Abstract
The first bystrowianids recorded in the Gamskian Horizon of the East European Platform, Vyushkoviana operta gen. et sp. nov. and Dromotectum abditum sp. nov. (Amphibia, Chroniosuchia), are described based on specimens from the Malaya Northern Dvina and Don river basins. The patterns of the dermal scute evolution observed in bystrowianids warrant the subdivision of the group into the subfamilies Bystrowianinae Vjuschkov 1957, Dromotectinae subfam. nov. and Axitectinae subfam. nov. The second and third show progressive reduction of the external (paraxial) zone of the interosteoderm articulation. New data supporting close proximity or phyletic succession of the chroniosuchid and bystrowianid types of the osteoderm design are revealed. The concept of the Chroniosuchidae as a paraphyletic group is rejected. Evolutionary significance of the key features distinguishing the bystrowianid subfamilies is analyzed.
This is a preview of subscription content, log in via an institution to check access.
Similar content being viewed by others
References
Buchwitz, M., Foth, C., Kogan, I., and Voigt, S., On the use of osteoderm features in a phylogenetic approach on the internal relationships of the Chroniosuchia (Tetrapoda: Reptiliomorpha), Palaeontology, 2012, vol. 55, part 3, pp. 1–18.
Bulanov, V.V. and Golubev, V.K., Aquatic block of Sundyr community of Late Permian vertebrates of Eastern Europe, in Permskaya sistema: stratigrafiya, paleontologiya, paleogeografiya. Sbornik materialov Mezhdunarodnoi nauchnoi konferentsii k 170-letiyu so dnya otkrytiya permskoi sistemy (Permian System: Stratigraphy, Paleontology, and Paleogeography: Collected Works of International Scientific Conference Devoted to the 170th Anniversary of the Discovery of the Permian System), Perm: Perm. Gos. Nats. Issled. Univ., 2011, pp. 47–55.
Golubev, V.K., Narrow-armor chroniosuchians (Amphibia, Anthracosauromorpha) of the Late Permian of Eastern Europe, Paleontol. Zh., 1998, no. 3, pp. 64–73.
Golubev, V.K., Permian and Triassic chroniosuchians and biostratigraphy of the Upper Tatarian deposits of Eastern Europe based on tetrapods, Tr. Paleontol. Inst. Ross. Akad. Nauk, 2000, vol. 276, pp. 1–174.
Ivakhnenko, M.F. and Tverdokhlebova, G.I., Sistematika, morfologiya i stratigraficheskoe znachenie verkhnepermskikh khroniozukhov vostoka Evropeiskoi chasti SSSR (Systematics, Morphology, and Stratigraphic Significance of Upper Permian chroniosuchians of the eastern European part of the USSR), Saratov: Saratov. Gos. Univ., 1980.
Li, J., The most primitive lower tetrapod fauna in China, Sci. China, Ser. D, 2001, vol. 44, no. 1, pp. 47–51.
Li, J. and Cheng, Z., New anthracosaur and temnospondyl amphibians from Gansu, China, Vertebr. PalAsiat., 1999, vol. 37, no. 7, pp. 234–247.
Novikov, I.V. and Shishkin, M.A., Paleozoic relict in Triassic tetrapod communities: The last anthracosaurian amphibians, Sixth Symposium on Mesozoic Terrestrial Ecosystems and Biota: Short Papers, Beijing: China Ocean Press, 1995, pp. 29–32.
Novikov, I.V. and Shishkin, M.A., Triassic Chroniosuchia (Amphibia, Anthracosauromorpha) and evolution of dermal thoracic scutes in the Bystrowianidae, Paleontol. J., 2000, vol. 34, suppl. no. 2, pp. 165–178.
Novikov, I.V., Shishkin, M.A., and Golubev, V.K., Permian and Triassic anthracosaurs from Eastern Europe, in The Age of Dinosaurs in Russia and Mongolia, Benton, M.J., Shishkin, M.A., Unwin, D.M., and Kurochkin, E.N., Eds., Cambridge: Cambridge Univ. Press, 2000, pp. 60–70.
Schoch, R., Voigt, S., and Buchwitz, M., A chroniosuchid from the Triassic of Kyrgyzstan and analysis of chroniosuchian relationships, Zool. J. Linn. Soc., 2010, vol. 160, pp. 515–530.
Shishkin, M.A. and Novikov, I.V., Relict anthracosaurs in the Early Mesozoic of Eastern Europe, Dokl. Akad. Nauk, 1992, vol. 325, no. 4, pp. 829–832.
Shishkin, M.A. and Ochev, V.G., Fauna of terrestrial vertebrates as a basis of stratigraphy of continental Triassic deposits of the USSR, in Stratigrafiya i paleontologiya mezozoiskikh i paleogen-neogenovykh kontinental’nykh otlozhenii Aziatskoi chasti SSSR (Stratigraphy and Paleontology of Mesozoic and Paleogene-Neogene Continental Deposits of the Asian Part of the USSR), Leningrad: Nauka, 1967, pp. 74–82.
Tatarinov, L.P., Seymouriamorphen aus der Fauna der USSR, in Handbuch der Palaeoherpetologie, vol. 5 Batrachosauria (Anthracosauria), part B Gephyrostegida-Chroniosuchida, Stuttgart: G. Fischer Verlag, 1972, pp. 70–80.
Vjuschkov, B.P., New unusual animals from the Tatarian Stage of the European part of the USSR, Dokl. Akad. Nauk SSSR, 1957a, vol. 113, no. 1, pp. 183–186.
Vjuschkov, B.P., New kotlassiomorphs from the Tatarian of the European part of the USSR, Tr. Paleontol. Inst. Akad. Nauk SSSR, 1957b, vol. 68 (Cotylosaurs and Batrachosaurs from the Upper Permian of the Soviet Union), pp. 89–107.
Witzmann, F., Schoch, R.R., and Maisch, M.W., A relic basal tetrapod from the Middle Triassic of Germany, Naturwissenschaften, 2007, no. 95, pp. 67–72.
Young, C.C., A new Late Permian fauna from Jiyuan, Henan, Vertebr. PalAsiat., 1979, vol. 17, no. 2, pp. 99–113.
Author information
Authors and Affiliations
Corresponding author
Additional information
Original Russian Text © M.A. Shishkin, I.V. Novikov, J. Fortuny, 2014, published in Paleontologicheskii Zhurnal, 2014, No. 5, pp. 60–71.
Rights and permissions
About this article
Cite this article
Shishkin, M.A., Novikov, I.V. & Fortuny, J. New bystrowianid chroniosuchians (Amphibia, Anthracosauromorpha) from the Triassic of Russia and diversification of Bystrowianidae. Paleontol. J. 48, 512–522 (2014). https://doi.org/10.1134/S0031030114050098
Received:
Published:
Issue Date:
DOI: https://doi.org/10.1134/S0031030114050098