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Die Mitschattierung

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Contrast and concealment colour patterns take the form of spotting in visually articulated surroundings. In visually more uniform surroundings, the same effects are achieved by means of shading. The present paper brings some new examples of ‘countershading’, and explains the phenomenon of ‘shadow accent’ (the reverse of countershading: emphasis of shadow areas by pigmentation, increasing the contrast between the animal and its environment).

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Albrecht, H. Die Mitschattierung. Experientia 18, 284–286 (1962). https://doi.org/10.1007/BF02148239

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