–144.Google Scholar
- Jansen, U.;Becker, G.;Plodowski, G.;Schindler, E.;Vogel, O. &Weddige, K. 2004. The Emsian to Eifelian near Foum Zguid (NE Dra Valley, Morocco). — In:El Hassani, A., ed., Devonian neritic-pelagic correlation and events in the Dra Valley (Western Anti-Atlas, Morocco). International Meeting on Stratigraphy, Rabat, March 1–10, 2004. — Documents de l’Institut Scientifique19: 19–28.Google Scholar
- Jansen, U.;Lazreq, N.;Plodowski, G.;Schemm-Gregory, M.;Schindler, E. &Weddige, K. 2007. Neritic-pelagic correlation in the Lower and basal Middle Devonian of the Dra Valley (S Anti-Atlas, Moroccan Pre-Sahara). — Geological Society, Special Publications278: 9–37.CrossRefGoogle Scholar
- Jansen, U.;Schemm-Gregory, M. &Chen Xiuqin 2008. Comparison of some spiriferid brachiopods from the Lower Devonian of South China and Europe. — Fossils and Strata54: 99–104.Google Scholar
- Johnson, J.G. 1970. Great Basin Lower Devonian Brachiopoda. — Memoirs of the Geological Society of America121: 1–421.Google Scholar
- Johnson, J.G. 1979. Devonian brachiopod biostratigraphy. — Special Papers in Palaeontology23: 291–306.Google Scholar
- Johnson, J.G. &Hou Hongfei 2006. Delthyridoidea. — In:Kaesler, R.L., ed., Treatise on Invertebrate Paleontology, Part H, Brachiopoda5 (revised): 1825–1847, Lawrence, Kansas (Geological Society of America and University of Kansas).Google Scholar
- Jones, B. &Boucot, A.J. 1983. Spiriferida from the Lower Devonian strata of Southwest Ellesmere Island, Arctic Canada. — Journal of Paleontology57 (2): 327–346.Google Scholar
- Kaplun, L.I. 1961. Brakhiopody niznego devona severnogo pribalkhashya. — Materialy po geologii i poleznym iskopaemym Kazakhstana. Stratigrafiya i paleontologiya1 (26): 64–114 [in Russian].Google Scholar
- Kozłowski, R. 1946.Howellella, a new name forCrispellaKozłowski, 1929. — Journal of Paleontology20 (3): 295.Google Scholar
- Krantz, A. 1857. Ueber ein neues bei Muenzenberg aufgeschlossenes Petrefakten-Lager in den devonischen Schichten. — Verhandlungen des Naturhistorischen Vereins der Preußischen Rheinlande und Westfalens14: 143–165.Google Scholar
- Nalivkin, D.V. 1924. — In:Frederiks, G. Paleontologitcheskie ztoudy, 2: O verkhne kamennougolnykh spiriferidakh Ourala. — Izvestiya Geologicheskogo Komiteta38 (3): 295–324 [in Russian].Google Scholar
- Phillips, J. 1841. Figures and description of the Palaeozoic fossils of Cornwall, Devon and West Somerset. — xii+231 p., London (Longman & Co.).Google Scholar
- Pitrat, C.W. 1965. Spiriferidina. — In:Moore, R.C., ed., Treatise on Invertebrate Paleontology, Part H, Brachiopoda2: H667-H728, Lawrence, Kansas (Geological Society of America and University of Kansas).Google Scholar
- Schemm-Gregory, M. 2007. The Lower Devonian brachiopod genusAcrospirifer Helmbrecht and Wedekind, 1923 — globally distributed or highly endemic? — In: Subcommission on Devonian Stratigraphy and IGCP 499 Devonian Land Sea Interaction: Devonian Global Change: compelling changes in the Devonian world, highlighting new findings in the terrestrial and marine biomes: fish, invertebrates, plants, terrestrial vertebrates, global warming, mass extinction, bolide strikes, and global correlation. Program and Abstracts; Eureka, Nevada, 9–17 September 2007: 78–79.Google Scholar
- Schemm-Gregory, M. 2008a. A new species ofFilispirifer (Brachiopoda: Delthyridoidea) from the Dra Valley, Morocco (Lower Devonian) — Zootaxa1739: 53–68.Google Scholar
- Schemm-Gregory, M. 2008b. A new terebratulid brachiopod species from the Siegenian (middle Lower Devonian) of the Dra Valley, Morocco. — Palaeontology.Google Scholar
- Schemm-Gregory, M. in press.Frequentispirifer, a new spiriferid genus and its phylogenetic position within the Delthyridoidea (Brachiopoda, Lower Devonian). — Neues Jahrbuch für Geologie und Paläontologie.Google Scholar
- Steininger, J. 1853. Geognostische Beschreibung der Eifel. — 144 p., Trier (Lintz’sche Buchhandlung).Google Scholar
- Struve, W. 1995. Die Riesen-Phacopiden aus dem Maïder, SE-marokkanische Prä-Sahara. — Senckenbergiana lethaea75 (2): 77–129.Google Scholar
- Su Yangzhen &Hou Hongfei 1993. The Early Devonian Spiriferid BrachiopodBorealispirifer n. gen. from the Balkash-Mongolia-Okhotsk Region. — In:Yang Zunyi, ed., Stratigraphy and Palaeontology of China2: 139–147, Beijing (Geological Publishing House).Google Scholar
- Termier, H. &Termier, G. 1949. Essai sur l’évolution des Spiriféridés: Maroc. Service Géologique. Division de Mines et Géologique. — Notes et Mémoires74: 85–1123.Google Scholar
- Verneuil, E. de 1850. Tableu des fossiles to terrain dévonien du départment de la Sarthe. — Bulletin de la Société Géologique de France (2)7: 778–787.Google Scholar
- Verneuil, E.de 1869. Appendice à la Faune dévonienne du Bosphore. — Extrait de l’Asie Mineure, de P.Tchihatcheff (Paléontologie: 425–495): 1–64.Google Scholar
- Waagen, W.H. 1883. Salt Range fossils, Part 4 (2) Brachiopoda. — Palaeontologica Indica, Memoires13: 329–390.Google Scholar
- Walliser, O.H. 1996. Global Events in the Devonian and Carboniferous. — In:Walliser, O.H., ed., Global Events and Event Stratigraphy in the Phanerozoic: 225–250, Berlin, Heidelberg, New York (Springer).Google Scholar
- Wang Yü;Yu Changmin &Wu Qi 1974. [Advances in the Devonian biostratigraphy of South China.] — Memoires of Nanking Institute of Geology and Palaeontology6: 1–71 [in Chinese].Google Scholar
- Wedekind, R. 1926. Die devonische Formation. — In:Salomon, W., ed., Grundzüge der Geologie. Band2 (Erdgeschichte): 194–226, Stuttgart (Enke).Google Scholar