1983, pp 265-278

Insect Species Diversity as a Function of Environmental Variability and Disturbance in Stream Systems

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Thienemann (1954) observed that the number of species inhabiting a given locality is highest in areas which have persisted largely unchanged for a long time and are characterized by diverse and predictable (i.e., not deviating substantially from the long-term norm) environmental conditions. The rhithron (i.e., the habitat of cold, rubble-bottom streams often found in the middle and upper reaches of river systems) epitomizes the ideal running water environment for co-existence of many species (Illies, 1969), most of which are usually insects (Hynes, 1970).